192 lines
18 KiB
XML
192 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.906.48255" ID-GBIF-Dataset="96b04a26-6084-45de-b832-bae14c14617b" ID-PMC="PMC6997243" ID-Pensoft-Pub="1313-2970-906-131" ID-Pensoft-UUID="06FA9B94737E597BA28734DCFA0FECCB" ID-PubMed="32042245" ID-ZooBank="AD702E753A08481E81B10203AEF362BE" ModsDocID="1313-2970-906-131" checkinTime="1580161947601" checkinUser="pensoft" docAuthor="Will, Kipling W." docDate="2020" docId="4D3EDE20AB6F57C3B79E98D70B15486C" docLanguage="en" docName="ZooKeys 906: 131-140" docOrigin="ZooKeys 906" docSource="http://dx.doi.org/10.3897/zookeys.906.48255" docTitle="Tiferonia leytensis Will 2020, sp. nov." docType="treatment" docUuid="939FDB9B-0561-473C-921D-D9AD409E87BF" docUuidSource="ZooBank" docVersion="4" id="06FA9B94737E597BA28734DCFA0FECCB" lastPageNumber="131" masterDocId="06FA9B94737E597BA28734DCFA0FECCB" masterDocTitle="Review and new species of Tiferonia Darlington, 1962 (Carabidae, Abacetini)" masterLastPageNumber="140" masterPageNumber="131" pageNumber="131" updateTime="1668168348473" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review and new species of Tiferonia Darlington, 1962 (Carabidae, Abacetini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Will, Kipling W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>906</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>131</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.906.48255</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.906.48255</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-906-131</mods:identifier>
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<mods:identifier type="ZooBank">AD702E753A08481E81B10203AEF362BE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">06FA9B94737E597BA28734DCFA0FECCB</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161840392" LSID="urn:lsid:zoobank.org:act:939FDB9B-0561-473C-921D-D9AD409E87BF" httpUri="http://treatment.plazi.org/id/4D3EDE20AB6F57C3B79E98D70B15486C" lastPageNumber="131" pageId="0" pageNumber="131">
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<subSubSection pageId="0" pageNumber="131" type="nomenclature">
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<paragraph pageId="0" pageNumber="131">
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<taxonomicName LSID="4D3EDE20-AB6F-57C3-B79E-98D70B15486C" authority="Will, 2020" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis" status="sp. nov.">Tiferonia leytensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="131">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tiferonia leytensis sp. nov. head, left lateral view. Arrows indicate a post-ocular sulcus and b posterior edge of the orbit." figureDoi="10.3897/zookeys.906.48255.figure1" httpUri="https://binary.pensoft.net/fig/374830" pageId="0" pageNumber="131">Figs 1</figureCitation>
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">, 2</figureCitation>
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<figureCitation captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">, 4</figureCitation>
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<figureCitation captionStart="Figures 6–9" captionStartId="F4" captionText="Figures 6 - 9. Tiferonia species dorsal habitus. 6 Tiferonia parva, male paratype 7 Tiferonia leytensis male holotype 8 Tiferonia schoutedeni 9 Tiferonia brunnea, male holotype." figureDoi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131">, 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="materials_examined">
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<paragraph pageId="0" pageNumber="131">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Holotype</emphasis>
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: Philippines • ♂, // "Plains of NE Leyte Is.,P.I. Nov
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<normalizedToken originalValue="’44-Jan’">'44-Jan'</normalizedToken>
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45 Darlington"// "MCZ Holotype 36215"// [deposited MCZ].
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</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Paratypes</emphasis>
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: Philippines • 1♂, 1 ♀, same data as holotype [MCZ].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="type locality">
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<paragraph pageId="0" pageNumber="131">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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As listed on locality label, type locality is estimated to be roughly centered on
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<geoCoordinate degrees="11.221" direction="north" orientation="latitude" precision="55" value="11.221">11.221N</geoCoordinate>
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,
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<geoCoordinate degrees="124.828" direction="east" orientation="longitude" precision="55" value="124.828">124.828E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="diagnosis">
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<paragraph pageId="0" pageNumber="131">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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The combination of two pairs of supraorbital setae, all elytral striae impressed, and the pronotum (Fig.
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<figureCitation captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">4</figureCitation>
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) basally narrowed with slightly sinuate lateral margins separates this species from all other species of
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131" start="Figures 4, 5" startId="F3">
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">Figures 4, 5.</emphasis>
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
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</taxonomicName>
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species pronota.
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<emphasis bold="true" pageId="0" pageNumber="131">4</emphasis>
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<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="131">5</emphasis>
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="description">
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<paragraph pageId="0" pageNumber="131">Description.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Size.</emphasis>
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Overall length (sbl) 3.96 mm [3.96-4.21 mm]; greatest width over elytra 1.65 mm [1.62-1.79 mm].
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Color.</emphasis>
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Dorsal and ventral surfaces piceous to dark rufous, head slightly darker, elytral interval 1 and apical fourth of elytra paler; legs, mouthparts, and antennae pale brunneous, tibiae darker rufous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Luster.</emphasis>
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Dorsally and ventrally distinctly shiny. Iridescence not evident on head, spectral iridescence slightly apparent on pronotum, distinctly evident on elytra, slightly apparent on ventral surface of body.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Head.</emphasis>
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Dorsal microsculpture evident nearly isodiametric mesh. Clypeal-ocular impressions clearly impressed, narrow, divergent, and extended to anterior supraocular seta. Ocular ratio 1.61 [1.55-1.58]. Eyes moderate size, prominent, with large posterior orbital area; gena slightly narrower than width of antennomere 1. Labrum with anterior margin straight. Mentum median tooth entire, triangular. Antennae, overall length moderately long, antennomeres 10-11 surpassing pronotal base.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Thorax.</emphasis>
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Pronotum dorsally impunctate, microsculpture not evident at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification in anterior half, irregular, very transverse mesh slightly evident near base; widest at middle, lateral margins with short, slight sinuation near hind angles; lateral marginal bead uniformly evident and narrow throughout; basal margin smooth, not beaded; anterior angles moderately produced; anterior submarginal sulcus broadly interrupted medially; hind angles right angled, denticulate; basal impressions linear, deeply impressed, slightly crenulate, shallowly reaching basal margin; seta at hind angle touching marginal bead. Elytra parallel sided; plica large and externally visible. Elytral striae well impressed, deeply, densely crenulate-punctate; intervals nearly flat, slightly convex basally. Elytral microsculpture scarcely evident as very transversely stretched sculpticells. Prosternal process rounded, margin not marked with bead; prosterna and proepisterna smooth. Mesosterna with few, coarse punctures. Metasternum laterally and metepisternum with shallow, coarse punctures.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Abdomen.</emphasis>
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Abdominal ventrites irregularly, coarsely punctate laterally, impunctate medially.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Male genitalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">2</figureCitation>
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), ostium dorsal, endophallus with light spine field in left apical position in repose.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Female ovipositor</emphasis>
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moderately long, slightly curved, two large ensiform setae, one dorsal one ventral, two long nematoform setae in well-developed groove.
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</paragraph>
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<caption doi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131" start="Figures 6–9" startId="F4">
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<paragraph pageId="0" pageNumber="131">
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<emphasis bold="true" pageId="0" pageNumber="131">Figures 6-9.</emphasis>
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
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</taxonomicName>
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species dorsal habitus.
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<emphasis bold="true" pageId="0" pageNumber="131">6</emphasis>
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
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</taxonomicName>
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, male paratype
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<emphasis bold="true" pageId="0" pageNumber="131">7</emphasis>
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<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
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</taxonomicName>
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male holotype
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<emphasis bold="true" pageId="0" pageNumber="131">8</emphasis>
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<taxonomicName authorityName="Will" authorityYear="2020" baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia schoutedeni</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="131">9</emphasis>
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<taxonomicName baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia brunnea" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia brunnea</emphasis>
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</taxonomicName>
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, male holotype.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="etymology">
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<paragraph pageId="0" pageNumber="131">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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The specific epithet
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<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
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<emphasis italics="true" pageId="0" pageNumber="131">leytensis</emphasis>
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</taxonomicName>
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is based on the type locality and is treated as an adjective.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="131" type="notes">
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<paragraph pageId="0" pageNumber="131">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="131">
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In his discussion of the species of
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<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
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</taxonomicName>
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,
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<bibRefCitation author="Darlington, PJ" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="131" pagination="321 - 564" refId="B5" refString="Darlington, PJ, 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126: 321 - 564" title="The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini." volume="126" year="1962">Darlington (1962</bibRefCitation>
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: 561) states that
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<taxonomicName lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
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<emphasis italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
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</taxonomicName>
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from the Philippines is distinguished by having only a single pair of supraorbital setae and then states that he has a series of that species from Leyte Island. However, this appears to be an error. Among
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<normalizedToken originalValue="Darlington’s">Darlington's</normalizedToken>
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specimens at the MCZ he has a series from Leyte Island, but they all have two pairs of supraorbital setae and are otherwise distinctly different from
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<taxonomicName lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
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<emphasis italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
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</taxonomicName>
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. These specimens comprise the holotype and paratypes of
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<taxonomicName lsidName="T. leytensis" pageId="0" pageNumber="131" rank="species" species="leytensis">
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<emphasis italics="true" pageId="0" pageNumber="131">T. leytensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |