236 lines
32 KiB
XML
236 lines
32 KiB
XML
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<mods:title id="7279BA83BCEE2322E32906B710124B3F">A molecular phylogeny of Pseudocrangonyx from Japan, including a new subterranean species (Crustacea, Amphipoda, Pseudocrangonyctidae)</mods:title>
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<mods:namePart id="C2DB91908B5E3A65FC01C3595D5B50A3">Tomikawa, Ko</mods:namePart>
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<mods:namePart id="AC082953A0EFA4CE1212DCA4301DF318">Nakano, Takafumi</mods:namePart>
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<mods:namePart id="0D2F05018A725C1238C297B46532B497">Sato, Anna</mods:namePart>
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<mods:namePart id="6B2A3408FDD3A66BF19A2A95EE5FB19F">Onodera, Yutaka</mods:namePart>
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<mods:namePart id="A5AE645B1F9A7AEBC29B6926ADF9B132">Ohtaka, Akifumi</mods:namePart>
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<mods:title id="2D84E1BCB16E1ADE5D8D2F0DCB86B9C9">Zoosystematics and Evolution</mods:title>
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<mods:part id="668BDB346B26FFB8B069B05FABE577BE">
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<mods:date id="BA96B8FEF9AB08D51CBB3845E13ECA53">2016</mods:date>
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<mods:number id="3BB40205A99B601AD85306970163AB55">92</mods:number>
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<mods:identifier id="0EF848266603A5ABA94D197D677EDAED" type="DOI">http://dx.doi.org/10.3897/zse.92.10176</mods:identifier>
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<treatment id="58B7513EAA34ED62280265FDF8CE1661" ID-GBIF-Taxon="127869981" LSID="urn:lsid:zoobank.org:act:A821F75A-4FCB-40F7-B8AE-48289A696725" httpUri="http://treatment.plazi.org/id/58B7513EAA34ED62280265FDF8CE1661" lastPageId="7" lastPageNumber="193" pageId="2" pageNumber="189">
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<subSubSection id="E7B67B6AF2AD9023144B98D30EAEAC0E" pageId="2" pageNumber="189" type="multiple">
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<paragraph id="810837C0492CDED3F84D5FD9348BC58B" pageId="2" pageNumber="189">Taxon classification Animalia Amphipoda Pseudocrangonyctidae</paragraph>
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</subSubSection>
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<subSubSection id="21C93BEC221E582B5332028F703F18DA" pageId="2" pageNumber="189" type="nomenclature">
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<paragraph id="225A2902209CB004C9DA3B5877E8CD38" pageId="2" pageNumber="189">
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<taxonomicName id="649AF3FDCB07D3945FB5698CC610E174" ID-CoL="4NV54" LSID="http://zoobank.org/A821F75A-4FCB-40F7-B8AE-48289A696725" authority="Tomikawa & Sato" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis Tomikawa & Sato</taxonomicName>
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<taxonomicNameLabel id="63A99F29CB704268DB7586B1C229AD0F" pageId="2" pageNumber="189">sp. n.</taxonomicNameLabel>
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Figs 2, 3, 4, 5, 6, 7, 8, 9
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</paragraph>
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</subSubSection>
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<subSubSection id="A8F24BB1F776A15FC0B284DC436B1473" pageId="2" pageNumber="189" type="reference_group">
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<paragraph id="A0F5502423CA70EF0CA52D46AFF74654" pageId="2" pageNumber="189">
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<taxonomicName id="27DD0FC757380432EB1887414E855D4C" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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New Japanese name: Gudarimekura-yokoebi
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</paragraph>
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</subSubSection>
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<subSubSection id="DA3FE9980E8964E931F71B6267A67808" lastPageId="3" lastPageNumber="190" pageId="2" pageNumber="189" type="type material">
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<paragraph id="598E426ABC4F36E291021EB8E9466943" pageId="2" pageNumber="189">Type materials.</paragraph>
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<paragraph id="8D5B66DC43083DDF5FB1F4C1F774F37D" lastPageId="3" lastPageNumber="190" pageId="2" pageNumber="189">
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Holotype: Male (3.9 mm), NSMT-Cr 24603, Gudari-numa Stream (
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<geoCoordinate id="20982FD34C1CB488EA5781E0AB6616F8" direction="north" orientation="latitude" precision="1" value="40.672558">40°40'21.2″N</geoCoordinate>
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,
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<geoCoordinate id="5F85194417C9EA82C534C883A2811EAF" direction="east" orientation="longitude" precision="1" value="140.93184">140°55'54.6″E</geoCoordinate>
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, elev. 589 m), Komagome, Aomori, Aomori Prefecture, Japan, 4 June 2014, collected by A. Ohtaka. Paratypes: 1 female (3.1 mm), NSMT-Cr 24604, locality same as for holotype, 21 June 2015, collected by
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<pageBreakToken id="2697BAE26C3619BEAB1DDDD0B6D07BF1" pageId="3" pageNumber="190" start="start">A</pageBreakToken>
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. Ohtaka; 1 female (4.4 mm), NSMT-Cr 24605, locality same as for holotype, 14 March 2015, collected by A. Ohtaka; 1 male (2.5 mm) and 3 females (1.4-2.8 mm), NSMT-Cr 24606, locality same as for holotype, 23 May 2015, collected by A. Ohtaka; 4 females (2.0-2.9 mm) KUZ Z1746, data same as for holotype, collected by A. Ohtaka.
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</paragraph>
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</subSubSection>
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<subSubSection id="396643107224B92B702B925B015700BE" pageId="3" pageNumber="190" type="type locality">
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<paragraph id="E58DF20A7593A102DCBC6260B4207E01" pageId="3" pageNumber="190">Type locality.</paragraph>
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<paragraph id="4F5132AB536534E65C7454200129BD3A" pageId="3" pageNumber="190">Japan, Aomori Prefecture: Aomori, Gudari-numa Stream (northern Honshu).</paragraph>
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</subSubSection>
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<subSubSection id="92351AB28A92A947EFBC10476C983F35" lastPageId="5" lastPageNumber="192" pageId="3" pageNumber="190" type="description">
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<paragraph id="F125DB0D29FC84AE7041DBD581FFFF1D" pageId="3" pageNumber="190">Description.</paragraph>
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<paragraph id="A78415A809585AD92ED7240F953763D2" pageId="3" pageNumber="190">
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Male [NSMT-Cr 24603, 3.9 mm]. Head (Fig. 2) with short dorsal setae; rostrum reduced; lateral cephalic lobe rounded; antennal sinus shallow with rounded angle; eyes absent. Pereonites 1-7 with short dorsal setae (Fig. 2); posterolateral margin of pereonites 5-7 with seta (Fig. 2). Dorsal margin of pleonites 1-3 with 5, 6, and 8 setae, respectively (Fig. 7
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<normalizedToken id="EF916F7CCE4C351D4C45FEA438D8B61F" originalValue="I–K">I-K</normalizedToken>
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). Dorsal margin of urosomites 1 and 2 each with 4 setae (Fig. 7L, M), dorsal margin of urosomite 3 lacking setae (Fig. 7N). Posterior margin of epimeral plate 1 with 2 setae, posteroventral corner rounded with 1 seta (Fig. 7P); ventral and posterior margins of plate 2 with 1 and 2 setae, respectively, posteroventral corner rounded, with 1 seta (Fig. 7Q); ventral and posterior margins of plate 3 each with 1 seta, posteroventral corner rounded, with 1 seta (Fig. 7R).
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</paragraph>
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<caption id="5C2181B95E44BA8F0BBA2571BFED788A" pageId="3" pageNumber="190">
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<paragraph id="770E879CFE8087466A3976A98CC9C5AD" pageId="3" pageNumber="190">
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Figure 2.
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<taxonomicName id="DB58BD2AE04A3780A0C6808C013B62FC" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. Habitus, lateral view.
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</paragraph>
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</caption>
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<paragraph id="610FC65B8C3E43A8A400A3430012F421" pageId="3" pageNumber="190">Antenna 1 (Fig. 3A) 0.47 times as long as body length, peduncular articles 1 to 3 in length ratio of 1.0: 0.6: 0.4; accessory flagellum 2-articulate, terminal article with 3 setae; primary flagellum 12-articulate, 1 aesthetasc on some articles. Antenna 2 (Fig. 3B) 0.60 times as long as antenna 1; peduncular article 5 with calceolus; flagellum 0.55 times as long as peduncular articles 4 and 5 combined, consisting of 4 articles, first 2 of which with calceolus.</paragraph>
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<paragraph id="F795B3E254EFE670F54E47E4FFFD287A" pageId="3" pageNumber="190">
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Upper lip (= labrum) (Fig. 3C) with rounded anterior margin, bearing fine setae. Mandibles (Fig. 3E, F) with left and right incisors both 5-dentate; left lacinia mobilis 4-dentate, right lacinia bifid, bearing many teeth; molar process triturative, molar of right mandible with accessory seta; accessory setal rows of left and right mandibles with 3 and 2 weakly pectinate setae, respectively; palp 3-articulate, article 3 with 1 A-, 4-5 D-, and 4 E-setae. Lower lip (Fig. 3D) with broad outer lobes, mandibular process of outer lobe rounded apically; inner lobes indistinct. Maxilla 1 (Fig. 3G) with inner and outer plates, and palp; inner plate subovate, its medial margin with 3 plumose setae; outer plate subrectangular with 7 serrate teeth apically; palp 2-articulate, longer than outer plate, article 1 lacking marginal setae, article 2 with 3 apical and 1 subapical robust setae. Maxilla 2 (Fig. 3H) with oblique inner row of 3 plumose setae on inner plate. Maxilliped (Fig. 4
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<normalizedToken id="53EA28ACCD2A7B36C5CBABEACCDE22DD" originalValue="A–C">A-C</normalizedToken>
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) with inner and outer plates, and palp; inner plate (Fig. 4C) with 3 apical and 2 subapical robust setae; outer plate (Fig. 4B) with 1 apical plumose seta and 1 apical robust seta and 5 medial slender setae; palp (Fig. 4A) 4-articulate, medial margin of article 2 lined with setae, article 4 with nail.
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</paragraph>
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<paragraph id="2E7FFAF37E2B7E2720DE266ACA7C7B8F" lastPageId="4" lastPageNumber="191" pageId="3" pageNumber="190">
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Gnathopod 1 (= pereopod 1) (Fig. 4D, E) with subquadrate coxa, bearing setae on anterodistal corner of coxa, width 2.0 times as long as depth; anterior margin of basis bare, posterior margin of basis with 5 setae; posterodistal corner of carpus with slender setae, some weakly pectinate; propodus stout, subchelate, palmar margin with 6 medial and 5 lateral robust setae, some distally notched (Fig. 4E); posterior margin of dactylus dentate (Fig. 4E). Gnathopod 2 (= pereopod 2) (Fig. 4F, G) with subquadrate coxa bearing setae on its anterior margin and posterodistal corner, width 1.9 times as long as depth; anterior and posterior margins of basis with 0 and 5 setae, respectively; posterodistal corner of carpus with slender seta, some weakly pectinate; propodus stout, subchelate with 6 medial and 8 lateral robust setae along palmar margin, some
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<pageBreakToken id="5F02408C3E8FAA98D4D3B330E57DB315" pageId="4" pageNumber="191" start="start">distally</pageBreakToken>
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notched (Fig. 4G); posterior margin of dactylus dentate (Fig. 4G). Pereopod 3 (Fig. 5A) with subquadrate coxa bearing setae on its anterodistal and and posteroventral corners, width 1.8 times as long as depth; anterior and posterior margins of basis with 2 and 5 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 1.0: 0.9; posterior margin of dactylus with 1 seta. Pereopod 4 (Fig. 5B) with subquadrate coxa bearing setae on its anterior margin, anterodistal and posteroventral corners, width 1.8 times as long as depth; anterior and posterior margins of basis each with 3 setae; merus, carpus, and propodus in length ratio of 1.0: 0.8: 0.8; posterior margin of dactylus with 1 seta. Pereopod 5 (Fig. 5C, D) with weakly bilobed coxa bearing setae on anterior and posterior lobes; anterior and posterior margins of basis with 3 and 4 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.8: 0.9; posterior margin of dactylus with 2 setae (Fig. 5D). Pereopod 6 (Fig. 6A, B) with coxa bearing concave lower margin, posteroproximal corner with 1 seta; anterior and posterior margins of basis with 4 and 3 setae, respectively; merus, carpus, and propodus in length ratio of 1.0: 0.9: 1.1; posterior margin of dactylus with 2 setae (Fig. 6B). Pereopod 7 (Fig. 6
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<normalizedToken id="277F11B9A6E3AAEF059A6B47340BF52B" originalValue="C–E">C-E</normalizedToken>
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) with subtriangular coxa, bearing 1 seta on posteroproximal corner; anterior and posterior margins of basis each with 3 setae; merus, carpus, and propodus in length ratio of 1.0: 1.1: 1.2; posterior margin of dactylus with 2 setae (Fig. 6E).
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</paragraph>
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<caption id="325F400B4FAB8CBDB1BBDD1184489A89" pageId="4" pageNumber="191">
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<paragraph id="9EBC0FA6C3792F4441C0317D6A716375" pageId="4" pageNumber="191">
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Figure 3.
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<taxonomicName id="10E2B66C710454D455327E9BA667DFFB" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. A antenna 1, medial view; B antenna 2, medial view; C upper lip, anterior view; D lower lip, ventral view; E left mandible, medial view; F right mandible, medial view; G maxilla 1, dorsal view; H maxilla 2, dorsal view.
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</paragraph>
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</caption>
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<caption id="7738551BDB668AFD7693AB72583599CD" pageId="4" pageNumber="191">
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<paragraph id="604A3A6AB92DE121DFE4A66616B49936" pageId="4" pageNumber="191">
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Figure 4.
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<taxonomicName id="DC37B0831B74E45E99DB78F76CD9CCE2" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. A maxilliped, dorsal view; B inner plate of maxilliped, dorsal view; C outer plate of maxilliped, dorsal view; D gnathopod 1, lateral view; E palmar margin of propodus and dactylus of gnathopod 1, medial view; F gnathopod 2, lateral view; G palmar margin of propodus and dactylus of gnathopod 2, medial view.
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</paragraph>
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</caption>
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<caption id="9908CB5FA1A68B3DAF5E9BCD2D319409" pageId="4" pageNumber="191">
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<paragraph id="F8AE017B704870800A28AE5CDB518DFF" pageId="4" pageNumber="191">
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Figure 5.
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<taxonomicName id="EE68870B62327AD0AB7F18D113551C41" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. A pereopod 3, lateral view; B pereopod 4, lateral view; C pereopod 5, lateral view; D dactylus of pereopod 5, lateral view.
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</paragraph>
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</caption>
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<caption id="722795273B0F125DD9E830473172E5AA" pageId="4" pageNumber="191">
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<paragraph id="E3D4C23D6C1D74D985AEE56BC55A88B3" pageId="4" pageNumber="191">
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Figure 6.
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<taxonomicName id="3D2F285F1E9E610389DEB51972CE70F2" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
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Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. A pereopod 6, lateral view; B dactylus of pereopod 6, lateral view; C
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<normalizedToken id="C9C4AB11AEEB50FD9890485B22DB3DE6" originalValue="coxa–carpus">coxa-carpus</normalizedToken>
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of pereopod 6, lateral view; D
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<normalizedToken id="CEA96ABC73D069277FED6630A9D5844B" originalValue="propodus–dactylus">propodus-dactylus</normalizedToken>
|
||
of pereopod 6, lateral view; E dactylus of pereopod 7, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AD804A945A0FFA1010CDB2F513782B87" pageId="4" pageNumber="191">Coxal gills (Fig. 2) on gnathopod 2 and pereopods 3-6; sternal gills absent.</paragraph>
|
||
<paragraph id="CEB829DDB65C21683D1BCA0A080E0E32" pageId="4" pageNumber="191">Peduncles of pleopods 1 and 2 (Fig. 7A, C) with 2 and 1 setae on outer margins, respectively; peduncle of pleopod 3 (Fig. 7D) lacking marginal setae. Pleopods 1-3 each with paired retinacula (Fig. 7B), and lacking bifid setae (clothes-pin setae) on inner basal margin of inner ramus.</paragraph>
|
||
<paragraph id="B4812BDFA7E8A207B8187568B3AB7563" pageId="4" pageNumber="191">Uropod 1 (Fig. 7E) with basofacial slender seta on peduncle; inner ramus 0.70 times as long as peduncle, inner margin of former with 2 robust setae, outer margin bare, basal part with 3 slender setae; outer ramus 0.76 times as long as inner, its inner and outer margins with 0 and 2 robust setae, respectively. Uropod 2 (Fig. 7F, G) with inner and outer rami; inner ramus 0.90 times as long as peduncle, its inner margin with 4 robust setae, outer margin bare, distal part with 2 serrate and 4 simple robust setae and 1 slender seta (Fig. 7G); outer ramus 0.89 times as long as inner ramus, its outer margin with 1 robust seta. Uropod 3 (Fig. 7H) with peduncle 0.33 times as long as outer ramus, with 1 dorsal and 3 ventral robust setae; inner ramus absent; outer ramus 2-articulate, proximal article with robust setae, terminal article 0.36 times as long as proximal article, with 3 distal setae.</paragraph>
|
||
<caption id="41DEAF446C63EC96C7357463501DD729" pageId="4" pageNumber="191">
|
||
<paragraph id="2C331EC6A2DD9E7A8657E7B36FAB2F17" pageId="4" pageNumber="191">
|
||
Figure 7.
|
||
<taxonomicName id="41840E13A0A6E8BAF465F9EBEB6AC8EB" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="4" pageNumber="191" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
Tomikawa & Sato, sp. n., holotype, male (3.9 mm), NSMT-Cr 24603. A pleopod 1, anterior view; B retinacula on peduncle of pleopod 1, anterior view; C pleopod 2, anterior view; D pleopod 3, anterior view; E uropod 1, dorsal view; F uropod 2, dorsal view; G distal part of inner ramus of uropod 2, dorsal view; H uropod 3, dorsal view;
|
||
<normalizedToken id="967FE868FB291A5D4A3144DFC8E4387E" originalValue="I–K">I-K</normalizedToken>
|
||
dorsal margins of pleonites 1-3, respectively, dorsal views;
|
||
<normalizedToken id="EC6B76EE324A86E082FA57DB8EEB3198" originalValue="L–N">L-N</normalizedToken>
|
||
dorsal margins of urosomites 1-3, respectively, dorsal views; O telson, dorsal view;
|
||
<normalizedToken id="B33402B697186C0B8EA38538E0E443B4" originalValue="P–R">P-R</normalizedToken>
|
||
epimeral plates 1-3, respectively, lateral views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="94AB75AF2EB72443A0C5EA54B2D4F0AC" pageId="5" pageNumber="192">
|
||
<pageBreakToken id="167DDE1FA1CF12873CE3BD00FE893A3E" pageId="5" pageNumber="192" start="start">Telson</pageBreakToken>
|
||
(Fig. 7O) length 1.2 times as long as wide, cleft for 0.08 times of length, each telson lobe with 2 lateral long penicillate setae, 2 apical robust and 1 apical short penicillate setae.
|
||
</paragraph>
|
||
<paragraph id="E32052896A70A13608331D69DD3DDE04" pageId="5" pageNumber="192">Female [NSMT-Cr 24604, 3.1 mm]. Antenna 1 (Fig. 8A) 0.58 times as long as body length, primary flagellum 12-articulate. Antenna 2 (Fig. 8B) 0.64 times as long as antenna 1, calceoli absent; flagellum 0.64 times as long as peduncular articles 4 and 5 combined, 5-articulate, first 2 of which with 1 robust seta, lacking calceoli.</paragraph>
|
||
<caption id="770B84E90F55808ACF1E8A552F0F7D1D" pageId="5" pageNumber="192">
|
||
<paragraph id="93E41C8F7F6E7E0007D708370CFA0AC2" pageId="5" pageNumber="192">
|
||
Figure 8.
|
||
<taxonomicName id="C9D59B16BBD9155A660D75CB9045E50A" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="5" pageNumber="192" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
Tomikawa & Sato, sp. n., paratype, female (3.1 mm), NSMT-Cr 24604. A antenna 1, medial view; B antenna 2, medial view; C gnathopod 1, lateral view; D palmar margin of propodus and dactylus of gnathopod 1, medial view; E gnathopod 2, lateral view; F palmar margin of propodus and dactylus of gnathopod 2, medial view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B350154918AE10EFFCCB9BA2358D9F44" pageId="5" pageNumber="192">Lacinia mobilis of left mandible 5-dentate.</paragraph>
|
||
<paragraph id="C748EC3C26B2D4F4BDEDA74B1CF09032" pageId="5" pageNumber="192">Gnathopod 1 (Fig. 8C, D) with coxa width 1.9 times as long as depth; palmar margin (Fig. 8D) with 4 medial and 4 lateral distally notched robust setae. Gnathopod 2 (Fig. 8E, F) with coxa width 1.8 times as long as depth; palmar margin (Fig. 8F) with 9 medial and 5 lateral robust setae, some distally notched.</paragraph>
|
||
<paragraph id="E4F34CB520C3B294A32B8504DF6730C4" pageId="5" pageNumber="192">Brood plates (Fig. 8G) slender, with numerous setae, on gnathopod 2 and pereopods 3-5.</paragraph>
|
||
<paragraph id="D003296C1CAF7942222528E985E7FD4C" pageId="5" pageNumber="192">Uropod 1 (Fig. 9A) with 3 robust setae on inner margin of inner ramus, basal part with 2 slender setae; outer ramus 0.80 times as long as inner. Uropod 2 (Fig. 9B) with 6 simple robust setae and 1 slender seta on distal part of inner ramus. Uropod 3 (Fig. 9C) with peduncle 0.32 times as long as outer ramus; terminal article of outer ramus 0.35 times as long as proximal article.</paragraph>
|
||
<caption id="FD1F74A52845373546A4720CDBA1E1CB" pageId="5" pageNumber="192">
|
||
<paragraph id="9E5E0D4E8879348D0D616B384E92E0D5" pageId="5" pageNumber="192">
|
||
Figure 9.
|
||
<taxonomicName id="04A8659A379D3C394C3D71AC4FB87CBD" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="5" pageNumber="192" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
Tomikawa & Sato, sp. n., paratype, female (3.1 mm), NSMT-Cr 24604.
|
||
<normalizedToken id="A8524E7144DF0E0C2FE3D2FF91FFEE61" originalValue="A–C">A-C</normalizedToken>
|
||
uropods 1-3, respectively, dorsal views.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="76A8FCBA6DBAE918545728493D2A3495" pageId="5" pageNumber="192" type="etymology">
|
||
<paragraph id="B82E1B65F204EE0D45D00AA1438C693A" pageId="5" pageNumber="192">Etymology.</paragraph>
|
||
<paragraph id="25BC05642E1E67493BF140AF65A30F7C" pageId="5" pageNumber="192">The specific name is an adjective derived from Gudari-numa, the type locality of the new species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55BF0E5865BE3C63DB58830C945B9547" pageId="5" pageNumber="192" type="distribution">
|
||
<paragraph id="C400E9E401AE8B7EC6118872EEECBBD4" pageId="5" pageNumber="192">Distribution and habitat.</paragraph>
|
||
<paragraph id="FE1DE2706B28645192E3A09F042F96E8" pageId="5" pageNumber="192">This species is known only from the type locality. The specimens were collected from interstitial water in the gravelly bottom. Water temperature of the habitat was stable and around 7°C throughout the year (Baba and Ohtaka unpublished).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DEF7A64BF5A91F07B0806364C902F67" lastPageId="7" lastPageNumber="194" pageId="6" pageNumber="193" type="remarks">
|
||
<paragraph id="42D220F4AC10AC601F43B0A94DA92224" pageId="6" pageNumber="193">
|
||
<pageBreakToken id="3ACEE365067FEAE20561AF201883D3F3" pageId="6" pageNumber="193" start="start">Remarks</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4CA743DBED37700FAF18556C13741402" pageId="6" pageNumber="193">
|
||
<taxonomicName id="F3D29EAE863308AF6B4AC3FD40A74578" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
is morphologically similar to
|
||
<taxonomicName id="55D9DBC8B99DB95A9AF3A21FBBF20694" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx coreanus" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="coreanus">Pseudocrangonyx coreanus</taxonomicName>
|
||
described from the Korean Peninsula. The deposited female paratypes of the latter species have calceoli on antenna 2 and pleopods without bifid setae on inner basal margin of inner ramus, which are features that were not mentioned in the original description (NSMT-Cr 13521-13522; Tomikawa and Onodera, personal observation).These two species share the following features: 1) relatively small body size (smaller than 6 mm), 2) eye completely absent, 3) carpus of gnathopod 2 without serrate robust setae on posterodistal corners, 4) outer margin or outer distal corner of pleopods 1 and 2 with setae, 5) inner basal margin of inner ramus of pleopods without bifid setae, and 6) small number of articles (less than 5) of rami of pleopods. However,
|
||
<taxonomicName id="3291674BEF7ED9C5CF7042B54FA8EAD9" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="F6D73B2E19702C716594BFC33AEF4D14" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx coreanus" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="coreanus">Pseudocrangonyx coreanus</taxonomicName>
|
||
by the following features (features of
|
||
<taxonomicName id="236CACD712B5B5C65DD6146963E4A4F0" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx coreanus" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="coreanus">Pseudocrangonyx coreanus</taxonomicName>
|
||
in parentheses): 1) antenna 2 of female without calceoli (present); 2) palmar margins of gnathopods 1 and 2 with distally notched robust setae (absent); 3) inner margin of inner ramus of uropod 2 with 4 (0 or 1) robust setae; and 4) basal part of inner ramus of uropod 2 without slender seta (present).
|
||
</paragraph>
|
||
<paragraph id="60E6FC7F54FC7FFF0C19F311CF9B7806" lastPageId="7" lastPageNumber="194" pageId="6" pageNumber="193">
|
||
<taxonomicName id="44B662FE910FF16895828F55C90D561C" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
is also similar to
|
||
<taxonomicName id="CB01C6EA8A974B9C4A2686D32F33DFF6" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx febras" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="febras">Pseudocrangonyx febras</taxonomicName>
|
||
from river basin of Primorye, Russia in having 1) relatively small body size (smaller than 6.5 mm), 2) eye completely absent, 3) palmar margins of gnathopods 1 and 2 with distally notched robust setae, 4) small number of articles (less than 6) of rami of pleopods, and 5) urosomite 1 without basal setae. However,
|
||
<taxonomicName id="937DD35C068F1E34B5E452910909318D" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx gudariensis" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="gudariensis">Pseudocrangonyx gudariensis</taxonomicName>
|
||
is distinguished from the latter by the following features (features of
|
||
<taxonomicName id="FEDA2AE7B323B2DACE2D54768669073A" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudocrangonyx febras" order="Amphipoda" pageId="6" pageNumber="193" phylum="Arthropoda" rank="species" species="febras">Pseudocrangonyx febras</taxonomicName>
|
||
in parentheses): 1) carpus of gnathopod 2 without serrate robust setae on posterodistal cor
|
||
<pageBreakToken id="5D4B2A967CD63BBA7AD8A6B057FBBAEC" pageId="7" pageNumber="194" start="start">ners</pageBreakToken>
|
||
(present), 2) peduncle of pleopods 1 and 2 with setae (absent), and 3) article 2 of uropod 3 longer (shorter) than setae on distal part of article 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |