treatments-xml/data/58/0E/87/580E87FAFFE9FFA6FC56099EFD6E8EE7.xml

<document id="0E006CB85B92DA5D753FE34405D06588" ID-DOI="10.5281/zenodo.4508022" ID-ISSN="2345-7600" ID-Zenodo-Dep="4508022" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612542827268" checkinUser="carolina" docAuthor="Li, Xinzheng" docDate="2006" docId="580E87FAFFE9FFA6FC56099EFD6E8EE7" docLanguage="en" docName="RafflesBZool.54.2.361-372.pdf" docOrigin="Raffles Bulletin of Zoology 54 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Heterocarpus chani Li 2006, new species" docType="treatment" docVersion="4" lastPageNumber="367" masterDocId="A437FF82FFE8FFA1FFE20A7AFF8C8C3F" masterDocTitle="Additional Pandaloid Shrimps From The South China Sea (Crustacea: Decapoda: Caridea), With Description Of One New Species" masterLastPageNumber="372" masterPageNumber="361" pageNumber="362" updateTime="1698867420658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0401631BD488FA981BF2A7553BD163AA">Additional Pandaloid Shrimps From The South China Sea (Crustacea: Decapoda: Caridea), With Description Of One New Species</mods:title>
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<mods:namePart id="CF167F9F3A7674428F8778BD4275149F">Li, Xinzheng</mods:namePart>
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<mods:date id="770C41F8EC4DA01A5B5CBDABECA2C503">2006</mods:date>
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<taxonomicName id="17A74D6FFFE9FFA0FC56099EFB1C8FC3" ID-CoL="9CRKZ" authority="Li, 2006" authorityName="Li" authorityYear="2006" box="[948,1168,996,1020]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="362" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="E2D3EAFEFFE9FFA0FC56099EFB1C8FC3" bold="true" box="[948,1168,996,1020]" italics="true" pageId="1" pageNumber="362">Heterocarpus chani</emphasis>
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, 
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(
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)
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–
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, female (cl 
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) (
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), 
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, “Panglao 2005”, stn. CP2358, 
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8 
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52.1'N
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123 
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37.1'E
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, 
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, sandy, 
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.
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Fig. 1. 
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<emphasis id="E2D3EAFEFFE9FFA0FF5D0257FEF3847D" box="[191,383,2093,2114]" italics="true" pageId="1" pageNumber="362">Heterocarpus chani</emphasis>
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, 
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, female, holotype (cl 24.4 mm), Philippines, body, right lateral view. Scale bar = 5 mm.
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: 
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:
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(cl 
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, 
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(cl 
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), 
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, 
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(cl 
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), 
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, 
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7 
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18.83'N
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<geoCoordinate id="B593502BFFEAFFA3FF190A80FEF18D30" box="[251,381,250,271]" degrees="116" direction="east" minutes="12.29" orientation="longitude" pageId="2" pageNumber="363" precision="9" value="116.204834">
116 
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12.29'E
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, 
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, soft mud, AT, 
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. 
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<collectingCountry id="A8B0767CFFEAFFA3FF6A0B6DFF738D13" box="[136,255,279,300]" name="Philippines" pageId="2" pageNumber="363">Philippines</collectingCountry>
: “ 
<location id="D5786037FFEAFFA3FEF70B6DFEE68D13" LSID="urn:lsid:plazi:treatment:580E87FAFFE9FFA6FC56099EFD6E8EE7:D5786037FFEAFFA3FEF70B6DFEE68D13" box="[277,362,279,300]" country="Philippines" latitude="9.646667" longLatPrecision="129" longitude="123.765" name="Panglao" pageId="2" pageNumber="363">Panglao</location>
2005”:
</emphasis>
1 ovigerous female (cl 
<quantity id="175F9B09FFEAFFA3FD440B6DFC8C8D13" box="[678,768,279,300]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="2" pageNumber="363" unit="mm" value="33.2">33.2 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FF6C0B4EFF4D8D76" box="[142,193,308,329]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="363">ZRC</collectionCode>
), stn. CP2332, 
<geoCoordinate id="B593502BFFEAFFA3FEB40B4EFE278D76" box="[342,427,307,329]" degrees="9" direction="north" minutes="38.8" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.646667">
9 
<emphasis id="E2D3EAFEFFEAFFA3FE800B49FEE08D77" bold="true" box="[354,364,307,328]" pageId="2" pageNumber="363">°</emphasis>
38.8'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FE500B4EFD938D76" box="[434,543,307,329]" degrees="123" direction="east" minutes="45.9" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.765">
123 
<emphasis id="E2D3EAFEFFEAFFA3FE340B49FE6C8D77" bold="true" box="[470,480,307,328]" pageId="2" pageNumber="363">°</emphasis>
45.9'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FDCB0B4EFD198D76" box="[553,661,308,329]" metricMagnitude="2" metricUnit="m" metricValue="4.475" metricValueMax="4.77" metricValueMin="4.18" pageId="2" pageNumber="363" unit="m" value="447.5" valueMax="477.0" valueMin="418.0">
<elevation id="5B8AD1DFFFEAFFA3FDCB0B4EFD198D76" box="[553,661,308,329]" metricMagnitude="2" metricUnit="m" metricValue="4.475" metricValueMax="4.77" metricValueMin="4.18" pageId="2" pageNumber="363" unit="m" value="447.5" valueMax="477.0" valueMin="418.0">418-477 m</elevation>
</quantity>
, muddy, 
<date id="A419102CFFEAFFA3FD0C0B4EFF608D59" pageId="2" pageNumber="363" value="2005-05-22">
<collectingDate id="B45DE9C4FFEAFFA3FD0C0B4EFF608D59" pageId="2" pageNumber="363" value="2005-05-22">22 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="60CF3CB1FFEAFFA3FF160B2BFD228DBD" collectingDate="2005-05-22" collectionCode="IOCAS" country="Philippines" elevation="448" latitude="9.646667" longLatPrecision="129" longitude="123.765" pageId="2" pageNumber="363" specimenCount="1" typeStatus="paratype">
1 ovigerous female (cl 
<quantity id="175F9B09FFEAFFA3FE290B2BFDAC8D59" box="[459,544,337,358]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="2" pageNumber="363" unit="mm" value="32.2">32.2 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FDD00B2BFDF28D59" box="[562,638,337,358]" country="China" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="363">IOCAS</collectionCode>
), stn. CP2332, 
<geoCoordinate id="B593502BFFEAFFA3FF6A0B17FF508DBD" box="[136,220,365,386]" degrees="9" direction="north" minutes="38.8" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.646667">
9 
<emphasis id="E2D3EAFEFFEAFFA3FF710B17FF118DBD" bold="true" box="[147,157,365,386]" pageId="2" pageNumber="363">°</emphasis>
38.8'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FF070B17FEDE8DBD" box="[229,338,365,386]" degrees="123" direction="east" minutes="45.9" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.765">
123 
<emphasis id="E2D3EAFEFFEAFFA3FEEB0B17FE9F8DBD" bold="true" box="[265,275,365,386]" pageId="2" pageNumber="363">°</emphasis>
45.9'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FEBC0B17FE408DBD" box="[350,460,365,386]" metricMagnitude="2" metricUnit="m" metricValue="4.475" metricValueMax="4.77" metricValueMin="4.18" pageId="2" pageNumber="363" unit="m" value="447.5" valueMax="477.0" valueMin="418.0">
<elevation id="5B8AD1DFFFEAFFA3FEBC0B17FE408DBD" box="[350,460,365,386]" metricMagnitude="2" metricUnit="m" metricValue="4.475" metricValueMax="4.77" metricValueMin="4.18" pageId="2" pageNumber="363" unit="m" value="447.5" valueMax="477.0" valueMin="418.0">418-477 m</elevation>
</quantity>
, muddy, 
<date id="A419102CFFEAFFA3FDCB0B17FD228DBD" box="[553,686,365,386]" pageId="2" pageNumber="363" value="2005-05-22">
<collectingDate id="B45DE9C4FFEAFFA3FDCB0B17FD228DBD" box="[553,686,365,386]" pageId="2" pageNumber="363" value="2005-05-22">22 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FD580B17FC8A8DBD" box="[698,774,365,386]" pageId="2" pageNumber="363" type="male">2 males</specimenCount>
(cl 18.8, 
<quantity id="175F9B09FFEAFFA3FF3A0BF0FEA18DA0" box="[216,301,394,415]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="2" pageNumber="363" unit="mm" value="20.3">20.3 mm</quantity>
), 
<materialsCitation id="60CF3CB1FFEAFFA3FEDE0BF0FE6E8DE6" collectingDate="2005-05-22" collectionCode="NTOU" country="Philippines" elevation="581" latitude="9.636666" longLatPrecision="129" longitude="123.725" pageId="2" pageNumber="363" specimenCount="8" specimenCount-female="8" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FEDE0BF0FE1A8DA0" box="[316,406,394,415]" pageId="2" pageNumber="363" type="female">8 females</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FE550BF0FDB18DA0" box="[439,573,394,415]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" metricValueMax="3.54" metricValueMin="2.18" pageId="2" pageNumber="363" unit="mm" value="28.6" valueMax="35.4" valueMin="21.8">21.8-35.4 mm</quantity>
), 3 ovigerous females (cl 26.4, 27.1, 
<quantity id="175F9B09FFEAFFA3FEE90BDDFED38D83" box="[267,351,423,444]" metricMagnitude="-2" metricUnit="m" metricValue="2.79" pageId="2" pageNumber="363" unit="mm" value="27.9">27.9 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FE920BDDFE3A8D83" box="[368,438,423,444]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="363">NTOU</collectionCode>
), stn. CP2333, 
<geoCoordinate id="B593502BFFEAFFA3FDA30BDDFD188D83" box="[577,660,422,444]" degrees="9" direction="north" minutes="38.2" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.636666">
9 
<emphasis id="E2D3EAFEFFEAFFA3FDAE0BDCFDDA8D84" bold="true" box="[588,598,422,443]" pageId="2" pageNumber="363">°</emphasis>
38.2'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FD7A0BDDFC8E8D83" box="[664,770,422,444]" degrees="123" direction="east" minutes="43.5" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.725">
123 
<emphasis id="E2D3EAFEFFEAFFA3FD590BDCFD498D84" bold="true" box="[699,709,422,443]" pageId="2" pageNumber="363">°</emphasis>
43.5'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FF6A0BBEFE898DE6" box="[136,261,452,473]" metricMagnitude="2" metricUnit="m" metricValue="5.8075" metricValueMax="5.96" metricValueMin="5.654999999999999" pageId="2" pageNumber="363" unit="m" value="580.75" valueMax="596.0" valueMin="565.5">
<elevation id="5B8AD1DFFFEAFFA3FF6A0BBEFE898DE6" box="[136,261,452,473]" metricMagnitude="2" metricUnit="m" metricValue="5.8075" metricValueMax="5.96" metricValueMin="5.654999999999999" pageId="2" pageNumber="363" unit="m" value="580.75" valueMax="596.0" valueMin="565.5">565.5-596 m</elevation>
</quantity>
, muddy, 
<date id="A419102CFFEAFFA3FEBC0BBEFE6E8DE6" box="[350,482,452,473]" pageId="2" pageNumber="363" value="2005-05-22">
<collectingDate id="B45DE9C4FFEAFFA3FEBC0BBEFE6E8DE6" box="[350,482,452,473]" pageId="2" pageNumber="363" value="2005-05-22">22 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="60CF3CB1FFEAFFA3FE0E0BBEFF618E2D" collectingDate="2005-05-22" collectionCode="ZRC" country="Philippines" elevation="606" latitude="9.63" longLatPrecision="129" longitude="123.685" pageId="2" pageNumber="363" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FE0E0BBEFDC68DE6" box="[492,586,452,473]" pageId="2" pageNumber="363" type="female">3 females</specimenCount>
(cl 21.5, 27.4, 
<quantity id="175F9B09FFEAFFA3FD3E0BBEFF238DC9" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="2" pageNumber="363" unit="mm" value="29.5">29.5 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FF230B9BFF788DC9" box="[193,244,481,502]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="363">ZRC</collectionCode>
), stn. CP2334, 
<geoCoordinate id="B593502BFFEAFFA3FE610B9BFE548DC9" box="[387,472,480,502]" degrees="9" direction="north" minutes="37.8" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.63">
9 
<emphasis id="E2D3EAFEFFEAFFA3FE6D0B9AFE158DCA" bold="true" box="[399,409,480,501]" pageId="2" pageNumber="363">°</emphasis>
37.8'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FE3F0B9BFDC68DC9" box="[477,586,480,502]" degrees="123" direction="east" minutes="41.1" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.685">
123 
<emphasis id="E2D3EAFEFFEAFFA3FDE30B9AFD878DCA" bold="true" box="[513,523,480,501]" pageId="2" pageNumber="363">°</emphasis>
41.1'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FDB10B9BFD2F8DC9" box="[595,675,481,502]" metricMagnitude="2" metricUnit="m" metricValue="6.062" pageId="2" pageNumber="363" unit="m" value="606.2">
<elevation id="5B8AD1DFFFEAFFA3FDB10B9BFD2F8DC9" box="[595,675,481,502]" metricMagnitude="2" metricUnit="m" metricValue="6.062" pageId="2" pageNumber="363" unit="m" value="606.2">606.2 m</elevation>
</quantity>
, sandy, 
<date id="A419102CFFEAFFA3FD0C0B9BFF618E2D" pageId="2" pageNumber="363" value="2005-05-22">
<collectingDate id="B45DE9C4FFEAFFA3FD0C0B9BFF618E2D" pageId="2" pageNumber="363" value="2005-05-22">22 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FF1B0B87FECA8E2D" box="[249,326,509,530]" pageId="2" pageNumber="363" type="male">5 males</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FE930B87FE738E2D" box="[369,511,509,530]" metricMagnitude="-2" metricUnit="m" metricValue="2.135" metricValueMax="2.56" metricValueMin="1.71" pageId="2" pageNumber="363" unit="mm" value="21.35" valueMax="25.6" valueMin="17.1">17.1-25.6 mm</quantity>
), 
<specimenCount id="C6A1FD65FFEAFFA3FDF10B87FDFF8E2D" box="[531,627,509,530]" pageId="2" pageNumber="363" type="female">3 females</specimenCount>
(cl 24.4, 29.2, 
<quantity id="175F9B09FFEAFFA3FF6A0860FF688E10" box="[136,228,538,559]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="2" pageNumber="363" unit="mm" value="29.4">29.4 mm</quantity>
), 
<materialsCitation id="60CF3CB1FFEAFFA3FF1B0860FD018E73" collectingDate="2005-05-23" collectionCode="IOCAS" country="Philippines" elevation="800" latitude="9.408334" longLatPrecision="129" longitude="123.82833" pageId="2" pageNumber="363" specimenCount="1" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FF1B0860FEB88E10" box="[249,308,538,559]" pageId="2" pageNumber="363" type="generic">1 juv.</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FEBD0860FE228E10" box="[351,430,538,559]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="2" pageNumber="363" unit="mm" value="9.9">9.9 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FE270860FD9F8E10" box="[453,531,538,559]" country="China" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="363">IOCAS</collectionCode>
), stn. CP2341, 
<geoCoordinate id="B593502BFFEAFFA3FD520860FC8A8E10" box="[688,774,538,559]" degrees="9" direction="north" minutes="24.5" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.408334">
9 
<emphasis id="E2D3EAFEFFEAFFA3FD5E0860FD4A8E10" bold="true" box="[700,710,538,559]" pageId="2" pageNumber="363">°</emphasis>
24.5'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FF6A084DFF798E73" box="[136,245,566,588]" degrees="123" direction="east" minutes="49.7" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.82833">
123 
<emphasis id="E2D3EAFEFFEAFFA3FF49084CFF398E74" bold="true" box="[171,181,566,587]" pageId="2" pageNumber="363">°</emphasis>
49.7'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FEE2084DFEE18E73" box="[256,365,567,588]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.879999999999999" metricValueMin="7.12" pageId="2" pageNumber="363" unit="m" value="800.0" valueMax="888.0" valueMin="712.0">
<elevation id="5B8AD1DFFFEAFFA3FEE2084DFEE18E73" box="[256,365,567,588]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.879999999999999" metricValueMin="7.12" pageId="2" pageNumber="363" unit="m" value="800.0" valueMax="888.0" valueMin="712.0">712-888 m</elevation>
</quantity>
, sandy/muddy, 
<date id="A419102CFFEAFFA3FDEA084DFD018E73" box="[520,653,567,588]" pageId="2" pageNumber="363" value="2005-05-23">
<collectingDate id="B45DE9C4FFEAFFA3FDEA084DFD018E73" box="[520,653,567,588]" pageId="2" pageNumber="363" value="2005-05-23">23 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FD7B084DFD698E73" box="[665,741,567,588]" pageId="2" pageNumber="363" type="male">7 males</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FF6A082EFE9F8E56" box="[136,275,596,617]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.36" metricValueMin="1.64" pageId="2" pageNumber="363" unit="mm" value="20.0" valueMax="23.6" valueMin="16.4">16.4-23.6 mm</quantity>
), 
<specimenCount id="C6A1FD65FFEAFFA3FEC6082EFE028E56" box="[292,398,596,617]" pageId="2" pageNumber="363" type="female">12 females</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FE56082EFDCC8E56" box="[436,576,596,617]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" metricValueMax="2.82" metricValueMin="1.92" pageId="2" pageNumber="363" unit="mm" value="23.7" valueMax="28.2" valueMin="19.2">19.2-28.2 mm</quantity>
), 
<materialsCitation id="60CF3CB1FFEAFFA3FDB3082EFD3D8EB9" collectingDate="2005-05-26" collectionCode="NTOU" country="Philippines" elevation="583" latitude="8.868333" location="Philippines" longLatPrecision="129" longitude="123.61833" pageId="2" pageNumber="363" specimenCount="5" specimenCount-juvenile="5" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FDB3082EFD368E56" box="[593,698,596,617]" pageId="2" pageNumber="363" type="juvenile">5 juveniles</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FD02082EFF6C8EB9" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.46" metricValueMin="0.84" pageId="2" pageNumber="363" unit="mm" value="11.5" valueMax="14.6" valueMin="8.4">8.4- 14.6 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FF16080BFEB08EB9" box="[244,316,625,646]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="363">NTOU</collectionCode>
), collection data same as the holotype
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FD59080BFC8A8EB9" box="[699,774,625,646]" pageId="2" pageNumber="363" type="male">8 males</specimenCount>
[cl 
<quantity id="175F9B09FFEAFFA3FF4508F7FEBC8E9D" box="[167,304,653,674]" metricMagnitude="-2" metricUnit="m" metricValue="2.235" metricValueMax="2.85" metricValueMin="1.6199999999999999" pageId="2" pageNumber="363" unit="mm" value="22.35" valueMax="28.5" valueMin="16.2">16.2-28.5 mm</quantity>
, 1 (cl 
<quantity id="175F9B09FFEAFFA3FE8808F7FE4C8E9D" box="[362,448,653,674]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="2" pageNumber="363" unit="mm" value="21.6">21.6 mm</quantity>
) for DNA research], 
<materialsCitation id="60CF3CB1FFEAFFA3FD6908F7FD7D8EE3" collectingDate="2005-05-26" collectionCode="NTOU" country="Philippines" elevation="440" latitude="8.831667" longLatPrecision="129" longitude="123.581665" pageId="2" pageNumber="363" specimenCount="8" specimenCount-female="8" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FD6908F7FD648E9D" box="[651,744,653,674]" pageId="2" pageNumber="363" type="female">8 females</specimenCount>
[cl 
<quantity id="175F9B09FFEAFFA3FF6A08D0FE988E80" box="[136,276,682,703]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="3.41" metricValueMin="2.02" pageId="2" pageNumber="363" unit="mm" value="27.15" valueMax="34.1" valueMin="20.2">20.2-34.1 mm</quantity>
, 1 (cl 
<quantity id="175F9B09FFEAFFA3FEBA08D0FE3D8E80" box="[344,433,682,703]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="2" pageNumber="363" unit="mm" value="29.1">29.1 mm</quantity>
) for DNA research] (
<collectionCode id="B6B6AE29FFEAFFA3FD6808D0FD5E8E80" box="[650,722,682,703]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="363">NTOU</collectionCode>
), stn. CP2359, 
<geoCoordinate id="B593502BFFEAFFA3FF0308BDFEB58EE3" box="[225,313,710,732]" degrees="8" direction="north" minutes="49.9" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="8.831667">
8 
<emphasis id="E2D3EAFEFFEAFFA3FF0F08BCFF7B8EE4" bold="true" box="[237,247,710,731]" pageId="2" pageNumber="363">°</emphasis>
49.9’N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FEDD08BDFE208EE3" box="[319,428,710,732]" degrees="123" direction="east" minutes="34.9" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.581665">
123 
<emphasis id="E2D3EAFEFFEAFFA3FE8108BCFEE18EE4" bold="true" box="[355,365,710,731]" pageId="2" pageNumber="363">°</emphasis>
34.9'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FE5408BDFDAD8EE3" box="[438,545,711,732]" metricMagnitude="2" metricUnit="m" metricValue="4.4" metricValueMax="4.43" metricValueMin="4.37" pageId="2" pageNumber="363" unit="m" value="440.0" valueMax="443.0" valueMin="437.0">
<elevation id="5B8AD1DFFFEAFFA3FE5408BDFDAD8EE3" box="[438,545,711,732]" metricMagnitude="2" metricUnit="m" metricValue="4.4" metricValueMax="4.43" metricValueMin="4.37" pageId="2" pageNumber="363" unit="m" value="440.0" valueMax="443.0" valueMin="437.0">437-443 m</elevation>
</quantity>
, sandy, 
<date id="A419102CFFEAFFA3FD8C08BDFD7D8EE3" box="[622,753,711,732]" pageId="2" pageNumber="363" value="2005-05-26">
<collectingDate id="B45DE9C4FFEAFFA3FD8C08BDFD7D8EE3" box="[622,753,711,732]" pageId="2" pageNumber="363" value="2005-05-26">26 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FD1808BDFCF78CE9" box="[762,891,193,732]" pageId="2" pageNumber="363" type="male">4 males</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FC7D0ABBFBA58CE9" box="[927,1065,193,214]" metricMagnitude="-2" metricUnit="m" metricValue="2.395" metricValueMax="2.67" metricValueMin="2.12" pageId="2" pageNumber="363" unit="mm" value="23.95" valueMax="26.7" valueMin="21.2">21.2-26.7 mm</quantity>
), 
<materialsCitation id="60CF3CB1FFEAFFA3FBDA0ABBFB7B8D30" collectingDate="2005-05-26" collectionCode="MNHN" country="Philippines" elevation="361" latitude="8.815" longLatPrecision="129" longitude="123.62666" pageId="2" pageNumber="363" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FBDA0ABBFB198CE9" box="[1080,1173,193,214]" pageId="2" pageNumber="363" type="female">7 females</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FB5A0ABBFACE8CE9" box="[1208,1346,193,214]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" metricValueMax="2.9899999999999998" metricValueMin="2.05" pageId="2" pageNumber="363" unit="mm" value="25.2" valueMax="29.9" valueMin="20.5">20.5-29.9 mm</quantity>
), 2 ovigerous females (cl 26.9, 
<quantity id="175F9B09FFEAFFA3FC1F0AA7FBD78CCD" box="[1021,1115,221,242]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="2" pageNumber="363" unit="mm" value="27.4">27.4 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FB910AA7FB488CCD" box="[1139,1220,221,242]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="363">MNHN</collectionCode>
), stn. CP2360, 
<geoCoordinate id="B593502BFFEAFFA3FA8A0AA7FA4E8CCD" box="[1384,1474,221,242]" degrees="8" direction="north" minutes="48.9" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="8.815">
8 
<emphasis id="E2D3EAFEFFEAFFA3FA970AA7FAF38CCD" bold="true" box="[1397,1407,221,242]" pageId="2" pageNumber="363">°</emphasis>
48.9'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FCA10A80FC3C8D30" box="[835,944,250,271]" degrees="123" direction="east" minutes="37.6" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.62666">
123 
<emphasis id="E2D3EAFEFFEAFFA3FC850A80FCFD8D30" bold="true" box="[871,881,250,271]" pageId="2" pageNumber="363">°</emphasis>
37.6'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FC580A80FBA98D30" box="[954,1061,250,271]" metricMagnitude="2" metricUnit="m" metricValue="3.6049999999999995" metricValueMax="3.6399999999999997" metricValueMin="3.5700000000000003" pageId="2" pageNumber="363" unit="m" value="360.5" valueMax="364.0" valueMin="357.0">
<elevation id="5B8AD1DFFFEAFFA3FC580A80FBA98D30" box="[954,1061,250,271]" metricMagnitude="2" metricUnit="m" metricValue="3.6049999999999995" metricValueMax="3.6399999999999997" metricValueMin="3.5700000000000003" pageId="2" pageNumber="363" unit="m" value="360.5" valueMax="364.0" valueMin="357.0">357-364 m</elevation>
</quantity>
, sandy, 
<date id="A419102CFFEAFFA3FB960A80FB7B8D30" box="[1140,1271,250,271]" pageId="2" pageNumber="363" value="2005-05-26">
<collectingDate id="B45DE9C4FFEAFFA3FB960A80FB7B8D30" box="[1140,1271,250,271]" pageId="2" pageNumber="363" value="2005-05-26">26 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="60CF3CB1FFEAFFA3FAE30A80FBB48D76" collectingDate="2005-05-26" collectionCode="MNHN" country="Philippines" elevation="682" latitude="8.941667" longLatPrecision="129" longitude="123.545" pageId="2" pageNumber="363" specimenCount="10" specimenCount-juvenile="10" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FAE30A80FAFA8D30" box="[1281,1398,250,271]" pageId="2" pageNumber="363" type="juvenile">10 juveniles</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FA7E0A80FC118D13" metricMagnitude="-2" metricUnit="m" metricValue="1.02" metricValueMax="1.19" metricValueMin="0.85" pageId="2" pageNumber="363" unit="mm" value="10.2" valueMax="11.9" valueMin="8.5">8.5- 11.9 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FC500B6DFB8C8D13" box="[946,1024,279,300]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="363">MNHN</collectionCode>
), stn. CP2362, 
<geoCoordinate id="B593502BFFEAFFA3FB780B6DFB7C8D13" box="[1178,1264,278,300]" degrees="8" direction="north" minutes="56.5" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="8.941667">
8 
<emphasis id="E2D3EAFEFFEAFFA3FB450B6CFB3D8D14" bold="true" box="[1191,1201,278,299]" pageId="2" pageNumber="363">°</emphasis>
56.5'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FB1B0B6DFAE98D13" box="[1273,1381,278,300]" degrees="123" direction="east" minutes="32.7" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.545">
123 
<emphasis id="E2D3EAFEFFEAFFA3FAFE0B6CFAAA8D14" bold="true" box="[1308,1318,278,299]" pageId="2" pageNumber="363">°</emphasis>
32.7'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FA900B6DFCD48D76" metricMagnitude="2" metricUnit="m" metricValue="6.815" metricValueMax="6.84" metricValueMin="6.79" pageId="2" pageNumber="363" unit="m" value="681.5" valueMax="684.0" valueMin="679.0">
<elevation id="5B8AD1DFFFEAFFA3FA900B6DFCD48D76" metricMagnitude="2" metricUnit="m" metricValue="6.815" metricValueMax="6.84" metricValueMin="6.79" pageId="2" pageNumber="363" unit="m" value="681.5" valueMax="684.0" valueMin="679.0">679-684 m</elevation>
</quantity>
, sandy, 
<date id="A419102CFFEAFFA3FC4D0B4EFBB48D76" box="[943,1080,308,329]" pageId="2" pageNumber="363" value="2005-05-26">
<collectingDate id="B45DE9C4FFEAFFA3FC4D0B4EFBB48D76" box="[943,1080,308,329]" pageId="2" pageNumber="363" value="2005-05-26">26 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="C6A1FD65FFEAFFA3FBA70B4EFB188D76" box="[1093,1172,308,329]" pageId="2" pageNumber="363" type="male">3 males</specimenCount>
(cl 18.3, 18.5, 
<quantity id="175F9B09FFEAFFA3FAD50B4EFA198D76" box="[1335,1429,308,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="2" pageNumber="363" unit="mm" value="20.4">20.4 mm</quantity>
), 
<materialsCitation id="60CF3CB1FFEAFFA3FA4B0B4EFA8D8DBD" collectingDate="2005-05-30" collectionCode="NTOU" country="Philippines" elevation="783" latitude="9.446667" longLatPrecision="129" longitude="123.60167" pageId="2" pageNumber="363" specimenCount="10" specimenCount-juvenile="10" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FA4B0B4EFC2D8D59" pageId="2" pageNumber="363" type="juvenile">10 juveniles</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FC330B2BFBD68D59" box="[977,1114,337,358]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="11.6" metricValueMin="7.9" pageId="2" pageNumber="363" unit="mm" value="9.75" valueMax="11.6" valueMin="7.9">7.9-11.6 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FB900B2BFB328D59" box="[1138,1214,337,358]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="2" pageNumber="363">NTOU</collectionCode>
), stn. CP2389, 
<geoCoordinate id="B593502BFFEAFFA3FA840B2BFA4D8D59" box="[1382,1473,336,358]" degrees="9" direction="north" minutes="26.8" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.446667">
9 
<emphasis id="E2D3EAFEFFEAFFA3FA910B2AFAF18D5A" bold="true" box="[1395,1405,336,357]" pageId="2" pageNumber="363">°</emphasis>
26.8'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FCA10B17FC238DBD" box="[835,943,365,386]" degrees="123" direction="east" minutes="36.1" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.60167">
123 
<emphasis id="E2D3EAFEFFEAFFA3FC850B17FCFD8DBD" bold="true" box="[871,881,365,386]" pageId="2" pageNumber="363">°</emphasis>
36.1'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FC5E0B17FBA58DBD" box="[956,1065,365,386]" metricMagnitude="2" metricUnit="m" metricValue="7.83" metricValueMax="7.859999999999999" metricValueMin="7.8" pageId="2" pageNumber="363" unit="m" value="783.0" valueMax="786.0" valueMin="780.0">
<elevation id="5B8AD1DFFFEAFFA3FC5E0B17FBA58DBD" box="[956,1065,365,386]" metricMagnitude="2" metricUnit="m" metricValue="7.83" metricValueMax="7.859999999999999" metricValueMin="7.8" pageId="2" pageNumber="363" unit="m" value="783.0" valueMax="786.0" valueMin="780.0">780-786 m</elevation>
</quantity>
, sandy, 
<date id="A419102CFFEAFFA3FB990B17FA8D8DBD" box="[1147,1281,365,386]" pageId="2" pageNumber="363" value="2005-05-30">
<collectingDate id="B45DE9C4FFEAFFA3FB990B17FA8D8DBD" box="[1147,1281,365,386]" pageId="2" pageNumber="363" value="2005-05-30">30 May.2005</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="60CF3CB1FFEAFFA3FAEE0B17FB9B8D83" collectingDate="2005-05-30" collectionCode="ZRC" country="Philippines" elevation="595" latitude="9.498333" longLatPrecision="129" longitude="123.69334" pageId="2" pageNumber="363" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="C6A1FD65FFEAFFA3FAEE0B17FAE28DBD" box="[1292,1390,365,386]" pageId="2" pageNumber="363" type="juvenile">1 juvenile</specimenCount>
(cl 
<quantity id="175F9B09FFEAFFA3FA7A0B17FCE68DA0" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="363" unit="mm" value="11.7">11.7 mm</quantity>
) (
<collectionCode id="B6B6AE29FFEAFFA3FC9E0BF0FC238DA0" box="[892,943,394,415]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="363">ZRC</collectionCode>
), stn. CP2394, 
<geoCoordinate id="B593502BFFEAFFA3FBDD0BF0FB198DA0" box="[1087,1173,394,415]" degrees="9" direction="north" minutes="29.9" orientation="latitude" pageId="2" pageNumber="363" precision="92" value="9.498333">
9 
<emphasis id="E2D3EAFEFFEAFFA3FBA90BF0FBD98DA0" bold="true" box="[1099,1109,394,415]" pageId="2" pageNumber="363">°</emphasis>
29.9'N
</geoCoordinate>
<geoCoordinate id="B593502BFFEAFFA3FB780BF0FA8B8DA0" box="[1178,1287,394,415]" degrees="123" direction="east" minutes="41.6" orientation="longitude" pageId="2" pageNumber="363" precision="92" value="123.69334">
123 
<emphasis id="E2D3EAFEFFEAFFA3FB5C0BF0FB448DA0" bold="true" box="[1214,1224,394,415]" pageId="2" pageNumber="363">°</emphasis>
41.6'E
</geoCoordinate>
, 
<quantity id="175F9B09FFEAFFA3FAF20BF0FAF68DA0" box="[1296,1402,394,415]" metricMagnitude="2" metricUnit="m" metricValue="5.95" metricValueMax="7.87" metricValueMin="4.029999999999999" pageId="2" pageNumber="363" unit="m" value="595.0" valueMax="787.0" valueMin="403.0">
<elevation id="5B8AD1DFFFEAFFA3FAF20BF0FAF68DA0" box="[1296,1402,394,415]" metricMagnitude="2" metricUnit="m" metricValue="5.95" metricValueMax="7.87" metricValueMin="4.029999999999999" pageId="2" pageNumber="363" unit="m" value="595.0" valueMax="787.0" valueMin="403.0">403-787 m</elevation>
</quantity>
, sandy/ muddy, 
<date id="A419102CFFEAFFA3FC710BDDFB9B8D83" box="[915,1047,423,444]" pageId="2" pageNumber="363" value="2005-05-30">
<collectingDate id="B45DE9C4FFEAFFA3FC710BDDFB9B8D83" box="[915,1047,423,444]" pageId="2" pageNumber="363" value="2005-05-30">30 May.2005</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="98BD6567FFEAFFA5FCA10B9EFF638F87" lastPageId="4" lastPageNumber="365" pageId="2" pageNumber="363" type="diagnosis">
<paragraph id="D01836ECFFEAFFA5FCA10B9EFF638F87" blockId="2.[835,1474,484,731]" lastBlockId="4.[136,775,193,952]" lastPageId="4" lastPageNumber="365" pageId="2" pageNumber="363">
<emphasis id="E2D3EAFEFFEAFFA3FCA10B9EFC468DC3" bold="true" box="[835,970,484,508]" pageId="2" pageNumber="363">
<emphasis id="E2D3EAFEFFEAFFA3FCA10B9EFC3B8DC3" bold="true" box="[835,951,484,508]" italics="true" pageId="2" pageNumber="363">Diagnosis.</emphasis>
–
</emphasis>
Rostrum far overreaching scaphocerite, curved upwards above antennular peduncle, distal part anterior to curvation straight, dorsoanteral pointed, dorsally armed with 8 or 9 teeth including 4-5 teeth on carapace posterior to orbital margin, distal 0.33-0.25 unarmed, ventrally armed with 13- 16 teeth along entire length, tooth size progressively reduced distally, subapical ones very small; carapace with postorbital longitudinal carina rounded, extending posteriorly almost to posterior margin of carapace, curved ventrally at middle of carina length, branchiostegal carina extends posteriorly to 72- 78% of carapace length, without intermediate carina, branchiostegal spine not overreaching antennal spine; abdomen unarmed posteriomedially on all somites, third somite with blunt longitudinal dorsomedian carina, of which 2.85-3.40 times longer than the maximum width, with lateral margins slightly convex; pleura of 4th and 5th somites with acute posteroventral tooth, telson with 4 pairs of dorsolateral and 3 pairs of posterior spines; stylocerite sharply acute, extending to 0.7-1.0 of second segment length; scaphocerite 0.52-0.57 times as long as carapace, 2.80-3.35 times as long as width, blade slightly overreaching distolateral spine; third maxilliped with short (less than 0.23 length of antipenultimate segment), apex usually nude exopod and strap-like epipod; anterior 4 pereiopods with strap-like epipods; second pereiopods markedly unequal, right one shorter and stouter, with 8 articles, left one longer and more slender, with 23 or 24 articles; third pereiopod overreaching scaphocerite by dactylus, propodus and distal 0.1-0.6 carpus; dactylus 0.36- 0.39 times as long as propodus, accessory distal spine 0.36- 0.52 times as long as main terminal spine, closely abutting main terminal spine; maximum carapace length more than 
<quantity id="175F9B09FFECFFA5FF6A09DBFF678F87" box="[136,235,929,952]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="4" pageNumber="365" unit="mm" value="35.5">35.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="84D86664FFEAFFA3FF6A0D8FFDCC847C" ID-DOI="http://doi.org/10.5281/zenodo.4508028" ID-Zenodo-Dep="4508028" httpUri="https://zenodo.org/record/4508028/files/figure.png" pageId="2" pageNumber="363" startId="2.[136,171,2037,2058]" targetBox="[137,1458,856,1943]" targetPageId="2">
<paragraph id="D01836ECFFEAFFA3FF6A0D8FFDCC847C" blockId="2.[136,1475,2037,2115]" pageId="2" pageNumber="363">
Fig. 2. 
<taxonomicName id="17A74D6FFFEAFFA3FF300D8FFE1D8435" authority="Li, 2006" authorityName="Li" authorityYear="2006" box="[210,401,2037,2058]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="363" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="E2D3EAFEFFEAFFA3FF300D8FFE1D8435" box="[210,401,2037,2058]" italics="true" pageId="2" pageNumber="363">Heterocarpus chani</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="F9E05785FFEAFFA3FE7C0D8FFD998435" box="[414,533,2037,2058]" pageId="2" pageNumber="363" rank="species">new species</taxonomicNameLabel>
: a, eye and anterior carapace, left lateral view; b, body, left lateral view; c, rostrum and carapace, left lateral view. Female, holotype (cl 24.4 mm), Philippines: a; male, paratype (cl 18.5 mm), South China Sea: b; male, paratype (cl 19.6 mm), South China Sea: c. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="84D86664FFEBFFA2FF930D89FD4B847D" ID-DOI="http://doi.org/10.5281/zenodo.4508030" ID-Zenodo-Dep="4508030" httpUri="https://zenodo.org/record/4508030/files/figure.png" pageId="3" pageNumber="364" startId="3.[113,148,2035,2056]" targetBox="[268,1294,242,1966]" targetPageId="3">
<paragraph id="D01836ECFFEBFFA2FF930D89FD4B847D" blockId="3.[113,1451,2035,2114]" pageId="3" pageNumber="364">
Fig. 3. 
<taxonomicName id="17A74D6FFFEBFFA2FF5F0D8EFEF18436" authority="Li, 2006" authorityName="Li" authorityYear="2006" box="[189,381,2036,2057]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="E2D3EAFEFFEBFFA2FF5F0D8EFEF18436" box="[189,381,2036,2057]" italics="true" pageId="3" pageNumber="364">Heterocarpus chani</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="F9E05785FFEBFFA2FE680D89FD8E8437" box="[394,514,2035,2056]" pageId="3" pageNumber="364" rank="species">new species</taxonomicNameLabel>
: a, third maxilliped; b, same, antipenultimate segment and exopod; c, chela of first pereiopod; d, right second pereiopod; e, dactylus of third pereiopod. Male, paratype (cl 18.5 mm), South China Sea: a-d; Male, paratype (cl 19.6 mm), South China Sea: e. Scale bars: a, d, e = 5 mm; b, c = 1 mm.
</paragraph>
</caption>
<subSubSection id="98BD6567FFECFFA4FF6A099BFCE38E87" lastPageId="5" lastPageNumber="366" pageId="4" pageNumber="365" type="description">
<paragraph id="D01836ECFFECFFA5FF6A099BFC8B8987" blockId="4.[136,775,992,1464]" pageId="4" pageNumber="365">
<emphasis id="E2D3EAFEFFECFFA5FF6A099BFEA88FC7" bold="true" box="[136,292,992,1017]" pageId="4" pageNumber="365">
<emphasis id="E2D3EAFEFFECFFA5FF6A099BFE828FC6" bold="true" box="[136,270,993,1017]" italics="true" pageId="4" pageNumber="365">Description.</emphasis>
–
</emphasis>
Rostrum far overreaching scaphocerite, 0.8 to 2.0 times as long as carapace, rostrum length shorter in larger individuals, curving upwards above antennular peduncle, becoming straight and directed anterodorsally; lateral carina gradually sharpened posterior to curvation but becoming round and indistinct anterior to curvation, postrostral carina extending to almost posterior margin of carapace; dorsally armed with 8-9 teeth including 4-5 teeth on carapace posterior to orbit, posteriormost tooth on carapace situated at proximal 0.37-0.45 of carapace length, with distinct basal suture, 0.25-0.33 of dorsal rostral margin unarmed; ventrally armed with 13-16 acute teeth along entire length, tooth size progressively larger from posteriormost tooth to 4th or 5th tooth, but reducing in size from 4th or 5th tooth to distal tooth, distal 2 or 3 teeth very small and obscure.
</paragraph>
<paragraph id="D01836ECFFECFFA5FF6A0F9BFE318AE7" blockId="4.[136,775,1505,1752]" pageId="4" pageNumber="365">Carapace with dorsal profile distinctly convex; postorbital longitudinal carina rounded, extends posteriorly almost to posterior margin of carapace, curved ventrally at middle of carina length; branchiostegal carina sharp on anterior part, becoming broadly rounded posteriorly, extending posteriorly to 72-78% of lower carapace length (at level of branchiostegal carina); without intermediate carina; branchiostegal spine not overreaching antennal spine.</paragraph>
<paragraph id="D01836ECFFECFFA5FF6A0D7BFB538DA7" blockId="4.[136,775,1793,2104]" lastBlockId="4.[835,1474,193,408]" pageId="4" pageNumber="365">
Abdomen unarmed posteriomedially on all somites, without sharp dorsal carina or tooth on any somite; third somite with blunt longitudinal dorsomedian carina, which disappearing on anterior 0.10-0.12 and posterior 0.09-0.14 of somite length, carina 2.85-3.40 times longer than maximum width and with lateral margins slightly convex. Fourth and 5th pleura with acute posteroventral tooth. Sixth somite 1.40-1.60 times as long as maximum height, armed with acute posteroventral tooth, posterolateral lobe elongate posteriorly, sharply spiniform. Telson 0.56-0.58 times as long as carapace length, 1.55-1.65 longer than 6th somite, usually with 4 pairs of dorsolateral spines at around 0.38, 0.56, 0.75 and 0.86 of telson length (
<typeStatus id="0F1C884EFFECFFA5FC020B7BFBB38D27" box="[992,1087,257,280]" pageId="4" pageNumber="365" type="holotype">holotype</typeStatus>
with only 3 dorsolateral spines on left side); posterior margin generally with 3 pairs of spines, lateral pair similar to dorsolateral spines, intermediate pair longest, about 0.11-0.13 as long as telson length, middle pair about half length of intermediate pair.
</paragraph>
<paragraph id="D01836ECFFECFFA5FCA10BBBFB148E27" blockId="4.[835,1474,449,536]" pageId="4" pageNumber="365">Eye subpyriform, maximum diameter 0.15-0.17 times carapace length; ocellus absent but cornea slightly protruded at position of supposed ocellus.</paragraph>
<paragraph id="D01836ECFFECFFA5FCA1083BFB328E87" blockId="4.[835,1474,577,696]" pageId="4" pageNumber="365">Antennular peduncle extending to proximal 0.60-0.70 of scaphocerite; stylocerite sharply acute, reaching proximal 0.7 or distal end of second segment; outer flagellum very long, more than 3 times carapace length.</paragraph>
<paragraph id="D01836ECFFECFFA5FCA1089BFBB88F67" blockId="4.[835,1474,737,856]" pageId="4" pageNumber="365">Scaphocerite 0.52-0.57 times as long as carapace, 2.80-3.35 times as long as wide, blade slightly overreaching distolateral spine, lateral margin feebly convex; flagellum more than 5.6 times carapace length.</paragraph>
<paragraph id="D01836ECFFECFFA5FCA109FBFA848807" blockId="4.[835,1474,897,1080]" pageId="4" pageNumber="365">Third maxilliped stout, overreaching scaphocerite by at least distal 0.33 of distal segment, penultimate segment 0.22-0.25 as long as carapace, 0.63-0.73 as long as distal segment, 0.39- 0.44 as long as antepenultimate segment; exopod reduced but distinct, short, apex nude, at most 0.23 of antipenultimate segment length; strap-like epipod present.</paragraph>
<paragraph id="D01836ECFFECFFA4FCA10E1BFD7D8827" blockId="4.[835,1474,1121,1560]" lastBlockId="5.[113,753,193,1048]" lastPageId="5" lastPageNumber="366" pageId="4" pageNumber="365">Pereiopods with strap-like epipods on 4 anterior pairs. First pereiopod slightly overreaching scaphocerite; fingers small but distinct; chela 0.23-0.27 times as long as carapace, 0.64- 0.78 times as long as carpus. Second pereiopods markedly unequal, right one shorter and stouter, overreaching carpocerite by fingers or by entire chela; chela 0.23-0.24 times as long as carapace, fingers 0.68-0.82 times as long as palm; carpus 1.30-1.42 times as long as chela, subdivided into 8 articles, proximal article longest, distal article second longest, others articles similar and short; merus 0.97-1.04 times as long as chela, ischium compressed and produced ventrally, 1.34-1.51 times as long as chela; left second pereiopod overreaching scaphocerite by chela and distal 0.37-0.53 carpus, chela 0.10-0.13 times as long as carpus, fingers 0.75- 1.01 times as long palm; carpus 0.73-0.77 times as long as carapace, subdivided into 23 or 24 articles; merus 0.48-0.58 times as long as carpus, with 8-9 articles; ischium 0.49-0.64 times as long as carpus, with longitudinal carina along outer surface, proximal half of ventral margin slightly swollen forming compressed lobe. Third pereiopod overreaching scaphocerite by dactylus, propodus and distal 0.1-0.6 carpus; dactylus 0.38-0.39 as long as propodus, bearing 5 spines along proximal 0.35 of flexor margin, accessory distal spine 0.36- 0.52 of main terminal spine, and strongly abutting latter, main terminal spine 0.07-0.10 of total dactylus length; propodus 0.52-0.58 as long as carapace, scattered with long setae, ventral margin with 13-15 spinules, distoventral spine accompanying with row of setae, distodorsal corner also with row of setae; carpus 0.73-0.77 as long as propodus, with 1- 3 strong spines and 4-10 spinules on flexor margin; merus 1.54-1.58 as long as propodus, armed with 4-7 mesial and 8- 20 lateral spines along flexor margin; ischium 0.34-0.39 as long as propodus, with 2 (seldom 3) spines on ventral margin. Fourth and fifth pereiopods similar to third, fourth overreaching scaphocerite by dactylus, propodus, and at most distal 0.4 carpus, carpus armed with 1 (seldom 2) spine(s), merus with 1 subdistomesial and 9-21 (avg 12) ventral spines, ischium with 2 ventral spines; fifth overreaching scaphocerite by dactylus and propodus, carpus with 0-2 (usually 1) spine, merus with 7-16 (avg 9) ventral spines, without distomesial spine; ischium unarmed or at most bearing 1 ventral tooth.</paragraph>
<caption id="84D86664FFECFFA5FCA10268FBE1847B" ID-DOI="http://doi.org/10.5281/zenodo.4508032" ID-Zenodo-Dep="4508032" httpUri="https://zenodo.org/record/4508032/files/figure.png" pageId="4" pageNumber="365" startId="4.[835,870,2066,2087]" targetBox="[835,1475,1623,2048]" targetPageId="4">
<paragraph id="D01836ECFFECFFA5FCA10268FBE1847B" blockId="4.[835,1474,2066,2116]" pageId="4" pageNumber="365">
Fig. 4. 
<taxonomicName id="17A74D6FFFECFFA5FC650268FBCA8418" authority="Li, 2006" authorityName="Li" authorityYear="2006" box="[903,1094,2066,2087]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="E2D3EAFEFFECFFA5FC650268FBCA8418" box="[903,1094,2066,2087]" italics="true" pageId="4" pageNumber="365">Heterocarpus chani</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="F9E05785FFECFFA5FBB00268FB448418" box="[1106,1224,2066,2087]" pageId="4" pageNumber="365" rank="species">new species</taxonomicNameLabel>
, female, holotype (cl 24.4 mm), Philippines, color photo.
</paragraph>
</caption>
<paragraph id="D01836ECFFEDFFA4FF930E3BFB948D67" blockId="5.[113,752,1089,1272]" lastBlockId="5.[812,1451,193,344]" pageId="5" pageNumber="366">Endopod of first pleopod in males broadened at distal half, mesial margin sinuous, with short setae along proximal 0.75 of endopod length; lateral margin slightly sinuous, with long plumose and simple setae; distal margin with distinct sinus, resulting in endopod somewhat bifurca, mesial lobe smaller, with cluster of small hooked protuberances. Appendix masculine on second pleopod subequal to appendix interna, armed with 16-18 long spines on anteromesial and distal margins. Uropod distinctly overreaching telson, exopod bearing fixed distolateral spine that accompanied with strong movable spine inside.</paragraph>
<paragraph id="D01836ECFFEDFFA4FCCE0BFBFCE38E87" blockId="5.[812,1452,384,696]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE0BFBFC328DA7" bold="true" box="[812,958,384,409]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE0BFBFC258DA6" bold="true" box="[812,937,385,409]" italics="true" pageId="5" pageNumber="366">Coloration.</emphasis>
–
</emphasis>
Body orange-red to reddish, with dorsal parts proportionally more orange-red while ventral parts including pleopods and tailfan generally reddish. Eye black-brown with golden reflections. Flagella reddish. Mouth parts and anterior 2 pereiopods varied from reddish to pinkish. Posterior 3 pereiopods with dactyli and distal portion of propodi always reddish, carpi and distal portion of meri as well as proximal portion of propodus always pinkish to light pink, basal segment to medial part of meri reddish or pinkish. Eggs dark green.
</paragraph>
</subSubSection>
<subSubSection id="98BD6567FFEDFFA4FCCE089BFBF58F67" pageId="5" pageNumber="366" type="etymology">
<paragraph id="D01836ECFFEDFFA4FCCE089BFBF58F67" blockId="5.[812,1451,736,856]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE089BFC308EC7" bold="true" box="[812,956,736,761]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE089BFC248EC6" bold="true" box="[812,936,737,761]" italics="true" pageId="5" pageNumber="366">Etymology.</emphasis>
–
</emphasis>
It is a pleasure to name this species after Tin- Yam Chan, carcinologist of the National 
<collectingCountry id="A8B0767CFFEDFFA4FAEB097BFAD78F27" box="[1289,1371,769,792]" name="Taiwan" pageId="5" pageNumber="366">Taiwan</collectingCountry>
Ocean University, who has had a series of works on the pandalid shrimps from the Indo-Pacific.
</paragraph>
</subSubSection>
<subSubSection id="98BD6567FFEDFFA4FCCE09FBFAA18FE7" pageId="5" pageNumber="366" type="distribution">
<paragraph id="D01836ECFFEDFFA4FCCE09FBFAA18FE7" blockId="5.[812,1451,897,984]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE09FBFC4B8FA6" bold="true" box="[812,967,897,921]" italics="true" pageId="5" pageNumber="366">Distribution. –</emphasis>
This species is known with certainty only from the Nansha Islands, southern South 
<collectingCountry id="A8B0767CFFEDFFA4FB4009DBFB6D8F87" box="[1186,1249,929,952]" name="China" pageId="5" pageNumber="366">China</collectingCountry>
Sea, and the 
<collectingRegion id="1263F80EFFEDFFA4FA8809DBFA278F87" box="[1386,1451,929,952]" country="Philippines" name="Bohol" pageId="5" pageNumber="366">Bohol</collectingRegion>
and 
<collectingRegion id="1263F80EFFEDFFA4FCB909BBFC008FE7" box="[859,908,961,984]" country="Philippines" name="Sulu" pageId="5" pageNumber="366">Sulu</collectingRegion>
Seas, 
<collectingCountry id="A8B0767CFFEDFFA4FC3109BBFBC28FE7" box="[979,1102,961,984]" name="Philippines" pageId="5" pageNumber="366">Philippines</collectingCountry>
, at 
<quantity id="175F9B09FFEDFFA4FB9709BBFB678FE7" box="[1141,1259,961,984]" metricMagnitude="2" metricUnit="m" metricValue="6.35" metricValueMax="8.879999999999999" metricValueMin="3.82" pageId="5" pageNumber="366" unit="m" value="635.0" valueMax="888.0" valueMin="382.0">382-888 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="98BD6567FFEDFFA6FCCE0E7BFD6E8EE7" lastPageId="7" lastPageNumber="368" pageId="5" pageNumber="366" type="discussion">
<paragraph id="D01836ECFFEDFFA4FCCE0E7BFBB58967" blockId="5.[812,1452,1025,1368]" pageId="5" pageNumber="366">
<emphasis id="E2D3EAFEFFEDFFA4FCCE0E7BFC378826" bold="true" box="[812,955,1025,1049]" italics="true" pageId="5" pageNumber="366">Remarks. –</emphasis>
This new species is closely related to 
<taxonomicName id="17A74D6FFFEDFFA4FCCE0E5BFB248807" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[812,1192,1057,1080]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEDFFA4FCCE0E5BFBAB8807" box="[812,1063,1057,1080]" italics="true" pageId="5" pageNumber="366">Heterocarpus gibbosus</emphasis>
Bate, 1888
</taxonomicName>
, 
<taxonomicName id="17A74D6FFFEDFFA4FB570E5BFC748867" authority="Alcock &amp; Anderson, 1894" authorityName="Alcock &amp; Anderson" authorityYear="1894" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis id="E2D3EAFEFFEDFFA4FB570E5BFAD98807" box="[1205,1365,1057,1080]" italics="true" pageId="5" pageNumber="366">H. tricarinatus</emphasis>
Alcock &amp; Anderson, 1894
</taxonomicName>
, and 
<taxonomicName id="17A74D6FFFEDFFA4FBD10E3BFAC98867" authority="de Man, 1917" authorityName="De Man" authorityYear="1917" box="[1075,1349,1089,1112]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="E2D3EAFEFFEDFFA4FBD10E3BFB2E8867" box="[1075,1186,1089,1112]" italics="true" pageId="5" pageNumber="366">H. lepidus</emphasis>
de Man, 1917
</taxonomicName>
. Besides the exopod of the third maxilliped (rudimentary vs. welldeveloped), 
<taxonomicName id="17A74D6FFFEDFFA4FC520EFBFB8788A7" authorityName="Li" authorityYear="2006" box="[944,1035,1153,1176]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="chani">
<emphasis id="E2D3EAFEFFEDFFA4FC520EFBFB8788A7" box="[944,1035,1153,1176]" italics="true" pageId="5" pageNumber="366">H. chani</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="17A74D6FFFEDFFA4FACE0EFBFA2788A7" authorityName="Bate" authorityYear="1888" box="[1324,1451,1153,1176]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEDFFA4FACE0EFBFA2788A7" box="[1324,1451,1153,1176]" italics="true" pageId="5" pageNumber="366">H. gibbosus</emphasis>
</taxonomicName>
by having a broader dorsomedian carina on the third abdominal somite (0.29-0.34 vs. 0.15-0.20 as carina length); from 
<taxonomicName id="17A74D6FFFEDFFA4FC910E9BFBA188C7" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[883,1069,1249,1272]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis id="E2D3EAFEFFEDFFA4FC910E9BFBA188C7" box="[883,1069,1249,1272]" italics="true" pageId="5" pageNumber="366">H. tricarinatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="17A74D6FFFEDFFA4FB910E9BFB7888C7" authorityName="De Man" authorityYear="1917" box="[1139,1268,1249,1272]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="E2D3EAFEFFEDFFA4FB910E9BFB7888C7" box="[1139,1268,1249,1272]" italics="true" pageId="5" pageNumber="366">H. lepidus</emphasis>
</taxonomicName>
by the longer branchiostegal carina (about 70% lower carapace length vs. about 50% in 
<taxonomicName id="17A74D6FFFEDFFA4FC240F5BFBEA8907" authorityName="Alcock &amp; Anderson" authorityYear="1894" box="[966,1126,1313,1336]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis id="E2D3EAFEFFEDFFA4FC240F5BFBEA8907" box="[966,1126,1313,1336]" italics="true" pageId="5" pageNumber="366">H. tricarinatus</emphasis>
</taxonomicName>
and about 60% in 
<taxonomicName id="17A74D6FFFEDFFA4FAD70F5BFA288907" authorityName="De Man" authorityYear="1917" box="[1333,1444,1313,1336]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="lepidus">
<emphasis id="E2D3EAFEFFEDFFA4FAD70F5BFA288907" box="[1333,1444,1313,1336]" italics="true" pageId="5" pageNumber="366">H. lepidus</emphasis>
</taxonomicName>
, also see 
<bibRefCitation id="B4364B1DFFEDFFA4FC6B0F3BFBA18967" author="Crosnier, A" box="[905,1069,1345,1368]" pageId="5" pageNumber="366" pagination="57 - 103" refId="ref11646" refString="Crosnier, A., 1988. Sur les Heterocarpus (Crustacea, Decapoda, Pandalidae) du sud-ouest de l'ocean Indien Remarques sur d'autres especes ouest-pacifiques du genre et description de quatre taxa nouveaux. Bulletin du Museum national d'Histoire Naturelle, Paris 4 e ser., 10, section A, no. 1: 57 - 103, pls. 1 - 4." type="journal article" year="1988">Crosnier, 1988</bibRefCitation>
).
</paragraph>
<paragraph id="D01836ECFFEDFFA7FCCE0FFBFC5C8D47" blockId="5.[812,1452,1409,1528]" lastBlockId="6.[835,1474,193,376]" lastPageId="6" lastPageNumber="367" pageId="5" pageNumber="366">
There were many specimens of this species collected by the “Panglao 2005” expedition to the 
<collectingCountry id="A8B0767CFFEDFFA4FB7C0FDBFA9A8987" box="[1182,1302,1441,1464]" name="Philippines" pageId="5" pageNumber="366">Philippines</collectingCountry>
and there are color photographs of the “Panglao 2005” specimens. It seems that the coloration of 
<taxonomicName id="17A74D6FFFEDFFA4FBF20F9BFBE789C7" authorityName="Li" authorityYear="2006" box="[1040,1131,1505,1528]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="chani">
<emphasis id="E2D3EAFEFFEDFFA4FBF20F9BFBE789C7" box="[1040,1131,1505,1528]" italics="true" pageId="5" pageNumber="366">H. chani</emphasis>
</taxonomicName>
(
<figureCitation id="489C2A69FFEDFFA4FB9B0F9BFB3489C7" box="[1145,1208,1505,1528]" captionStart="Fig" captionStartId="4.[835,870,2066,2087]" captionTargetBox="[835,1475,1623,2048]" captionTargetId="figure-802@4.[835,1475,1623,2048]" captionTargetPageId="4" captionText="Fig. 4. Heterocarpus chani, new species, female, holotype (cl 24.4 mm), Philippines, color photo." figureDoi="http://doi.org/10.5281/zenodo.4508032" httpUri="https://zenodo.org/record/4508032/files/figure.png" pageId="5" pageNumber="366">Fig. 4</figureCitation>
) and 
<taxonomicName id="17A74D6FFFEDFFA4FB110F9BFAFE89C7" authorityName="Bate" authorityYear="1888" box="[1267,1394,1505,1528]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEDFFA4FB110F9BFAFE89C7" box="[1267,1394,1505,1528]" italics="true" pageId="5" pageNumber="366">H. gibbosus</emphasis>
</taxonomicName>
(
<figureCitation id="489C2A69FFEDFFA7FA620F9BFF1A8CE7" captionStart="Fig" captionStartId="5.[812,847,2007,2028]" captionTargetBox="[812,1452,1566,1991]" captionTargetId="figure-311@5.[812,1452,1566,1991]" captionTargetPageId="5" captionText="Fig. 6. Heterocarpus gibbossus Bate, 1888, ovigerous female (cl 30.7 mm), Philippines, Bohol Sea, Panglao 2005 Expedition, station CA2337, 9º31.5'N 123º41.7'E, 336 m depth, 22 May.2005, color photo." figureDoi="http://doi.org/10.5281/zenodo.4508036" httpUri="https://zenodo.org/record/4508036/files/figure.png" lastPageId="6" lastPageNumber="367" pageId="5" pageNumber="366">Fig. 6</figureCitation>
) are very similar, but there are consistant differences. 
<taxonomicName id="17A74D6FFFEEFFA7FF6A0A9BFEF38CC7" authorityName="Bate" authorityYear="1888" box="[136,383,225,248]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEEFFA7FF6A0A9BFEF38CC7" box="[136,383,225,248]" italics="true" pageId="6" pageNumber="367">Heterocarpus gibbosus</emphasis>
</taxonomicName>
has a distinct red ring at the subdistal part of the meri of the posterior three pereiopods. In 
<taxonomicName id="17A74D6FFFEEFFA7FD4E0B7BFC8A8D27" authorityName="Li" authorityYear="2006" box="[684,774,257,280]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="chani">
<emphasis id="E2D3EAFEFFEEFFA7FD4E0B7BFC8A8D27" box="[684,774,257,280]" italics="true" pageId="6" pageNumber="367">H. chani</emphasis>
</taxonomicName>
the posterior three pereiopods never show distinct red rings on the meri. Furthermore, in freshly preserved material of 
<taxonomicName id="17A74D6FFFEEFFA7FF6A0B1BFF6F8D47" authorityName="Li" authorityYear="2006" box="[136,227,353,376]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="chani">
<emphasis id="E2D3EAFEFFEEFFA7FF6A0B1BFF6F8D47" box="[136,227,353,376]" italics="true" pageId="6" pageNumber="367">H. chani</emphasis>
</taxonomicName>
, the dorsal margin of the basal rostral teeth and the dactyli of the posterior pereiopods are reddish. On the other hand, preserved specimens of 
<taxonomicName id="17A74D6FFFEEFFA7FE360BDBFDDB8D87" authorityName="Bate" authorityYear="1888" box="[468,599,417,440]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEEFFA7FE360BDBFDDB8D87" box="[468,599,417,440]" italics="true" pageId="6" pageNumber="367">H. gibbosus</emphasis>
</taxonomicName>
have the dorsal margin of the basal rostral teeth not red in color but with almost the entire posterior pereiopods being reddish. The third maxilliped exopod tends to be shorter in smaller specimens of 
<taxonomicName id="17A74D6FFFEEFFA7FECE085BFE448E07" box="[300,456,545,568]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="E2D3EAFEFFEEFFA7FECE085BFE448E07" box="[300,456,545,568]" italics="true" pageId="6" pageNumber="367">Heterocarpus</emphasis>
</taxonomicName>
. Nevertheless, of the 160 “Panglao 2005” specimens examined, there is a clear separation of the exopod/antipenultimate segment of third maxilliped ratio in the two forms for those with a carapace length longer than 
<quantity id="175F9B09FFEEFFA7FEAA08DBFE1C8E87" box="[328,400,673,696]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="367" unit="mm" value="15.0">15 mm</quantity>
(
<figureCitation id="489C2A69FFEEFFA7FE4008DBFE6D8E87" box="[418,481,673,696]" captionStart="Fig" captionStartId="5.[114,149,2036,2057]" captionTargetBox="[116,750,1334,2000]" captionTargetId="figure-654@5.[102,765,1308,2013]" captionTargetPageId="5" captionText="Fig. 5. Relationships between the postorbital carapace length and the ratio of exopod/antipenultimate segment of third maxilliped in Heterocarpus chani, new species, and H. gibbosus Bate, 1888." figureDoi="http://doi.org/10.5281/zenodo.4508038" httpUri="https://zenodo.org/record/4508038/files/figure.png" pageId="6" pageNumber="367">Fig. 5</figureCitation>
). In the 75 larger specimens (i.e. cl longer than 
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) of the short exopod form the exopod/antipenultimate segment of third maxilliped ratio is 0.13-0.23 (avg 0.18). On the other hand, the 45 larger specimens (cl&gt; 
<quantity id="175F9B09FFEEFFA7FED9095BFE058F07" box="[315,393,801,824]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="367" unit="mm" value="15.0">15 mm</quantity>
) of the long exopod form have the ratio ranging from 0.23-0.53 (avg 0.37) (fig. 7a). The separation of the two forms is very distinct for ovigerous females, with an average ratio of 0.20 for the short exopod form (n=7) and 0.42 for the long exopod form (n=18). In small juveniles (cl&lt;
<quantity id="175F9B09FFEEFFA7FBFE0B5BFBE78D07" box="[1052,1131,289,312]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="367" unit="mm" value="15.0">15 mm</quantity>
), there is still a slight difference in the average ratio (0.15 (0.09-0.20, n=26) vs. 0.17 (0.11- 0.24, n=13)).
</paragraph>
<caption id="84D86664FFEDFFA4FCCE0DADFCE6847C" ID-DOI="http://doi.org/10.5281/zenodo.4508036" ID-Zenodo-Dep="4508036" httpUri="https://zenodo.org/record/4508036/files/figure.png" pageId="5" pageNumber="366" startId="5.[812,847,2007,2028]" targetBox="[812,1452,1566,1991]" targetPageId="5">
<paragraph id="D01836ECFFEDFFA4FCCE0DADFCE6847C" blockId="5.[812,1451,2007,2115]" pageId="5" pageNumber="366">
Fig. 6. 
<taxonomicName id="17A74D6FFFEDFFA4FC970DADFB598BD3" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[885,1237,2007,2028]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gibbossus">
<emphasis id="E2D3EAFEFFEDFFA4FC970DADFBEC8BD3" box="[885,1120,2007,2028]" italics="true" pageId="5" pageNumber="366">Heterocarpus gibbossus</emphasis>
Bate, 1888
</taxonomicName>
, ovigerous female (cl 30.7 mm), Philippines, Bohol Sea, Panglao 2005 Expedition, station CA2337, 9º31.5'N 123º41.7'E, 336 m depth, 22 May.2005, color photo.
</paragraph>
</caption>
<caption id="84D86664FFEDFFA4FF900D8EFD5A847C" ID-DOI="http://doi.org/10.5281/zenodo.4508038" ID-Zenodo-Dep="4508038" httpUri="https://zenodo.org/record/4508038/files/figure.png" pageId="5" pageNumber="366" startId="5.[114,149,2036,2057]" targetBox="[116,750,1334,2000]" targetPageId="5">
<paragraph id="D01836ECFFEDFFA4FF900D8EFD5A847C" blockId="5.[114,753,2036,2115]" pageId="5" pageNumber="366">
Fig. 5. Relationships between the postorbital carapace length and the ratio of exopod/antipenultimate segment of third maxilliped in 
<taxonomicName id="17A74D6FFFEDFFA4FF900254FEBE847C" authority="Li, 2006" authorityName="Li" authorityYear="2006" box="[114,306,2094,2115]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="chani" status="sp. nov.">
<emphasis id="E2D3EAFEFFEDFFA4FF900254FEBE847C" box="[114,306,2094,2115]" italics="true" pageId="5" pageNumber="366">Heterocarpus chani</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="F9E05785FFEDFFA4FEA20254FE3B847C" box="[320,439,2094,2115]" pageId="5" pageNumber="366" rank="species">new species</taxonomicNameLabel>
, and 
<taxonomicName id="17A74D6FFFEDFFA4FE090254FD5E847C" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[491,722,2094,2115]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="366" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEDFFA4FE090254FDD3847C" box="[491,607,2094,2115]" italics="true" pageId="5" pageNumber="366">H. gibbosus</emphasis>
Bate, 1888
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="D01836ECFFEEFFA7FCA10BDBFB638E27" blockId="6.[835,1474,417,536]" pageId="6" pageNumber="367">
These two forms occur in similar depths (
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for the short exopod form and 
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for the long exopod form) and collected together in 6 stations (i.e. CP2332, CP 2333, CP2341, CP 2359, CP 2360, CP 2362).
</paragraph>
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<taxonomicName id="17A74D6FFFEEFFA7FCA1083BFBB08E67" authorityName="Bate" authorityYear="1888" box="[835,1084,577,600]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="367" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEEFFA7FCA1083BFBB08E67" box="[835,1084,577,600]" italics="true" pageId="6" pageNumber="367">Heterocarpus gibbosus</emphasis>
</taxonomicName>
is generally believed to have a long exopod at the third maxilliped (
<bibRefCitation id="B4364B1DFFEEFFA7FB63081BFA928E47" author="Crosnier, A" box="[1153,1310,609,632]" pageId="6" pageNumber="367" pagination="57 - 103" refId="ref11646" refString="Crosnier, A., 1988. Sur les Heterocarpus (Crustacea, Decapoda, Pandalidae) du sud-ouest de l'ocean Indien Remarques sur d'autres especes ouest-pacifiques du genre et description de quatre taxa nouveaux. Bulletin du Museum national d'Histoire Naturelle, Paris 4 e ser., 10, section A, no. 1: 57 - 103, pls. 1 - 4." type="journal article" year="1988">Crosnier, 1988</bibRefCitation>
; 
<bibRefCitation id="B4364B1DFFEEFFA7FAC5081BFA368E47" author="Tavares" box="[1319,1466,609,632]" pageId="6" pageNumber="367" pagination="671 - 677" refId="ref12411" refString="Tavares, Marcos, 1999. New species and new records of deep-water caridean shrimps from the South Atlantic Ocean (Crustacea, Decapoda). Zoosystema, 21 (4): 671 - 677, figs. 1 - 4." type="journal article" year="1999">Tavares, 1999</bibRefCitation>
) and widely distributed from eastern Africa to 
<collectingCountry id="A8B0767CFFEEFFA7FAB108FBFA1C8EA7" box="[1363,1424,641,664]" name="Japan" pageId="6" pageNumber="367">Japan</collectingCountry>
and 
<collectingCountry id="A8B0767CFFEEFFA7FCA108DBFC0A8E87" box="[835,902,673,696]" name="Tonga" pageId="6" pageNumber="367">Tonga</collectingCountry>
(
<bibRefCitation id="B4364B1DFFEEFFA7FC7008DBFB958E87" author="Chace, F" box="[914,1049,673,696]" pageId="6" pageNumber="367" pagination="1 - 143" refId="ref11314" refString="Chace, F. A., Jr., 1985. The Caridean Shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 3: Families Thalassocarididae and Pandalidae. Smithsonian Contributions to Zoology, 411: 1 - 143, Fig. 1 - 62." type="journal article" year="1985">Chace, 1985</bibRefCitation>
; Chan &amp; Yu, 1987). However, it appears that there are actually two distinct forms, at least in the specimens obtained from the 
<collectingCountry id="A8B0767CFFEEFFA7FB78089BFA908EC7" box="[1178,1308,737,760]" name="Philippines" pageId="6" pageNumber="367">Philippines</collectingCountry>
. The original description and figures given by Bate (1888) was based on a single juvenile type specimen (cl 
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, “Challenger” Expedition, station 207, off Tables Island, 
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, 
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12 
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21'N
</geoCoordinate>
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122 
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15'E
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, 700 fathoms, 
<date id="A419102CFFEFFFA6FE3D0ABBFDE98CE7" box="[479,613,193,216]" pageId="7" pageNumber="368" value="1875-01-16">16 Jan.1875</date>
) and has no information on the status of the exopod of the third maxilliped. The type has been re-examined (NHM 88.22, The Natural History Museum, London) and was found to be badly damaged, soft and decalcified (including the abdomen). In fact, both the third maxillipeds are missing (also see 
<bibRefCitation id="B4364B1DFFEFFFA6FD740B1BFF258DA7" author="Calman, W" pageId="7" pageNumber="368" pagination="183 - 224" refId="ref11276" refString="Calman, W. T., 1939. Crustacea: Caridea. The John Murray Expedition 1933 - 1934, Scientific Report 6 (4): 183 - 224, Figs. 1 - 8." type="journal article" year="1939">Calman, 1939</bibRefCitation>
; 
<bibRefCitation id="B4364B1DFFEFFFA6FF560BFBFEB18DA7" author="Chace, F" box="[180,317,385,408]" pageId="7" pageNumber="368" pagination="1 - 143" refId="ref11314" refString="Chace, F. A., Jr., 1985. The Caridean Shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 3: Families Thalassocarididae and Pandalidae. Smithsonian Contributions to Zoology, 411: 1 - 143, Fig. 1 - 62." type="journal article" year="1985">Chace, 1985</bibRefCitation>
). The 
<typeStatus id="0F1C884EFFEFFFA6FE640BFBFE6F8DA7" box="[390,483,385,408]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="17A74D6FFFEFFFA6FDE50BFBFD0A8DA7" authorityName="Bate" authorityYear="1888" box="[519,646,385,408]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEFFFA6FDE50BFBFD0A8DA7" box="[519,646,385,408]" italics="true" pageId="7" pageNumber="368">H. gibbosus</emphasis>
</taxonomicName>
is now of no use in deciding whether the short or long exopod form should bear Bate’s (1888) name. In order to fix the identity of 
<taxonomicName id="17A74D6FFFEFFFA6FF720B9BFE9D8DC7" authorityName="Bate" authorityYear="1888" box="[144,273,481,504]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEFFFA6FF720B9BFE9D8DC7" box="[144,273,481,504]" italics="true" pageId="7" pageNumber="368">H. gibbosus</emphasis>
</taxonomicName>
, an application to the International Code of Zoological Nomenclature has been made to replace the 
<typeStatus id="0F1C884EFFEFFFA6FF93085BFF428E07" box="[113,206,545,568]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
with a 
<typeStatus id="0F1C884EFFEFFFA6FEC1085BFEFB8E07" box="[291,375,545,568]" pageId="7" pageNumber="368" type="neotype">neotype</typeStatus>
selected from the “Panglao 2005” material with a long exopod on the third maxilliped (Li et al., in press). The short exopod form is here treated as a new species. Records of 
<taxonomicName id="17A74D6FFFEFFFA6FEB108FBFE588EA7" authorityName="Bate" authorityYear="1888" box="[339,468,641,664]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="368" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="E2D3EAFEFFEFFFA6FEB108FBFE588EA7" box="[339,468,641,664]" italics="true" pageId="7" pageNumber="368">H. gibbosus</emphasis>
</taxonomicName>
by previous workers (e.g. 
<bibRefCitation id="B4364B1DFFEFFFA6FF9308DBFF788E87" author="Chace, F" box="[113,244,673,696]" pageId="7" pageNumber="368" pagination="1 - 143" refId="ref11314" refString="Chace, F. A., Jr., 1985. The Caridean Shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 3: Families Thalassocarididae and Pandalidae. Smithsonian Contributions to Zoology, 411: 1 - 143, Fig. 1 - 62." type="journal article" year="1985">Chace, 1985</bibRefCitation>
) will need to be re-examined in order to establish the exact distribution of these two closely related species.
</paragraph>
</subSubSection>
</treatment>
</document>