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<document id="48CC903F1341B7735325F69984859D30" ID-CLB-Dataset="4250" ID-DOI="10.11646/palaeoentomology.4.1.6" ID-GBIF-Dataset="ab05a0bd-5d5c-4175-a2c8-61a10cf04d08" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507696" ID-ZooBank="0D058DAF-F3D8-4C8F-913C-55908F8E827D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631644204445" checkinUser="carolina" docAuthor="HEISS, ERNST &amp; GUILBERT, ERIC" docDate="2021" docId="4E015A1BFFBFFFE561F1F82EFC4FF7AA" docLanguage="en" docName="palaeoentomology.4.1.6.pdf" docOrigin="Palaeoentomology 4 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.1.6" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Burmacader grandis HEISS &amp; GUILBERT 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="B2382263FFBFFFE46179FF8BF92BFFAC" masterDocTitle="A new species of Burmacader Heiss &amp; Guilbert, 2013 from Cretaceous Burmese amber (Hemiptera, Heteroptera, Tingidae)" masterLastPageNumber="33" masterPageNumber="30" pageNumber="30" updateTime="1699448745859" updateUser="ExternalLinkService">
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<mods:title id="56D318A1B739F2B1A7D7E9A1AD6D86BB">A new species of Burmacader Heiss &amp; Guilbert, 2013 from Cretaceous Burmese amber (Hemiptera, Heteroptera, Tingidae)</mods:title>
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<mods:name id="97CC49273B33105EEE8C72785991DEAA" type="personal">
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<mods:namePart id="9DE6715861C839B3C68D9B2A9A57110A">HEISS, ERNST</mods:namePart>
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<mods:namePart id="69436069BCA0BDF57D03F5B2EAB30060">GUILBERT, ERIC</mods:namePart>
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<mods:title id="541EB951280E5154B5652DFB4B022127">Palaeoentomology</mods:title>
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<mods:part id="31339CA68A47AA439308F98FECDD1A01">
<mods:date id="F8347F360016F4C987A8345BF45EA7BE">2021</mods:date>
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<treatment id="4E015A1BFFBFFFE561F1F82EFC4FF7AA" ID-DOI="http://doi.org/10.5281/zenodo.5515435" ID-GBIF-Taxon="188169264" ID-Zenodo-Dep="5515435" LSID="urn:lsid:plazi:treatment:4E015A1BFFBFFFE561F1F82EFC4FF7AA" httpUri="http://treatment.plazi.org/id/4E015A1BFFBFFFE561F1F82EFC4FF7AA" lastPageId="1" lastPageNumber="31" pageId="0" pageNumber="30">
<subSubSection id="8EB2B886FFBFFFE461F1F82EF8E3F812" box="[136,456,1957,1982]" pageId="0" pageNumber="30" type="nomenclature">
<paragraph id="C617EB0DFFBFFFE461F1F82EF8E3F812" blockId="0.[136,456,1957,2019]" box="[136,456,1957,1982]" pageId="0" pageNumber="30">
<heading id="9D5F5C61FFBFFFE461F1F82EF8E3F812" bold="true" box="[136,456,1957,1982]" fontSize="10" level="2" pageId="0" pageNumber="30" reason="3">
<emphasis id="F4DC371FFFBFFFE461F1F82EF8E3F812" bold="true" box="[136,456,1957,1982]" pageId="0" pageNumber="30">
<taxonomicName id="01A8908EFFBFFFE461F1F82EF847F811" authority="HEISS &amp; GUILBERT, 2021" authorityName="HEISS &amp; GUILBERT" authorityYear="2021" box="[136,364,1957,1981]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="30" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="F4DC371FFFBFFFE461F1F82EF847F811" bold="true" box="[136,364,1957,1981]" italics="true" pageId="0" pageNumber="30">Burmacader grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFEF8A64FFBFFFE4600BF82DF8E3F812" box="[370,456,1958,1982]" pageId="0" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8EB2B886FFBFFFE461F1F841F821F84F" box="[136,266,1994,2019]" pageId="0" pageNumber="30" type="description">
<paragraph id="C617EB0DFFBFFFE461F1F841F821F84F" blockId="0.[136,456,1957,2019]" box="[136,266,1994,2019]" pageId="0" pageNumber="30">
(
<figureCitation id="5E93F788FFBFFFE461E9F841F9F8F84F" box="[144,211,1994,2019]" captionStart="FIGURE 1" captionStartId="1.[136,235,1777,1801]" captionTargetBox="[151,1436,189,1753]" captionTargetId="figure-80@1.[151,1436,189,1753]" captionTargetPageId="1" captionText="FIGURE 1. Burmacader grandis sp. nov. A and B, Habitus holotype, dorsal view. C, Ditto, head and pronotum. D, Burmese amber stone with B. grandis inclusion. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5507700" httpUri="https://zenodo.org/record/5507700/files/figure.png" pageId="0" pageNumber="30">Figs 1</figureCitation>
,
<figureCitation id="5E93F788FFBFFFE46199F840F829F84F" box="[224,258,1995,2019]" captionStart="FIGURE 2" captionStartId="2.[136,235,1021,1045]" captionTargetBox="[174,1400,196,975]" captionTargetId="figure-433@2.[151,1436,171,997]" captionTargetPageId="2" captionText="FIGURE 2. Burmacader, reconstruction of habitus of holotypes, dorsal view. A, B. grandis sp. nov. B, B. multivenosus. C, B. lativentris. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5507702" httpUri="https://zenodo.org/record/5507702/files/figure.png" pageId="0" pageNumber="30">2A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8EB2B886FFBFFFE461F1F79FFC87FC3D" pageId="0" pageNumber="30" type="materials_examined">
<paragraph id="C617EB0DFFBFFFE461F1F79FFC87FC3D" blockId="0.[136,778,2068,2092]" lastBlockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE461F1F79FF9DCF780" bold="true" box="[136,247,2068,2092]" pageId="0" pageNumber="30">
<typeStatus id="191355AFFFBFFFE461F1F79FF9D9F780" box="[136,242,2068,2092]" pageId="0" pageNumber="30" type="holotype">Holotype</typeStatus>
.
</emphasis>
<typeStatus id="191355AFFFBFFFE46071F79FF840F780" box="[264,363,2068,2092]" pageId="0" pageNumber="30" type="holotype">Holotype</typeStatus>
male in a 20×24×
<quantity id="015046E8FFBFFFE46323F79FFBBAF780" box="[602,657,2068,2092]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="0" pageNumber="30" unit="mm" value="5.0">5mm</quantity>
cabochon
<materialsCitation id="76C0E150FFBFFFE46253FD6EFC83FC3D" ID-GBIF-Occurrence="3353679301" collectionCode="CEHI" pageId="0" pageNumber="30" specimenCount="1" typeStatus="holotype">
shaped piece of Burmese amber, dorsal and ventral surface structures partly obscured by detritus, air bubbles and untransparent layer of resin. The specimen SC3798 is designated as
<typeStatus id="191355AFFFBFFFE462BAFCDFFD0BFCC0" box="[963,1056,852,876]" pageId="0" pageNumber="30" type="holotype">holotype</typeStatus>
and deposited in authors collection at the Tiroler Landesmuseum (
<collectionCode id="A0B973C8FFBFFFE465F0FCF2FDE1FC3D" box="[1161,1226,889,913]" pageId="0" pageNumber="30">CEHI</collectionCode>
) as BUB-TING-12
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8EB2B886FFBFFFE46226FC15FA4EFC53" pageId="0" pageNumber="30" type="etymology">
<paragraph id="C617EB0DFFBFFFE46226FC15FA4EFC53" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226FC15FACBFC1A" bold="true" box="[863,992,926,950]" pageId="0" pageNumber="30">Etymology.</emphasis>
The epithet of this species refers to its relatively large size from grandis (Latin), meaning large, wide.
</paragraph>
</subSubSection>
<subSubSection id="8EB2B886FFBFFFE46226FB87FCB3FB3F" pageId="0" pageNumber="30" type="diagnosis">
<paragraph id="C617EB0DFFBFFFE46226FB87FCB3FB3F" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226FB87FAFFFB88" bold="true" box="[863,980,1036,1060]" pageId="0" pageNumber="30">Diagnosis.</emphasis>
<taxonomicName id="01A8908EFFBFFFE462A4FB87FD96FB88" authority="HEISS &amp; GUILBERT, 2021" authorityName="HEISS &amp; GUILBERT" authorityYear="2021" box="[989,1213,1036,1060]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="30" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="F4DC371FFFBFFFE462A4FB87FD96FB88" box="[989,1213,1036,1060]" italics="true" pageId="0" pageNumber="30">Burmacader grandis</emphasis>
</taxonomicName>
<emphasis id="F4DC371FFFBFFFE465BEFB87FC34FB88" bold="true" box="[1223,1311,1036,1060]" pageId="0" pageNumber="30">
<taxonomicNameLabel id="EFEF8A64FFBFFFE465BEFB87FC34FB88" box="[1223,1311,1036,1060]" pageId="0" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from both congeners at first glance by larger size
<quantity id="015046E8FFBFFFE46473FBBAFC7FFBE5" box="[1290,1364,1073,1097]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="0" pageNumber="30" unit="mm" value="4.5">4.5mm</quantity>
(
<emphasis id="F4DC371FFFBFFFE4641CFBB9FCA9FBE5" box="[1381,1410,1074,1097]" italics="true" pageId="0" pageNumber="30">vs.</emphasis>
<quantity id="015046E8FFBFFFE464F3FBBAFA78FBC1" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="0" pageNumber="30" unit="mm" value="2.8">2.8 mm</quantity>
in
<taxonomicName id="01A8908EFFBFFFE46202FBDDFD0CFBC2" authorityName="Heiss &amp; Guilbert" authorityYear="2013" box="[891,1063,1110,1134]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="30" phylum="Arthropoda" rank="species" species="multivenosus">
<emphasis id="F4DC371FFFBFFFE46202FBDDFD0CFBC2" box="[891,1063,1110,1134]" italics="true" pageId="0" pageNumber="30">B. multivenosus</emphasis>
</taxonomicName>
,
<quantity id="015046E8FFBFFFE4654EFBDDFDB2FBC1" box="[1079,1177,1110,1134]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" pageId="0" pageNumber="30" unit="mm" value="3.15">3.15 mm</quantity>
in
<taxonomicName id="01A8908EFFBFFFE465B8FBDDFC65FBC2" authorityName="Heiss &amp; Guilbert" authorityYear="2018" box="[1217,1358,1110,1134]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="30" phylum="Arthropoda" rank="species" species="lativentris">
<emphasis id="F4DC371FFFBFFFE465B8FBDDFC65FBC2" box="[1217,1358,1110,1134]" italics="true" pageId="0" pageNumber="30">B. lativentris</emphasis>
</taxonomicName>
) and by larger size and different shape of paranota (
<figureCitation id="5E93F788FFBFFFE46581FBF0FCA7FB3F" box="[1272,1420,1147,1171]" captionStart="FIGURE 2" captionStartId="2.[136,235,1021,1045]" captionTargetBox="[174,1400,196,975]" captionTargetId="figure-433@2.[151,1436,171,997]" captionTargetPageId="2" captionText="FIGURE 2. Burmacader, reconstruction of habitus of holotypes, dorsal view. A, B. grandis sp. nov. B, B. multivenosus. C, B. lativentris. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5507702" httpUri="https://zenodo.org/record/5507702/files/figure.png" pageId="0" pageNumber="30">Fig. 2A, B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8EB2B886FFBFFFE56226FB14FC4FF7AA" lastPageId="1" lastPageNumber="31" pageId="0" pageNumber="30" type="description">
<paragraph id="C617EB0DFFBFFFE46226FB14FD7EFAAD" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226FB14FAC1FB1B" bold="true" box="[863,1002,1183,1207]" pageId="0" pageNumber="30">Description.</emphasis>
Body length
<quantity id="015046E8FFBFFFE465FDFB14FDF0FB1B" box="[1156,1243,1183,1207]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="0" pageNumber="30" unit="mm" value="4.5">4.5 mm</quantity>
, lateral margins of paranota and hemelytra beset with small setigerous tubercles and erect setae on antennae.
</paragraph>
<paragraph id="C617EB0DFFBFFFE46226FA85FCA9FA39" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226FA85FA8BFA8A" box="[863,928,1294,1318]" italics="true" pageId="0" pageNumber="30">Head.</emphasis>
Declivous (partly obscured, reconstruction in
<figureCitation id="5E93F788FFBFFFE464F8FA85FA65FAE7" captionStart="FIGURE 2" captionStartId="2.[136,235,1021,1045]" captionTargetBox="[174,1400,196,975]" captionTargetId="figure-433@2.[151,1436,171,997]" captionTargetPageId="2" captionText="FIGURE 2. Burmacader, reconstruction of habitus of holotypes, dorsal view. A, B. grandis sp. nov. B, B. multivenosus. C, B. lativentris. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5507702" httpUri="https://zenodo.org/record/5507702/files/figure.png" pageId="0" pageNumber="30">Fig. 2A</figureCitation>
), surface punctate, without spines or tubercles; antennae slender, segments II and IV subequal in length, segment III longest (exact measurement not possible); eyes globose.
</paragraph>
<paragraph id="C617EB0DFFBFFFE46226FA29FD28F9DE" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226FA29FAE2FA15" box="[863,969,1442,1465]" italics="true" pageId="0" pageNumber="30">Pronotum</emphasis>
. About 1.7 times as wide as long, attenuated anteriorly, lateral margins bisinuate with large wing-shaped expanded paranota with 7 rows of areolae at widest part, extending along 4/5 of lateral margin, surface of disk punctate and raised on posterior lobe, without carina and posterior projection.
</paragraph>
<paragraph id="C617EB0DFFBFFFE46226F9F4FAA7F917" blockId="0.[810,1452,741,2092]" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226F9F4FAFBF93B" box="[863,976,1663,1687]" italics="true" pageId="0" pageNumber="30">Scutellum.</emphasis>
Triangular with knob like apex, surface punctate.
</paragraph>
<paragraph id="C617EB0DFFBFFFE56226F943FB75F815" blockId="0.[810,1452,741,2092]" lastBlockId="1.[136,778,1917,2049]" lastPageId="1" lastPageNumber="31" pageId="0" pageNumber="30">
<emphasis id="F4DC371FFFBFFFE46226F943FAFCF94C" box="[863,983,1736,1760]" italics="true" pageId="0" pageNumber="30">Hemelytra.</emphasis>
Submacropterous distinctly surpassing apex of abdomen; clavus triangular with 67 rows of punctures as pronotum; costal area with 4 rows of areolae at middle and 5 at anterior indentation, these areolae larger than those of subcostal- and discoidal area; subcostal area wide with 910 rows of areolae at middle and 1012 transverse veinlets; discoidal area lanceolate with 68 rows of areolae and 56 transverse veinlets; sutural area with two rows of areolae along discoidal area widening to three rows at membrane but posteriorly reduced to one areola; a small hypocostal lamina developed with 23 small areolae on anterior 1/3 of ventral side of costal vein; stenocostal area lacking.
</paragraph>
<caption id="92D7BB85FFBEFFE561F1F97AFBF1F881" ID-DOI="http://doi.org/10.5281/zenodo.5507700" ID-Zenodo-Dep="5507700" httpUri="https://zenodo.org/record/5507700/files/figure.png" pageId="1" pageNumber="31" startId="1.[136,235,1777,1801]" targetBox="[151,1436,189,1753]" targetPageId="1">
<paragraph id="C617EB0DFFBEFFE561F1F97AFBF1F881" blockId="1.[136,1451,1777,1837]" pageId="1" pageNumber="31">
<emphasis id="F4DC371FFFBEFFE561F1F97AF82DF8A5" bold="true" box="[136,262,1777,1801]" pageId="1" pageNumber="31">FIGURE 1.</emphasis>
<taxonomicName id="01A8908EFFBEFFE56076F97AF8C8F8A5" authority="HEISS &amp; GUILBERT, 2021" authorityName="HEISS &amp; GUILBERT" authorityYear="2021" box="[271,483,1777,1801]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="F4DC371FFFBEFFE56076F97AF8C8F8A5" box="[271,483,1777,1801]" italics="true" pageId="1" pageNumber="31">Burmacader grandis</emphasis>
</taxonomicName>
<emphasis id="F4DC371FFFBEFFE56095F97AFB71F8A5" bold="true" box="[492,602,1777,1801]" pageId="1" pageNumber="31">
<taxonomicNameLabel id="EFEF8A64FFBEFFE56095F97AFB14F8A5" box="[492,575,1777,1801]" pageId="1" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
and
<emphasis id="F4DC371FFFBEFFE563E9F97AFB8AF8A5" bold="true" box="[656,673,1777,1801]" pageId="1" pageNumber="31">B</emphasis>
, Habitus holotype, dorsal view.
<emphasis id="F4DC371FFFBEFFE5628DF97AFD2DF8A5" bold="true" box="[1012,1030,1777,1801]" pageId="1" pageNumber="31">C</emphasis>
, Ditto, head and pronotum.
<emphasis id="F4DC371FFFBEFFE56449F97AFC69F8A5" bold="true" box="[1328,1346,1777,1801]" pageId="1" pageNumber="31">D</emphasis>
, Burmese amber stone with
<taxonomicName id="01A8908EFFBEFFE56044F89DF88DF881" authorityName="HEISS &amp; GUILBERT" authorityYear="2021" box="[317,422,1813,1837]" class="Insecta" family="Tingidae" genus="Burmacader" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F4DC371FFFBEFFE56044F89DF867F881" box="[317,332,1814,1837]" italics="true" pageId="1" pageNumber="31">B</emphasis>
.
<emphasis id="F4DC371FFFBEFFE56020F89EF88DF881" box="[345,422,1813,1837]" italics="true" pageId="1" pageNumber="31">grandis</emphasis>
</taxonomicName>
inclusion. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="C617EB0DFFBEFFE561C4F84EFB72F7AC" blockId="1.[136,778,1917,2049]" pageId="1" pageNumber="31">
<emphasis id="F4DC371FFFBEFFE561C4F84EF82AF870" box="[189,257,1989,2012]" italics="true" pageId="1" pageNumber="31">Venter</emphasis>
. Pro-, meso- and metasternum with median labial groove, labium reaching to posterior coxae.
</paragraph>
<paragraph id="C617EB0DFFBEFFE56226F8F4FD5AF817" blockId="1.[810,1452,1918,2055]" pageId="1" pageNumber="31">
<emphasis id="F4DC371FFFBEFFE56226F8F4FAB2F83A" box="[863,921,1919,1942]" italics="true" pageId="1" pageNumber="31">Legs.</emphasis>
Long and slender, tarsi two-segmented with long claws, pulvilli not discernable.
</paragraph>
<paragraph id="C617EB0DFFBEFFE56226F841FC4FF7AA" blockId="1.[810,1452,1918,2055]" pageId="1" pageNumber="31">
<emphasis id="F4DC371FFFBEFFE56226F841FD2AF84D" box="[863,1025,1994,2017]" italics="true" pageId="1" pageNumber="31">Measurements.</emphasis>
Length
<quantity id="015046E8FFBEFFE5651EF842FDE8F84C" box="[1127,1219,1993,2017]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="31" unit="mm" value="4.5">4.5 mm</quantity>
; width of pronotum across paranota
<quantity id="015046E8FFBEFFE562ACF865FD13F7AA" box="[981,1080,2030,2054]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="31" unit="mm" value="2.15">2.15 mm</quantity>
; width of abdomen
<quantity id="015046E8FFBEFFE56472F865FC4BF7AA" box="[1291,1376,2030,2054]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="31" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>