655 lines
93 KiB
XML
655 lines
93 KiB
XML
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<mods:title id="F9F291A75BABDBBB4EA89D5D621D823D">Unveiling the Rosetta Stone of syllids: redescription and neotype designation of Syllis monilaris Savigny in Lamarck, 1818, type species of type genus of family Syllidae Grube, 1850 (Annelida)</mods:title>
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<treatment id="03BB87D0FFCC0725FF4E8386D6AFFADD" ID-DOI="http://doi.org/10.5281/zenodo.3507808" ID-GBIF-Taxon="119644514" ID-Zenodo-Dep="3507808" LSID="urn:lsid:plazi:treatment:03BB87D0FFCC0725FF4E8386D6AFFADD" httpUri="http://treatment.plazi.org/id/03BB87D0FFCC0725FF4E8386D6AFFADD" lastPageId="10" lastPageNumber="327" pageId="5" pageNumber="322">
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<paragraph id="8BAD36C6FFCC072AFF4E8386D4C0FF07" blockId="5.[151,672,151,211]" box="[151,672,151,178]" pageId="5" pageNumber="322">
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<taxonomicName id="4C124D45FFCC072AFF4E8386D7DDFF07" ID-CoL="7B73R" authority="Savigny" authorityName="Savigny" box="[151,445,151,178]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E8386D733FF04" bold="true" box="[151,339,151,177]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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Savigny
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<emphasis id="B966EAD4FFCC072AFE1C8386D7BEFF04" bold="true" box="[453,478,151,177]" italics="true" pageId="5" pageNumber="322">in</emphasis>
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<bibRefCitation id="EF834B37FFCC072AFE3D8389D4C0FF07" author="Lamarck" box="[484,672,152,178]" pageId="5" pageNumber="322" refString="Lamarck, J. B. (1818) Histoire Naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedes d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin l'exposition des principes fondamentaux de la zoologie. Vol. 5. Deterville, Paris, 612 pp." type="book" year="1818">Lamarck, 1818</bibRefCitation>
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<paragraph id="8BAD36C6FFCC072AFF4E83ABD77CFF66" blockId="5.[151,672,151,211]" box="[151,284,186,211]" pageId="5" pageNumber="322">
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<figureCitation id="13292A43FFCC072AFF4E83ABD77CFF66" box="[151,284,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1366,1388]" captionStartId-1="5.[151,250,1530,1552]" captionStartId-2="7.[151,250,1897,1919]" captionStartId-3="8.[151,250,1846,1868]" captionStartId-4="9.[151,250,1725,1747]" captionTargetBox-1="[256,1330,903,1505]" captionTargetBox-2="[161,1428,419,1862]" captionTargetBox-3="[159,1413,433,1822]" captionTargetBox-4="[193,1394,183,1698]" captionTargetId-1="figure@5.[253,1333,900,1509]" captionTargetId-2="figure@7.[151,1436,409,1876]" captionTargetId-3="figure@8.[151,1436,413,1835]" captionTargetId-4="figure@9.[176,1405,169,1709]" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 1. Syllis monilaris. Original plate by Savigny (1817)." captionText-1="FIGURE 2. Syllis monilaris. Photographs of neotype (ZMB F 1941). A, complete specimen, with original ZMB’s label. B, anterior end, dorsal view. C, midbody, dorsal view. D, compound chaetae, anterior parapodia. E, compound chaetae, midbody." captionText-2="FIGURE 3. Syllis monilaris. Neotype (ZMB F 1941). Anterior end, dorsal view. Scale: 0.4 mm." captionText-3="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." captionText-4="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri-1="https://zenodo.org/record/238850/files/figure.png" httpUri-2="https://zenodo.org/record/238851/files/figure.png" httpUri-3="https://zenodo.org/record/238852/files/figure.png" httpUri-4="https://zenodo.org/record/238853/files/figure.png" pageId="5" pageNumber="322">Figures 1–5</figureCitation>
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<taxonomicName id="4C124D45FFCC072AFF4E8213D7EEFEAD" ID-CoL="7B73R" authority="Savigny" authorityName="Savigny" box="[151,398,258,280]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E8213D755FEAD" box="[151,309,258,280]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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Savigny
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<emphasis id="B966EAD4FFCC072AFE4D8212D7C8FEAD" box="[404,424,259,280]" italics="true" pageId="5" pageNumber="322">in</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFE768213D418FEAD" author="Lamarck" box="[431,632,258,280]" page="318" pageId="5" pageNumber="322" year="1818">
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: 318
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.
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<taxonomicName id="4C124D45FFCC072AFF4E8230D78FFE82" ID-CoL="7B73R" authority="Savigny 1822: 44" authorityName="Savigny" authorityPageNumber="44" authorityYear="1822" box="[151,495,289,311]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E8230D755FE82" box="[151,309,289,311]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFEE28230D78FFE82" author="Savigny" box="[315,495,289,311]" page="44" pageId="5" pageNumber="322" year="1822">
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<bibRefCitation id="EF834B37FFCC072AFEE28230D7ABFE82" author="Savigny" box="[315,459,289,311]" pageId="5" pageNumber="322" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny 1822</bibRefCitation>
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: 44
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</treatmentCitation>
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–45, pl. 4, fig. 3.
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<taxonomicName id="4C124D45FFCC072AFF4E8251D796FEE3" ID-CoL="7B73R" authority="Audouin 1826: 71" authorityName="Audouin" authorityPageNumber="71" authorityYear="1826" box="[151,502,320,342]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E8251D755FEE3" box="[151,309,320,342]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFEE28251D796FEE3" author="Audouin" box="[315,502,320,342]" page="71" pageId="5" pageNumber="322" year="1826">
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<bibRefCitation id="EF834B37FFCC072AFEE28251D7B0FEE3" author="Audouin" box="[315,464,320,342]" pageId="5" pageNumber="322" refString="Audouin, V. (1826) Explication sommaire des planches d'annelides de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, membre de l'Institut; offrant un expose des caracteres naturelles des genres, avec la distinction et quelquefois le nom des especes. In: Audouin V. Explication sommaire des planches dont les dessins ont ete fournis par M. J. C. Savigny, pour l'histoire naturelle de l'ouvrage. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (4), pp. 57 - 76." type="journal article" year="1826">Audouin 1826</bibRefCitation>
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: 71
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAD36C6FFCC072AFF4E824FD78CFE07" blockId="5.[151,737,258,526]" pageId="5" pageNumber="322">
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<taxonomicName id="4C124D45FFCC072AFF4E824FD79BFEC1" ID-CoL="7B73R" authority="Savigny 1826: 372" authorityName="Savigny" authorityPageNumber="372" authorityYear="1826" box="[151,507,350,372]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E824FD755FEC1" box="[151,309,350,372]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFEE2824FD79BFEC1" author="Savigny" box="[315,507,350,372]" page="372" pageId="5" pageNumber="322" year="1826">
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: 372
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</treatmentCitation>
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</taxonomicName>
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–373, pl. IV, fig. 3.
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<taxonomicName id="4C124D45FFCC072AFF4E826CD46EFE26" ID-CoL="7B73R" authority="Blainville 1828: 473" authorityName="Blainville" authorityPageNumber="473" authorityYear="1828" box="[151,526,381,403]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E826CD755FE26" box="[151,309,381,403]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFEE2826CD46EFE26" author="Blainville" box="[315,526,381,403]" page="473" pageId="5" pageNumber="322" year="1828">
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<bibRefCitation id="EF834B37FFCC072AFEE2826CD7BBFE26" author="Blainville" box="[315,475,381,403]" pageId="5" pageNumber="322" refString="Blainville, H. M. (1828) Dictionnaire des Sciences Naturelles dans lequel on traite methodiquement des differents etres de la nature, consideres soit en eux-memes, d'apres l'etat actuel de nos connaissances, soit relativement a l'utilite qu'en peuvent retirer la medecine, l'agriculture, le commerce et les arts. Suivi d'une biographie des plus celebres naturalistes, 57, 1 - 628." type="journal article" year="1828">Blainville 1828</bibRefCitation>
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: 473
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</treatmentCitation>
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</taxonomicName>
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, pl. 17, fig. 2.
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||
<taxonomicName id="4C124D45FFCC072AFF4E828DD788FE07" ID-CoL="7B73R" authority="Peters 1855: 613" authorityName="Peters" authorityPageNumber="613" authorityYear="1855" box="[151,488,412,434]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E828DD755FE07" box="[151,309,412,434]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
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<treatmentCitation id="0AB310D7FFCC072AFEE2828CD788FE07" author="Peters" box="[315,488,412,434]" page="613" pageId="5" pageNumber="322" year="1855">
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<bibRefCitation id="EF834B37FFCC072AFEE2828CD7D6FE07" author="Peters" box="[315,438,412,434]" pageId="5" pageNumber="322" refString="Peters, W. (1855) Uber die Gattung Bdella Savigny, (Limnatis, Moquin-Tandon) und die in Mossambique beobachteten Anneliden. Bericht uber die zur Bekanntmachung geeigneten Verhandlungen Koniglich Preußischen Akademie der Wissenschaften zu Berlin, 1854, 607 - 614 + corrigenda." type="journal article" year="1855">Peters 1855</bibRefCitation>
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: 613
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</treatmentCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BAD36C6FFCC072AFF4E82ABD790FE65" blockId="5.[151,737,258,526]" box="[151,496,442,464]" pageId="5" pageNumber="322">
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<taxonomicName id="4C124D45FFCC072AFF4E82ABD78CFE65" ID-CoL="7B73R" authority="Licher 1999: 292" authorityName="Licher" authorityPageNumber="292" authorityYear="1999" box="[151,492,442,464]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="B966EAD4FFCC072AFF4E82ABD755FE65" box="[151,309,442,464]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
|
||
<treatmentCitation id="0AB310D7FFCC072AFEE282ABD78CFE65" author="Licher" box="[315,492,442,464]" page="292" pageId="5" pageNumber="322" year="1999">
|
||
<bibRefCitation id="EF834B37FFCC072AFEE282ABD7D9FE65" author="Licher" box="[315,441,442,464]" pageId="5" pageNumber="322" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
|
||
: 292
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFCC072AFF4E82C8D471FDBB" blockId="5.[151,737,258,526]" pageId="5" pageNumber="322">
|
||
<treatmentCitationGroup id="AB0211E8FFCC072AFF4E82C8D471FDBB" pageId="5" pageNumber="322">
|
||
<taxonomicName id="4C124D45FFCC072AFF4E82C8D736FE5A" ID-CoL="85S3S" box="[151,342,473,495]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="moniliformis">
|
||
<emphasis id="B966EAD4FFCC072AFF4E82C8D736FE5A" box="[151,342,473,495]" italics="true" pageId="5" pageNumber="322">Syllis moniliformis</emphasis>
|
||
</taxonomicName>
|
||
[sic;
|
||
<emphasis id="B966EAD4FFCC072AFE4D82C8D440FE5A" box="[404,544,473,495]" italics="true" pageId="5" pageNumber="322">lapsus calami</emphasis>
|
||
]
|
||
<treatmentCitation id="0AB310D7FFCC072AFDF682C8D4BDFE5A" author="Grube" box="[559,733,473,495]" page="502" pageId="5" pageNumber="322" year="1869">
|
||
<bibRefCitation id="EF834B37FFCC072AFDF682C8D4CAFE5A" author="Grube" box="[559,682,473,495]" pageId="5" pageNumber="322" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube 1869</bibRefCitation>
|
||
: 502
|
||
</treatmentCitation>
|
||
.
|
||
<taxonomicName id="4C124D45FFCC072AFF4E82E9D46DFDBB" ID-CoL="85S3S" authority="Licher 1999: 306" authorityName="Licher" authorityPageNumber="306" authorityYear="1999" box="[151,525,504,526]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="moniliformis">
|
||
<emphasis id="B966EAD4FFCC072AFF4E82E9D736FDBB" box="[151,342,504,526]" italics="true" pageId="5" pageNumber="322">Syllis moniliformis</emphasis>
|
||
<treatmentCitation id="0AB310D7FFCC072AFE8582E9D46DFDBB" author="Licher" box="[348,525,504,526]" page="306" pageId="5" pageNumber="322" year="1999">
|
||
<bibRefCitation id="EF834B37FFCC072AFE8582E9D7BAFDBB" author="Licher" box="[348,474,504,526]" pageId="5" pageNumber="322" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
|
||
: 306
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308654DFFCC072AFF4E812BD417FCFA" pageId="5" pageNumber="322" type="materials_examined">
|
||
<paragraph id="8BAD36C6FFCC072AFF4E812BD6A8FD2F" blockId="5.[151,1437,570,847]" pageId="5" pageNumber="322">
|
||
<emphasis id="B966EAD4FFCC072AFF4E812BD7E6FDE6" bold="true" box="[151,390,570,595]" pageId="5" pageNumber="322">Material examined.</emphasis>
|
||
El Tor (
|
||
<collectingCountry id="F3057656FFCC072AFE2B812AD459FDE6" box="[498,569,571,595]" name="Egypt" pageId="5" pageNumber="322">Egypt</collectingCountry>
|
||
, Sinai Peninsula, Gulf of Suez), C.G. Ehrenberg coll., id. A.E. Grube,
|
||
<specimenCount id="9D14FD4FFFCC072AFA57812AD770FDC2" pageId="5" pageNumber="322" type="generic">2 specimens</specimenCount>
|
||
:
|
||
<collectionCode id="ED03AE03FFCC072AFEC4814ED739FDC3" box="[285,345,607,630]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="5" pageNumber="322">ZMB</collectionCode>
|
||
F1941, 1 bigger specimen, designated as
|
||
<typeStatus id="54A98864FFCC072AFCE7814ED5FBFDC2" box="[830,923,607,631]" pageId="5" pageNumber="322" type="neotype">neotype</typeStatus>
|
||
;
|
||
<collectionCode id="ED03AE03FFCC072AFC71814ED584FDC3" box="[936,996,607,630]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="5" pageNumber="322">ZMB</collectionCode>
|
||
11529, 1 smaller specimen, broken in two.
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFCC072AFF1E81B7D417FCFA" blockId="5.[151,1437,570,847]" pageId="5" pageNumber="322">
|
||
<emphasis id="B966EAD4FFCC072AFF1E81B7D7DEFD0A" bold="true" box="[199,446,678,703]" pageId="5" pageNumber="322">Additional material.</emphasis>
|
||
<collectingCountry id="F3057656FFCC072AFE1081B7D449FD0A" box="[457,553,678,703]" name="United Kingdom" pageId="5" pageNumber="322">England</collectingCountry>
|
||
, off Folkestone, eastern English Channel,
|
||
<geoCoordinate id="EE265001FFCC072AFBC781B7D2D2FD0B" box="[1054,1202,678,703]" direction="north" orientation="latitude" pageId="5" pageNumber="322" precision="1" value="51.02726">51.02726º N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE265001FFCC072AFB1B81B6D325FD0B" box="[1218,1349,679,703]" direction="east" orientation="longitude" pageId="5" pageNumber="322" precision="1" value="1.30379">1.30379º E</geoCoordinate>
|
||
, coarse sediment with
|
||
<taxonomicName id="4C124D45FFCC072AFE9B81DBD44CFD56" box="[322,556,714,739]" class="Polychaeta" family="Sabellariidae" genus="Sabellaria" kingdom="Animalia" order="Sabellida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="spinulosa">
|
||
<emphasis id="B966EAD4FFCC072AFE9B81DBD44CFD56" box="[322,556,714,739]" italics="true" pageId="5" pageNumber="322">Sabellaria spinulosa</emphasis>
|
||
</taxonomicName>
|
||
crust,
|
||
<quantity id="4CEA9B23FFCC072AFDA481DAD4D9FD57" box="[637,697,715,739]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="322" unit="m" value="33.0">33 m</quantity>
|
||
depth,
|
||
<date id="FFAC1006FFCC072AFCCB81DAD5C4FD56" box="[786,932,715,739]" pageId="5" pageNumber="322" value="2014-06-22">22 Jun 2014</date>
|
||
, Fokestone Pomerania Survey,
|
||
<specimenCount id="9D14FD4FFFCC072AFAC981DAD3FDFD56" box="[1296,1437,714,739]" pageId="5" pageNumber="322" type="generic">3 specimens</specimenCount>
|
||
(collection of
|
||
<collectionCode id="ED03AE03FFCC072AFEEF81FED7E4FCB2" box="[310,388,751,775]" pageId="5" pageNumber="322">APEM</collectionCode>
|
||
Ltd.).
|
||
<collectingCountry id="F3057656FFCC072AFE0A81FFD45DFCB2" box="[467,573,750,775]" name="Australia" pageId="5" pageNumber="322">Australia</collectingCountry>
|
||
, Western
|
||
<collectingCountry id="F3057656FFCC072AFD7481FFD577FCB2" box="[685,791,750,775]" name="Australia" pageId="5" pageNumber="322">Australia</collectingCountry>
|
||
, Houtman Abrolhos, NE entrance Goss Passage, Beacon Island,
|
||
<geoCoordinate id="EE265001FFCC072AFF308002D71CFC9E" box="[233,380,786,811]" direction="south" orientation="latitude" pageId="5" pageNumber="322" precision="15" value="-28.465">28° 27' 54" S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE265001FFCC072AFE498002D454FC9F" box="[400,564,786,811]" direction="east" orientation="longitude" pageId="5" pageNumber="322" precision="15" value="113.778336">113° 46' 42" E</geoCoordinate>
|
||
,
|
||
<date id="FFAC1006FFCC072AFDE68003D4B6FC9E" box="[575,726,786,811]" pageId="5" pageNumber="322" value="1994-05-25">
|
||
25 May
|
||
<quantity id="4CEA9B23FFCC072AFD428002D4B6FC9E" box="[667,726,787,811]" metricMagnitude="3" metricUnit="m" metricValue="1.99433" pageId="5" pageNumber="322" unit="m" value="1994.33">1994</quantity>
|
||
</date>
|
||
, 33 m depth, underneath boulders embedded in coral sand, coll. Pat Hutchings,
|
||
<specimenCount id="9D14FD4FFFCC072AFE9A8026D7A2FCFA" box="[323,450,822,847]" pageId="5" pageNumber="322" type="generic">1 specimen</specimenCount>
|
||
(AM W29492).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6D664EFFCC072AFF4E86EBD3F4F99B" httpUri="https://zenodo.org/record/238850/files/figure.png" pageId="5" pageNumber="322" targetBox="[256,1330,903,1505]" targetPageId="5">
|
||
<paragraph id="8BAD36C6FFCC072AFF4E86EBD3F4F99B" blockId="5.[151,1435,1530,1582]" pageId="5" pageNumber="322">
|
||
<emphasis id="B966EAD4FFCC072AFF4E86EBD775F9BA" bold="true" box="[151,277,1530,1552]" pageId="5" pageNumber="322">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C124D45FFCC072AFEC786EBD7DFF9A5" box="[286,447,1530,1552]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="322" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCC072AFEC786EBD7DFF9A5" box="[286,447,1530,1552]" italics="true" pageId="5" pageNumber="322">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
. Photographs of neotype (ZMB F1941). A, complete specimen, with original ZMB’s label. B, anterior end, dorsal view. C, midbody, dorsal view. D, compound chaetae, anterior parapodia. E, compound chaetae, midbody.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C308654DFFCC0729FF1E8544D486FCA0" lastPageId="6" lastPageNumber="323" pageId="5" pageNumber="322" type="description">
|
||
<paragraph id="8BAD36C6FFCC0729FF1E8544D202FD79" blockId="5.[151,1437,1621,2006]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="323" pageId="5" pageNumber="322">
|
||
<emphasis id="B966EAD4FFCC072AFF1E8544D495F9DB" bold="true" box="[199,757,1621,1646]" pageId="5" pageNumber="322">
|
||
Redescription based on
|
||
<typeStatus id="54A98864FFCC072AFE338544D427F9DB" box="[490,583,1621,1646]" pageId="5" pageNumber="322" type="neotype">neotype</typeStatus>
|
||
(ZMB F1941)
|
||
</emphasis>
|
||
. Body of large size, elongated (
|
||
<figureCitation id="13292A43FFCC072AFBBD8544D2CCF9DB" box="[1124,1196,1621,1646]" captionStart="FIGURE 2" captionStartId="5.[151,250,1530,1552]" captionTargetBox="[256,1330,903,1505]" captionTargetId="figure@5.[253,1333,900,1509]" captionTargetPageId="5" captionText="FIGURE 2. Syllis monilaris. Photographs of neotype (ZMB F 1941). A, complete specimen, with original ZMB’s label. B, anterior end, dorsal view. C, midbody, dorsal view. D, compound chaetae, anterior parapodia. E, compound chaetae, midbody." httpUri="https://zenodo.org/record/238850/files/figure.png" pageId="5" pageNumber="322">Figs 2</figureCitation>
|
||
A–C, 3), incomplete, 5.8 cm and 296 chaetigers long, almost
|
||
<quantity id="4CEA9B23FFCC072AFD8A856BD4F4F927" box="[595,660,1658,1682]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="322" unit="mm" value="1.0">1 mm</quantity>
|
||
wide on anterior body (
|
||
<figureCitation id="13292A43FFCC072AFC46856BD582F927" box="[927,994,1658,1683]" captionStart="FIGURE 3" captionStartId="7.[151,250,1897,1919]" captionTargetBox="[161,1428,419,1862]" captionTargetId="figure@7.[151,1436,409,1876]" captionTargetPageId="7" captionText="FIGURE 3. Syllis monilaris. Neotype (ZMB F 1941). Anterior end, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238851/files/figure.png" pageId="5" pageNumber="322">Fig. 3</figureCitation>
|
||
), wider on mid-posterior segments (1.4 mm) (
|
||
<figureCitation id="13292A43FFCC072AFF04858CD743F903" box="[221,291,1693,1718]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="5" pageNumber="322">Fig. 4</figureCitation>
|
||
C), slender posteriorly (0.7 mm) (
|
||
<figureCitation id="13292A43FFCC072AFD7F858CD48CF903" box="[678,748,1693,1718]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="5" pageNumber="322">Fig. 4</figureCitation>
|
||
D). Body now very dark, opaque (
|
||
<figureCitation id="13292A43FFCC072AFBAF858CD2A5F903" box="[1142,1221,1693,1718]" captionStart="FIGURE 2" captionStartId="5.[151,250,1530,1552]" captionTargetBox="[256,1330,903,1505]" captionTargetId="figure@5.[253,1333,900,1509]" captionTargetPageId="5" captionText="FIGURE 2. Syllis monilaris. Photographs of neotype (ZMB F 1941). A, complete specimen, with original ZMB’s label. B, anterior end, dorsal view. C, midbody, dorsal view. D, compound chaetae, anterior parapodia. E, compound chaetae, midbody." httpUri="https://zenodo.org/record/238850/files/figure.png" pageId="5" pageNumber="322">Figs. 2</figureCitation>
|
||
A–C). Prostomium pentagonal; 4 eyes in trapezoidal arrangement. Palps robust, similar in length to prostomium. Median antenna on posterior part of prostomium, between posterior eyes, with 16 articles, slightly longer than combined length of prostomium and palps; lateral antennae shorter than median one, with 13–14 articles each. Peristomium shorter than subsequent segments (
|
||
<figureCitation id="13292A43FFCC072AFE1E843CD46BF8F3" box="[455,523,1837,1862]" captionStart="FIGURE 3" captionStartId="7.[151,250,1897,1919]" captionTargetBox="[161,1428,419,1862]" captionTargetId="figure@7.[151,1436,409,1876]" captionTargetPageId="7" captionText="FIGURE 3. Syllis monilaris. Neotype (ZMB F 1941). Anterior end, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238851/files/figure.png" pageId="5" pageNumber="322">Fig. 3</figureCitation>
|
||
). Dorsal tentacular cirri longer than median antenna, about twice as long as lateral ones, with about 25 articles each; ventral tentacular cirri about half length of dorsal ones, with 18 articles each. Dorsal cirri longer than body width on anterior segments, becoming similar to shorter than body width from midbody backwards, and more or less fusiform, especially from midbody backwards (
|
||
<figureCitation id="13292A43FFCC072AFB4C848BD2BCF807" box="[1173,1244,1946,1971]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="5" pageNumber="322">Fig. 4</figureCitation>
|
||
A–C), with well defined cirrophores and articles; articles basally and distally smaller than medially; anteriormost dorsal cirri longer than dorsal tentacular cirri, with 28–22–29–32–20–26–21–19–27 articles (left side), 29–19–26–32–22–22–21–23– 23 articles (right side) (
|
||
<figureCitation id="13292A43FFCF0729FE7983ADD785FF61" box="[416,485,188,213]" captionStart="FIGURE 3" captionStartId="7.[151,250,1897,1919]" captionTargetBox="[161,1428,419,1862]" captionTargetId="figure@7.[151,1436,409,1876]" captionTargetPageId="7" captionText="FIGURE 3. Syllis monilaris. Neotype (ZMB F 1941). Anterior end, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238851/files/figure.png" pageId="6" pageNumber="323">Fig. 3</figureCitation>
|
||
); at proventricular level, all cirri becoming shorter and similar in length, with about 14–18 articles (
|
||
<figureCitation id="13292A43FFCF0729FE9083CED7EFFF4D" box="[329,399,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="6" pageNumber="323">Fig. 4</figureCitation>
|
||
A); 12–14 articles on midbody (
|
||
<figureCitation id="13292A43FFCF0729FCDB83CED528FF4D" box="[770,840,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="6" pageNumber="323">Fig. 4</figureCitation>
|
||
B), with similar cirri alternating irregularly between long and short; on midbody-posterior segments, dorsal cirri proportionally thicker and more fusiform, with 12–17 articles (
|
||
<figureCitation id="13292A43FFCF0729FF208236D75FFEF5" box="[249,319,295,320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="6" pageNumber="323">Fig. 4</figureCitation>
|
||
C); on posteriormost segments, dorsal cirri smaller, with 10–14 articles each (
|
||
<figureCitation id="13292A43FFCF0729FB638236D360FEF5" box="[1210,1280,295,320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="6" pageNumber="323">Fig. 4</figureCitation>
|
||
D). Parapodia conical, distally bilobed on dorsal view (
|
||
<figureCitation id="13292A43FFCF0729FDBE825DD4CCFED1" box="[615,684,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1897,1919]" captionTargetBox="[161,1428,419,1862]" captionTargetId="figure@7.[151,1436,409,1876]" captionTargetPageId="7" captionText="FIGURE 3. Syllis monilaris. Neotype (ZMB F 1941). Anterior end, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238851/files/figure.png" pageId="6" pageNumber="323">Fig. 3</figureCitation>
|
||
). Ventral cirri digitiform, longer than parapodial lobes on anterior parapodia, becoming shorter from midbody onwards. Chaetae mostly broken but some parapodia with complete fascicles all along the body. All chaetae heterogomph compound, with thick shafts and short, triangular, unidentate blades sometimes with short spines on margin, but blades usually smooth, similar in shape and size throughout body (
|
||
<figureCitation id="13292A43FFCF0729FF0782CDD745FE41" box="[222,293,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1530,1552]" captionTargetBox="[256,1330,903,1505]" captionTargetId="figure@5.[253,1333,900,1509]" captionTargetPageId="5" captionText="FIGURE 2. Syllis monilaris. Photographs of neotype (ZMB F 1941). A, complete specimen, with original ZMB’s label. B, anterior end, dorsal view. C, midbody, dorsal view. D, compound chaetae, anterior parapodia. E, compound chaetae, midbody." httpUri="https://zenodo.org/record/238850/files/figure.png" pageId="6" pageNumber="323">Figs 2</figureCitation>
|
||
D–E, 5A–C). Upper blades 35 µm long, lower ones 25 µm long; usually dorsal-most falciger on each parapodium with slightly thicker shaft than remaining ones. Anterior parapodia each with 9 compound chaetae (
|
||
<figureCitation id="13292A43FFCF0729FF468135D683FD88" box="[159,227,548,573]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
A); occasionally some chaetae slightly bidentate; midbody parapodia with 8 compound chaetae (
|
||
<figureCitation id="13292A43FFCF0729FAED8135D31AFD88" box="[1332,1402,548,573]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
B); posterior parapodia with 5 compound chaetae (
|
||
<figureCitation id="13292A43FFCF0729FD7D8156D487FDD5" box="[676,743,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
C). Dorsal and ventral simple chaetae not observed (specimen incomplete). Anterior parapodia with 7 aciculae (
|
||
<figureCitation id="13292A43FFCF0729FD3C817DD54EFD30" box="[741,814,620,645]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
D), 4 at midbody (
|
||
<figureCitation id="13292A43FFCF0729FBCF817DD23FFD30" box="[1046,1119,620,645]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
E), and
|
||
<quantity id="4CEA9B23FFCF0729FB1A817DD295FD30" box="[1219,1269,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="323" unit="in" value="2.0">2 in</quantity>
|
||
posteriormost parapodia, one distally blunt and other slightly acuminate (
|
||
<figureCitation id="13292A43FFCF0729FCFF819ED508FD1D" box="[806,872,655,680]" captionStart="FIGURE 5" captionStartId="9.[151,250,1725,1747]" captionTargetBox="[193,1394,183,1698]" captionTargetId="figure@9.[176,1405,169,1709]" captionTargetPageId="9" captionText="FIGURE 5. Syllis monilaris. Neotype (ZMB F 1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm." httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="6" pageNumber="323">Fig. 5</figureCitation>
|
||
F). Pharynx extending through about 13 segments; pharyngeal tooth on anterior margin of pharynx. Proventricle from chaetigers 13 to 21.
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFCF0729FF1E81C6D486FCA0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="323">
|
||
Second specimen (ZMB 11529) much smaller than
|
||
<typeStatus id="54A98864FFCF0729FCD081C9D504FD45" box="[777,868,728,752]" pageId="6" pageNumber="323" type="neotype">neotype</typeStatus>
|
||
, broken in two pieces and also incomplete,
|
||
<quantity id="4CEA9B23FFCF0729FA9781C9D3FDFD5A" box="[1358,1437,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="323" unit="mm" value="13.0">13 mm</quantity>
|
||
long and 76 chaetigers,
|
||
<quantity id="4CEA9B23FFCF0729FE7B81EDD782FCA1" box="[418,482,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="323" unit="mm" value="1.0">1 mm</quantity>
|
||
wide on anterior body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308654DFFCF0729FF1E800ED7A8FC59" pageId="6" pageNumber="323" type="distribution">
|
||
<paragraph id="8BAD36C6FFCF0729FF1E800ED341FCE8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="323">
|
||
<emphasis id="B966EAD4FFCF0729FF1E800ED739FC8D" bold="true" box="[199,345,799,824]" pageId="6" pageNumber="323">Distribution</emphasis>
|
||
. Only known from the Gulf of Suez. In agreement with the ICZN article 76.3, the
|
||
<typeStatus id="54A98864FFCF0729FB2A8031D343FC8D" box="[1267,1315,800,824]" pageId="6" pageNumber="323">type</typeStatus>
|
||
locality of the species becomes El Tor (
|
||
<collectingCountry id="F3057656FFCF0729FE0F8055D47DFCE9" box="[470,541,836,860]" name="Egypt" pageId="6" pageNumber="323">Egypt</collectingCountry>
|
||
, Sinai Peninsula, Gulf of Suez), locality of collection of the
|
||
<typeStatus id="54A98864FFCF0729FB198054D37CFCE8" box="[1216,1308,837,861]" pageId="6" pageNumber="323" type="neotype">neotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFCF0729FF1E8076D7A8FC59" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="323">
|
||
<emphasis id="B966EAD4FFCF0729FF1E8076D743FC35" bold="true" box="[199,291,871,896]" pageId="6" pageNumber="323">Habitat</emphasis>
|
||
. Stated to be common on the coasts of the Red Sea (
|
||
<bibRefCitation id="EF834B37FFCF0729FCB08076D268FC35" author="Savigny" box="[873,1032,871,896]" pageId="6" pageNumber="323" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny 1822</bibRefCitation>
|
||
). Collected at El Tor under flattened grey coloured sponges, between corals (“
|
||
<emphasis id="B966EAD4FFCF0729FD5B809DD75EFC7D" italics="true" pageId="6" pageNumber="323">Sub tegmine gelatinoso-carneo-cinereo lineari lapidibus affini inter corallia ad Tor</emphasis>
|
||
”;
|
||
<bibRefCitation id="EF834B37FFCF0729FE8080BED781FC7D" author="Grube" box="[345,481,943,968]" pageId="6" pageNumber="323" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube 1869</bibRefCitation>
|
||
).. “
|
||
<emphasis id="B966EAD4FFCF0729FDD480BED760FC59" italics="true" pageId="6" pageNumber="323">Elle se déplace en serpentant avec beaucoup d’agilité et remuant continuellement ses cirres</emphasis>
|
||
” (
|
||
<bibRefCitation id="EF834B37FFCF0729FEC580C5D7DCFC59" author="Savigny" box="[284,444,980,1005]" pageId="6" pageNumber="323" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny 1822</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308654DFFCF0729FF1E80E6D757FBC8" pageId="6" pageNumber="323" type="etymology">
|
||
<paragraph id="8BAD36C6FFCF0729FF1E80E6D757FBC8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="323">
|
||
<emphasis id="B966EAD4FFCF0729FF1E80E6D726FBA5" bold="true" box="[199,326,1015,1040]" pageId="6" pageNumber="323">Etymology</emphasis>
|
||
. According to WoRMS (
|
||
<bibRefCitation id="EF834B37FFCF0729FD8780E6D4A4FBA5" author="Gil" box="[606,708,1015,1040]" pageId="6" pageNumber="323" refString="Gil, J. (2015) Syllis monilaris Savigny in Lamarck, 1818. In: Read, G. & Fauchald, K. (Eds.) World Polychaeta database. Available from: http: // www. marinespecies. org / polychaeta / aphia. php? p = taxdetails & id = 155116 (accessed 10 June 2015)" type="book chapter" year="2015">Gil 2015</bibRefCitation>
|
||
) the specific epithet
|
||
<taxonomicName id="4C124D45FFCF0729FC7480E6D278FBA5" box="[941,1048,1015,1040]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCF0729FC7480E6D278FBA5" box="[941,1048,1015,1040]" italics="true" pageId="6" pageNumber="323">monilaris</emphasis>
|
||
</taxonomicName>
|
||
is formed by the Latin root
|
||
<emphasis id="B966EAD4FFCF0729FA8A80E6D3FCFBA5" box="[1363,1436,1015,1040]" italics="true" pageId="6" pageNumber="323">monil-</emphasis>
|
||
, meaning "string of beads", and the Latin suffix
|
||
<emphasis id="B966EAD4FFCF0729FD63870CD48FFB81" box="[698,751,1053,1076]" italics="true" pageId="6" pageNumber="323">-aris</emphasis>
|
||
, which is used to form an adjective, usually by adding it to a noun, indicating a relationship or a pertaining to, referring to the presence of moniliform antennae and cirri typical of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308654DFFCF0727FF1E8796D4B7FE3D" lastPageId="8" lastPageNumber="325" pageId="6" pageNumber="323" type="discussion">
|
||
<paragraph id="8BAD36C6FFCF0728FF1E8796D448FED0" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="324" pageId="6" pageNumber="323">
|
||
<emphasis id="B966EAD4FFCF0729FF1E8796D754FB15" bold="true" box="[199,308,1159,1184]" pageId="6" pageNumber="323">Remarks</emphasis>
|
||
. The specimens studied by Grube agree with the descriptions and drawings provided by Savigny and Lamarck, and were collected in the same area (Gulf of Suez), so we have no doubt that they represent the same species. Since the
|
||
<typeStatus id="54A98864FFCF0729FEB587C0D7FCFB5C" box="[364,412,1233,1257]" pageId="6" pageNumber="323">type</typeStatus>
|
||
material used for the original description was lost, and that with the exception of
|
||
<bibRefCitation id="EF834B37FFCF0729FA8F87DED681FAB8" author="Grube" pageId="6" pageNumber="323" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube (1869)</bibRefCitation>
|
||
nobody redescribed the species, we designate here a
|
||
<typeStatus id="54A98864FFCF0729FC9A87E4D5FCFAB8" box="[835,924,1269,1293]" pageId="6" pageNumber="323" type="neotype">neotype</typeStatus>
|
||
and provide a redescription. Both specimens from the ZMB are in excellent condition, flexible, and well preserved, although most chaetae are now broken; however, specimens are very darkened, and both pharynx and proventricle are difficult to see by transparency; in order to preserve as undamaged as possible these two important specimens, we have not removed parapodia nor dissected them, so it is not possible to detail the number of muscle cell rows of the proventricle. Savigny’s description of
|
||
<taxonomicName id="4C124D45FFCF0729FEE086B6D787FA75" box="[313,487,1447,1472]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCF0729FEE086B6D787FA75" box="[313,487,1447,1472]" italics="true" pageId="6" pageNumber="323">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
is much more extensive than the one published in
|
||
<bibRefCitation id="EF834B37FFCF0729FBF986B6D2B7FA75" author="Lamarck" box="[1056,1239,1447,1472]" pageId="6" pageNumber="323" refString="Lamarck, J. B. (1818) Histoire Naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedes d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin l'exposition des principes fondamentaux de la zoologie. Vol. 5. Deterville, Paris, 612 pp." type="book" year="1818">Lamarck (1818)</bibRefCitation>
|
||
, but even so very incomplete, not detailing some important characters, especially the size and shape of the chaetae; the chaetae are described by
|
||
<bibRefCitation id="EF834B37FFCF0729FEF786FED7BCF9BD" author="Savigny" box="[302,476,1519,1544]" pageId="6" pageNumber="323" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny (1822)</bibRefCitation>
|
||
as “
|
||
<emphasis id="B966EAD4FFCF0729FDD686E0D51FF9BD" box="[527,895,1519,1544]" italics="true" pageId="6" pageNumber="323">assez grosses, obtuses, jaunâtres</emphasis>
|
||
” (“thick enough, obtuse, yellowish”), while the genus
|
||
<taxonomicName id="4C124D45FFCF0729FF388505D77DF998" box="[225,285,1556,1581]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFCF0729FF388505D77DF998" box="[225,285,1556,1581]" italics="true" pageId="6" pageNumber="323">Syllis</emphasis>
|
||
</taxonomicName>
|
||
is stated to have “
|
||
<emphasis id="B966EAD4FFCF0729FE2D8504D4E9F998" box="[500,649,1556,1581]" italics="true" pageId="6" pageNumber="323">soies simples</emphasis>
|
||
” (“simple chaetae”); in the drawings only the bases of the shafts are shown (see
|
||
<figureCitation id="13292A43FFCF0729FEFD8526D70AF9E5" box="[292,362,1591,1616]" captionStart="FIGURE 1" captionStartId="1.[151,250,1366,1388]" captionText="FIGURE 1. Syllis monilaris. Original plate by Savigny (1817)." pageId="6" pageNumber="323">Fig. 1</figureCitation>
|
||
). However,
|
||
<bibRefCitation id="EF834B37FFCF0729FE268526D4FBF9E5" author="Grube" box="[511,667,1591,1616]" pageId="6" pageNumber="323" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube (1869)</bibRefCitation>
|
||
describes the chaetae as “
|
||
<emphasis id="B966EAD4FFCF0729FC018526D3E9F9E5" box="[984,1417,1591,1616]" italics="true" pageId="6" pageNumber="323">falce plus minus brevi apice simplici</emphasis>
|
||
”, meaning that the chaetae did have a more or less short, simple, sickle-shaped apex. Savigny figured a dark, opaque, very long, complete specimen, with 341 segments and more than
|
||
<quantity id="4CEA9B23FFCF0729FCA4856ED5ADF92D" box="[893,973,1663,1688]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="323" unit="mm" value="76.0">76 mm</quantity>
|
||
long (from text; about
|
||
<quantity id="4CEA9B23FFCF0729FB0C8591D345F92D" box="[1237,1317,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="6" pageNumber="323" unit="mm" value="87.0">87 mm</quantity>
|
||
long from figure), and described to be grey-reddish coloured with some iridescence.
|
||
<bibRefCitation id="EF834B37FFCF0729FBCB85B5D2CFF908" author="Grube" box="[1042,1199,1700,1725]" pageId="6" pageNumber="323" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube (1869)</bibRefCitation>
|
||
reported about 300 segments, and we counted 296 chaetigers in the incomplete
|
||
<typeStatus id="54A98864FFCF0729FCE785D8D5FAF954" box="[830,922,1737,1761]" pageId="6" pageNumber="323" type="neotype">neotype</typeStatus>
|
||
. Both Grube’s and Savigny’s specimens have elongated dorsal cirri on anterior segments, becoming shorter, somewhat fusiform, on midbody and posterior segments, where they are shorter than body width (see
|
||
<figureCitation id="13292A43FFCF0729FD23841ED55EF89D" box="[762,830,1807,1832]" captionStart="FIGURE 4" captionStartId="8.[151,250,1846,1868]" captionTargetBox="[159,1413,433,1822]" captionTargetId="figure@8.[151,1436,413,1835]" captionTargetPageId="8" captionText="FIGURE 4. Syllis monilaris. Neotype (ZMB F 1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm." httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="6" pageNumber="323">Fig. 4</figureCitation>
|
||
). This arrangement of dorsal cirri is similar to those of
|
||
<taxonomicName id="4C124D45FFCF0729FF4E8425D7ADF8F9" authority="Grube, 1840" authorityName="Grube" authorityYear="1840" box="[151,461,1844,1869]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="gracilis">
|
||
<emphasis id="B966EAD4FFCF0729FF4E8425D752F8F8" box="[151,306,1844,1869]" italics="true" pageId="6" pageNumber="323">Syllis gracilis</emphasis>
|
||
<bibRefCitation id="EF834B37FFCF0729FEE58425D7ADF8F9" author="Grube" box="[316,461,1844,1869]" pageId="6" pageNumber="323" refString="Grube, A. E. (1840) Actinien, Echinodermen und Wurmer des Adriatischen- und Mittelmeers, nach eigenen Sammlungen beschrieben. J. H. Bon, Konigsberg, 92 pp., 1 pl." type="book" year="1840">Grube, 1840</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D45FFCF0729FE038425D517F8F8" authority="O.F. Muller, 1776" authorityName="O.F. Muller" authorityYear="1776" box="[474,887,1844,1869]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="armillaris">
|
||
<emphasis id="B966EAD4FFCF0729FE038425D4F0F8F8" box="[474,656,1844,1869]" italics="true" pageId="6" pageNumber="323">Syllis armillaris</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B37FFCF0729FD788425D50FF8F8" author="Muller" box="[673,879,1844,1869]" pageId="6" pageNumber="323" refString="Muller, O. F. (1776) Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae indigenarum characteres, nomina, et synonyma imprimis popularium. Hallageriis, Havniae (Copenhagen), xxxii + 274 pp. http: // dx. doi. org / 10.5962 / bhl. title. 13268" type="book" year="1776">O.F. Müller, 1776</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and, especially,
|
||
<taxonomicName id="4C124D45FFCF0729FBEE8425D30CF8F8" authority="Grube, 1863" authorityName="Grube" authorityYear="1863" box="[1079,1388,1844,1869]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="hyalina">
|
||
<emphasis id="B966EAD4FFCF0729FBEE8425D2B1F8F8" box="[1079,1233,1844,1869]" italics="true" pageId="6" pageNumber="323">Syllis hyalina</emphasis>
|
||
<bibRefCitation id="EF834B37FFCF0729FB028425D30CF8F8" author="Grube" box="[1243,1388,1844,1869]" pageId="6" pageNumber="323" refString="Grube, A. E. (1863) Beschreibung neuer oder wenig bekannter Anneliden. Sechster Beitrag. Archiv fur Naturgeschichte, 29, 37 - 69, pls. IV - VI." type="journal article" year="1863">Grube, 1863</bibRefCitation>
|
||
</taxonomicName>
|
||
; the general aspect of the body of
|
||
<taxonomicName id="4C124D45FFCF0729FE318446D4F8F8C5" box="[488,664,1879,1904]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCF0729FE318446D4F8F8C5" box="[488,664,1879,1904]" italics="true" pageId="6" pageNumber="323">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
is quite similar to the latter, but the compound chaetae are closer to those of
|
||
<taxonomicName id="4C124D45FFCF0729FF21846CD7E4F820" box="[248,388,1916,1941]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="armillaris">
|
||
<emphasis id="B966EAD4FFCF0729FF21846CD7E4F820" box="[248,388,1916,1941]" italics="true" pageId="6" pageNumber="323">S. armillaris</emphasis>
|
||
</taxonomicName>
|
||
, although clearly different. All these species have thick compound chaetae with short blades, which are fused with the shafts in the midbody parapodia only in
|
||
<taxonomicName id="4C124D45FFCF0729FCA984B0D581F80C" box="[880,993,1952,1977]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="gracilis">
|
||
<emphasis id="B966EAD4FFCF0729FCA984B0D581F80C" box="[880,993,1952,1977]" italics="true" pageId="6" pageNumber="323">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
. The three are probably closely related, since they also share the pentacerous reproductive stolon; however, none of the examined specimens of
|
||
<taxonomicName id="4C124D45FFCF0728FA8684D5D763FF05" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" lastPageId="7" lastPageNumber="324" order="Phyllodocida" pageId="6" pageNumber="323" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCF0728FA8684D5D763FF05" italics="true" lastPageId="7" lastPageNumber="324" pageId="6" pageNumber="323">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
were developing stolons, and none of them is useful for molecular analyses. This way, we cannot assure the phylogenetic relationships of
|
||
<taxonomicName id="4C124D45FFCE0728FDF983ACD4CDFF60" box="[544,685,188,213]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="324" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCE0728FDF983ACD4CDFF60" box="[544,685,188,213]" italics="true" pageId="7" pageNumber="324">S. monilaris</emphasis>
|
||
</taxonomicName>
|
||
with other species of
|
||
<taxonomicName id="4C124D45FFCE0728FC6583ADD599FF60" box="[956,1017,188,213]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="324" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFCE0728FC6583ADD599FF60" box="[956,1017,188,213]" italics="true" pageId="7" pageNumber="324">Syllis</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, the three species are reported worldwide and each one of them represents likely a complex of different species. Future researches will provide further details for the knowledge of
|
||
<taxonomicName id="4C124D45FFCE0728FD4C8214D57EFEA8" box="[661,798,260,285]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="324" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCE0728FD4C8214D57EFEA8" box="[661,798,260,285]" italics="true" pageId="7" pageNumber="324">S. monilaris</emphasis>
|
||
</taxonomicName>
|
||
(number of rows in the proventricle, dorsal and ventral simple capillary chaetae,
|
||
<typeStatus id="54A98864FFCE0728FE6A8239D783FEF5" box="[435,483,296,320]" pageId="7" pageNumber="324">type</typeStatus>
|
||
of stolon, molecular data), but the present description is sufficient and adequate for a correct identification of the species.
|
||
</paragraph>
|
||
<caption id="DF6D664EFFCE0728FF4E8478D201F8CA" box="[151,1121,1897,1919]" httpUri="https://zenodo.org/record/238851/files/figure.png" pageId="7" pageNumber="324" targetBox="[161,1428,419,1862]" targetPageId="7">
|
||
<paragraph id="8BAD36C6FFCE0728FF4E8478D201F8CA" blockId="7.[151,1121,1897,1919]" box="[151,1121,1897,1919]" pageId="7" pageNumber="324">
|
||
<emphasis id="B966EAD4FFCE0728FF4E8478D772F8CB" bold="true" box="[151,274,1897,1919]" pageId="7" pageNumber="324">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C124D45FFCE0728FEC08478D7D7F8CA" box="[281,439,1897,1919]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="324" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCE0728FEC08478D7D7F8CA" box="[281,439,1897,1919]" italics="true" pageId="7" pageNumber="324">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
. Neotype (ZMB F1941). Anterior end, dorsal view. Scale: 0.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD36C6FFCE0727FF1E84B7D4B7FE3D" blockId="7.[151,1436,1958,2019]" lastBlockId="8.[151,1437,151,392]" lastPageId="8" lastPageNumber="325" pageId="7" pageNumber="324">
|
||
Three specimens from
|
||
<collectingCountry id="F3057656FFCE0728FE0F84B7D453F80A" box="[470,563,1958,1983]" name="United Kingdom" pageId="7" pageNumber="324">England</collectingCountry>
|
||
and one from Beacon Island (Western
|
||
<collectingCountry id="F3057656FFCE0728FBD084B7D211F80A" box="[1033,1137,1958,1983]" name="Australia" pageId="7" pageNumber="324">Australia</collectingCountry>
|
||
) agree perfectly with the description of
|
||
<taxonomicName id="4C124D45FFCE0728FE9984DBD793F856" box="[320,499,1994,2019]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="324" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFCE0728FE9984DBD793F856" box="[320,499,1994,2019]" italics="true" pageId="7" pageNumber="324">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
, although being smaller. Whether they belong or not to the same species is not possible to assess at this moment.
|
||
<bibRefCitation id="EF834B37FFC10727FDCA8386D4CDFF05" author="Licher" box="[531,685,151,176]" pageId="8" pageNumber="325" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher (1999)</bibRefCitation>
|
||
reports this species from La Rochelle (
|
||
<collectingCountry id="F3057656FFC10727FBB88389D2D1FF05" box="[1121,1201,152,176]" name="France" pageId="8" pageNumber="325">France</collectingCountry>
|
||
) based on specimens collected by d’Orbigny (MNHN A–74,
|
||
<specimenCount id="9D14FD4FFFC10727FD8D83ADD410FF60" box="[596,624,188,213]" pageId="8" pageNumber="325" type="generic">15</specimenCount>
|
||
specimens), and also from Sicily, as
|
||
<taxonomicName id="4C124D45FFC10727FBC983ADD282FF60" box="[1040,1250,188,213]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="325" phylum="Annelida" rank="species" species="moniliformis">
|
||
<emphasis id="B966EAD4FFC10727FBC983ADD282FF60" box="[1040,1250,188,213]" italics="true" pageId="8" pageNumber="325">Syllis moniliformis</emphasis>
|
||
</taxonomicName>
|
||
(NHMW 621,
|
||
<specimenCount id="9D14FD4FFFC10727FA5783ADD3FCFF61" box="[1422,1436,188,212]" pageId="8" pageNumber="325" type="generic">3</specimenCount>
|
||
specimens). The question if the records of the species from Northern
|
||
<collectingCountry id="F3057656FFC10727FC4183F1D584FF42" box="[920,996,224,247]" name="France" pageId="8" pageNumber="325">France</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3057656FFC10727FBC083CED219FF4D" box="[1049,1145,223,248]" name="United Kingdom" pageId="8" pageNumber="325">England</collectingCountry>
|
||
, or the Mediterranean Sea represent sibling species or introduced populations remains unsolved until further detailed studies can be carried out. The Australian specimen however, comes from the same Ocean as
|
||
<taxonomicName id="4C124D45FFC10727FC1A8239D22BFEF5" box="[963,1099,295,320]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="325" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFC10727FC1A8239D22BFEF5" box="[963,1099,295,320]" italics="true" pageId="8" pageNumber="325">S. monilaris</emphasis>
|
||
</taxonomicName>
|
||
, although from a very distant locality. The present redescription will make the recognition of the species easier, and additional material will probably provide an answer to the above questions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6D664EFFC10727FF4E8427D585F8DE" httpUri="https://zenodo.org/record/238852/files/figure.png" pageId="8" pageNumber="325" targetBox="[159,1413,433,1822]" targetPageId="8">
|
||
<paragraph id="8BAD36C6FFC10727FF4E8427D585F8DE" blockId="8.[151,1436,1846,1899]" pageId="8" pageNumber="325">
|
||
<emphasis id="B966EAD4FFC10727FF4E8427D775F8F9" bold="true" box="[151,277,1846,1868]" pageId="8" pageNumber="325">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C124D45FFC10727FEC48427D7DEF8F9" box="[285,446,1846,1868]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="325" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFC10727FEC48427D7DEF8F9" box="[285,446,1846,1868]" italics="true" pageId="8" pageNumber="325">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
. Neotype (ZMB F1941). A, dorsal cirri, proventricular level. B, dorsal cirri, midbody. C, midposterior segments, dorsal view. D, posterior segments, dorsal view. Scale: 0.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6D664EFFC00726FF4E85ACD41DF947" httpUri="https://zenodo.org/record/238853/files/figure.png" pageId="9" pageNumber="326" targetBox="[193,1394,183,1698]" targetPageId="9">
|
||
<paragraph id="8BAD36C6FFC00726FF4E85ACD41DF947" blockId="9.[151,1435,1725,1778]" pageId="9" pageNumber="326">
|
||
<emphasis id="B966EAD4FFC00726FF4E85ACD774F966" bold="true" box="[151,276,1725,1747]" pageId="9" pageNumber="326">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C124D45FFC00726FEC285ACD7DBF966" box="[283,443,1725,1747]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFC00726FEC285ACD7DBF966" box="[283,443,1725,1747]" italics="true" pageId="9" pageNumber="326">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
. Neotype (ZMB F1941). Compound chaetae: A, anterior. B, midbody. C, posterior. Aciculae: D, anterior. E, midbody. F, posterior. Scale: 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C308654DFFC00725FF1E8408D6AFFADD" lastPageId="10" lastPageNumber="327" pageId="9" pageNumber="326" type="reference_group">
|
||
<paragraph id="8BAD36C6FFC00725FF1E8408D27FFEE2" blockId="9.[151,1436,1817,2009]" lastBlockId="10.[151,1437,151,1384]" lastPageId="10" lastPageNumber="327" pageId="9" pageNumber="326">
|
||
<taxonomicName id="4C124D45FFC00726FF1E8408D718F887" ID-CoL="7B73R" box="[199,376,1817,1842]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFC00726FF1E8408D718F887" box="[199,376,1817,1842]" italics="true" pageId="9" pageNumber="326">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
is the
|
||
<typeStatus id="54A98864FFC00726FE15840BD79CF887" box="[460,508,1818,1842]" pageId="9" pageNumber="326">type</typeStatus>
|
||
species of the
|
||
<typeStatus id="54A98864FFC00726FD77840BD4BEF887" box="[686,734,1818,1842]" pageId="9" pageNumber="326">type</typeStatus>
|
||
genus of the family
|
||
<taxonomicName id="4C124D45FFC00726FC0D8408D253F887" ID-CoL="84KTJ" box="[980,1075,1817,1842]" class="Polychaeta" family="Syllidae" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="family">Syllidae</taxonomicName>
|
||
, but it was poorly known, as a detailed description was lacking. This lack of knowledge caused numerous synonymies (see above list) and taxonomic problems. The most remarkable attempt to simplify the situation was done by
|
||
<bibRefCitation id="EF834B37FFC00726FB51844DD33FF8C0" author="Langerhans" box="[1160,1375,1884,1909]" pageId="9" pageNumber="326" refString="Langerhans, P. (1879) Die Wurmfauna von Madeira. Zeitschrift fur Wissenschaftliche Zoologie, 32, 513 - 592, pls. XXXI - XXXIII." type="journal article" year="1879">Langerhans (1879)</bibRefCitation>
|
||
, who split the genus into four subgenera:
|
||
<taxonomicName id="4C124D45FFC00726FD96846CD4AFF823" ID-CoL="84NYL" box="[591,719,1917,1942]" class="Polychaeta" family="Syllidae" genus="Haplosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC00726FD96846CD4AFF823" box="[591,719,1917,1942]" italics="true" pageId="9" pageNumber="326">Haplosyllis</emphasis>
|
||
</taxonomicName>
|
||
, only with simple chaetae;
|
||
<taxonomicName id="4C124D45FFC00726FBFA846CD2F2F823" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[1059,1170,1917,1942]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC00726FBFA846CD2F2F823" box="[1059,1170,1917,1942]" italics="true" pageId="9" pageNumber="326">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
, only with compound falcigerous chaetae;
|
||
<taxonomicName id="4C124D45FFC00726FE55848FD7A9F802" ID-CoL="9G24W" box="[396,457,1950,1975]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC00726FE55848FD7A9F802" box="[396,457,1950,1975]" italics="true" pageId="9" pageNumber="326">Syllis</emphasis>
|
||
</taxonomicName>
|
||
, with both compound falcigers and some secondarily simple, thick chaetae; and
|
||
<taxonomicName id="4C124D45FFC00726FF4E84D1D696F86C" ID-CoL="f0afee23-2530-46a8-937e-bd4bb18d10f5" box="[151,246,1984,2009]" class="Polychaeta" family="Syllidae" genus="Ehlersia" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="326" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC00726FF4E84D1D696F86C" box="[151,246,1984,2009]" italics="true" pageId="9" pageNumber="326">Ehlersia</emphasis>
|
||
</taxonomicName>
|
||
, with spinigerous and falcigerous compound chaetae; this division was followed by
|
||
<bibRefCitation id="EF834B37FFC00726FB1084D1D30AF86C" author="Fauvel" box="[1225,1386,1984,2009]" pageId="9" pageNumber="326" refString="Fauvel, P. (1923) Faune de France Vol. 5. Polychetes Errantes. Librairie de la Faculte des Sciences Paul Le Chevalier, Paris, 488 pp." type="book" year="1923">Fauvel (1923)</bibRefCitation>
|
||
and subsequent numerous different systems of classification, depending on the authors, who considered them as subgenera or raised them to genera.
|
||
<bibRefCitation id="EF834B37FFC30725FDEE83A8D563FF64" author="San" box="[567,771,185,209]" pageId="10" pageNumber="327" refString="San Martin, G. (1984) Estudio biogeografico, faunistico y sistematico de los Poliquetos de la familia Silidos (Syllidae: Polychaeta) en Baleares. Ediciones de la Universidad Complutense de Madrid, Madrid, 187, 1 - 529." type="journal article" year="1984">San Martín (1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FCC983A8D52BFF64" author="Martin" box="[784,843,185,209]" pageId="10" pageNumber="327" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Polychaeta: Syllidae: Syllinae) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FC8183A8D5F2FF64" author="Martin" box="[856,914,185,209]" pageId="10" pageNumber="327" refString="San Martin, G. (2003) Annelida Polychaeta II. Syllidae. Fauna Iberica, 21, 554 pp. [Museo Nacional de Ciencias Naturales. CSIC, Madrid]" type="book" year="2003">2003</bibRefCitation>
|
||
) offered a detailed discussion and revision of this problem, and with the exception of
|
||
<taxonomicName id="4C124D45FFC30725FD8F83CBD4B5FF46" ID-CoL="84NYL" box="[598,725,218,243]" class="Polychaeta" family="Syllidae" genus="Haplosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FD8F83CBD4B5FF46" box="[598,725,218,243]" italics="true" pageId="10" pageNumber="327">Haplosyllis</emphasis>
|
||
</taxonomicName>
|
||
, considered as a genus
|
||
<emphasis id="B966EAD4FFC30725FC0683CDD242FF46" box="[991,1058,220,243]" italics="true" pageId="10" pageNumber="327">per se</emphasis>
|
||
, proposed the combination of the remaining three genera under the genus
|
||
<taxonomicName id="4C124D45FFC30725FDA183EDD4D4FEA0" ID-CoL="9G24W" box="[632,692,252,277]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FDA183EDD4D4FEA0" box="[632,692,252,277]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
, at least until more species were better known and more data available: “[...]
|
||
<emphasis id="B966EAD4FFC30725FE95820CD791FE83" box="[332,497,285,310]" italics="true" pageId="10" pageNumber="327">the division of</emphasis>
|
||
<taxonomicName id="4C124D45FFC30725FE22820CD45BFE83" ID-CoL="9G24W" box="[507,571,285,310]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">Syllis</taxonomicName>
|
||
[...]
|
||
<emphasis id="B966EAD4FFC30725FDAE820CD567FEE3" italics="true" pageId="10" pageNumber="327">both as subgenera or genera, is an artificial division and it has been followed for practical reasons more than scientific ones</emphasis>
|
||
[...]” (
|
||
<bibRefCitation id="EF834B37FFC30725FC88822ED272FEE2" author="San" box="[849,1042,319,343]" pageId="10" pageNumber="327" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Polychaeta: Syllidae: Syllinae) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín 1992</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFC30725FF1E824ED7E2FDBD" blockId="10.[151,1437,151,1384]" pageId="10" pageNumber="327">
|
||
Furthermore,
|
||
<bibRefCitation id="EF834B37FFC30725FEB18271D425FECD" box="[360,581,352,376]" pageId="10" pageNumber="327" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Polychaeta: Syllidae: Syllinae) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article">San Martín (1992)</bibRefCitation>
|
||
stated that “
|
||
<emphasis id="B966EAD4FFC30725FD3E824ED283FECD" box="[743,1251,351,376]" italics="true" pageId="10" pageNumber="327">All these considerations do not mean that</emphasis>
|
||
<taxonomicName id="4C124D45FFC30725FB28824ED351FECD" ID-CoL="9G24W" box="[1265,1329,351,376]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">Syllis</taxonomicName>
|
||
<emphasis id="B966EAD4FFC30725FAE78270D537FE75" italics="true" pageId="10" pageNumber="327">must be considered as a homogeneous genus. There are many other characters, which have not been described for most of the species of this genus, but could be useful for segregating</emphasis>
|
||
<taxonomicName id="4C124D45FFC30725FCB882B6D5C1FE75" ID-CoL="9G24W" box="[865,929,423,448]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">Syllis</taxonomicName>
|
||
<emphasis id="B966EAD4FFC30725FC7282B8D70FFDBD" italics="true" pageId="10" pageNumber="327">into groups of species or subgenera. These characters are mainly three: the shape of posterior aciculae, shape of the solitary dorsal setae, and the kind of reproductive stolon</emphasis>
|
||
.”
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFC30725FF1E8105D7A7FC69" blockId="10.[151,1437,151,1384]" pageId="10" pageNumber="327">
|
||
Some recent authors (e.g.
|
||
<bibRefCitation id="EF834B37FFC30725FE358105D417FD99" author="Licher" box="[492,631,532,557]" pageId="10" pageNumber="327" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FD5A8105D5A7FD99" author="Tovar-Hernandez" box="[643,967,532,557]" pageId="10" pageNumber="327" refString="Tovar-Hernandez, M. A., Granados-Barba, A. & Solis-Weiss, V. (2002) Typosyllis papillosus, a new species (Annelida: Polychaeta: Syllidae) from the southwest Gulf of Mexico. Proceedings of the Biological Society of Washington, 115 (4), 760 - 768." type="journal article" year="2002">
|
||
Tovar-Hernández
|
||
<emphasis id="B966EAD4FFC30725FC958104D51EFD98" box="[844,894,532,557]" italics="true" pageId="10" pageNumber="327">et al</emphasis>
|
||
. 2002
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FC0B8105D2CCFD98" author="Aguado" box="[978,1196,532,557]" pageId="10" pageNumber="327" refString="Aguado, M. T., Helm, C., Weidhase, M. & Bleidorn, C. (2015) Description of a new syllid species as a model for evolutionary research of reproduction and regeneration in annelids. Organisms Diversity & Evolution, 15 (1), 1 - 21. http: // dx. doi. org / 10.1007 / s 13127 - 014 - 0183 - 5" type="journal article" year="2015">
|
||
Aguado
|
||
<emphasis id="B966EAD4FFC30725FBEB8104D203FD98" box="[1074,1123,532,557]" italics="true" pageId="10" pageNumber="327">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
) have maintained the existence of two genera:
|
||
<taxonomicName id="4C124D45FFC30725FE6B8126D78FFDE5" ID-CoL="9G24W" box="[434,495,567,592]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FE6B8126D78FFDE5" box="[434,495,567,592]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
, with some simple chaetae in midbody resulting from the fusion of the blades with the shafts, and
|
||
<taxonomicName id="4C124D45FFC30725FE99814DD7CFFDC0" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[320,431,604,629]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FE99814DD7CFFDC0" box="[320,431,604,629]" italics="true" pageId="10" pageNumber="327">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
, with the rest of the species, including those with thick simple chaetae resulting from the loss of blades, and the ones with pseudospinigers (previously considered as spinigers). This contradicts the opinion expressed by
|
||
<bibRefCitation id="EF834B37FFC30725FEEA81B5D79FFD09" author="San" box="[307,511,676,700]" pageId="10" pageNumber="327" refString="San Martin, G. (1984) Estudio biogeografico, faunistico y sistematico de los Poliquetos de la familia Silidos (Syllidae: Polychaeta) en Baleares. Ediciones de la Universidad Complutense de Madrid, Madrid, 187, 1 - 529." type="journal article" year="1984">San Martín (1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FDD581B5D427FD09" author="Martin" box="[524,583,676,700]" pageId="10" pageNumber="327" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Polychaeta: Syllidae: Syllinae) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B37FFC30725FD8D81B5D4F0FD09" author="Martin" box="[596,656,676,700]" pageId="10" pageNumber="327" refString="San Martin, G. (2003) Annelida Polychaeta II. Syllidae. Fauna Iberica, 21, 554 pp. [Museo Nacional de Ciencias Naturales. CSIC, Madrid]" type="book" year="2003">2003</bibRefCitation>
|
||
), in that “
|
||
<emphasis id="B966EAD4FFC30725FCDA81B5D69FFD55" italics="true" pageId="10" pageNumber="327">the presence of thick simple setae, which would define the subgenus</emphasis>
|
||
<taxonomicName id="4C124D45FFC30725FEDC81D6D725FD55" ID-CoL="9G24W" box="[261,325,711,736]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">Syllis</taxonomicName>
|
||
,
|
||
<emphasis id="B966EAD4FFC30725FE8A81D8D363FD55" box="[339,1283,711,736]" italics="true" pageId="10" pageNumber="327">cannot be used as a generic or subgeneric character, but only as a specific character</emphasis>
|
||
” (
|
||
<bibRefCitation id="EF834B37FFC30725FAC781D9D6B3FCB1" author="San" pageId="10" pageNumber="327" refString="San Martin, G. (1992) Syllis Savigny in Lamarck, 1818 (Polychaeta: Syllidae: Syllinae) from Cuba, the Gulf of Mexico, Florida and North Carolina, with a revision of several species described by Verrill. Bulletin of Marine Science, 51 (2), 167 - 196." type="journal article" year="1992">San Martín 1992</bibRefCitation>
|
||
). The fusion of blades and shafts is progressive during the ontogeny, and younger specimens of
|
||
<taxonomicName id="4C124D45FFC30725FB2581FDD3F4FCB0" ID-CoL="53MT5" box="[1276,1428,748,773]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="species" species="gracilis">
|
||
<emphasis id="B966EAD4FFC30725FB2581FDD3F4FCB0" box="[1276,1428,748,773]" italics="true" pageId="10" pageNumber="327">Syllis gracilis</emphasis>
|
||
</taxonomicName>
|
||
, for instance, do not have fused chaetae, and they become more markedly fused as the specimen grows; the presence of thick simple chaetae is thus a secondary character. This species would be a special case in the Animal Kingdom, changing of genus during its ontogeny, being
|
||
<taxonomicName id="4C124D45FFC30725FD488046D560FCC5" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[657,768,855,880]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FD488046D560FCC5" box="[657,768,855,880]" italics="true" pageId="10" pageNumber="327">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
during the early stages, and
|
||
<taxonomicName id="4C124D45FFC30725FBE78046D21AFCC5" ID-CoL="9G24W" box="[1086,1146,855,880]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FBE78046D21AFCC5" box="[1086,1146,855,880]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
when adult. Furthermore, there are some other similar species with only partially fused chaetae, which would not fit clearly in the two genera. There are two more considerations supporting
|
||
<taxonomicName id="4C124D45FFC30725FD6D808ED543FC0D" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[692,803,927,952]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FD6D808ED543FC0D" box="[692,803,927,952]" italics="true" pageId="10" pageNumber="327">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
as a non-valid genus, and just one more among many other synonymies of
|
||
<taxonomicName id="4C124D45FFC30725FE5B80D5D7DEFC68" ID-CoL="9G24W" box="[386,446,964,989]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FE5B80D5D7DEFC68" box="[386,446,964,989]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFC30725FF1E80F9D234FB91" blockId="10.[151,1437,151,1384]" pageId="10" pageNumber="327">
|
||
1. The
|
||
<typeStatus id="54A98864FFC30725FECC80F9D725FBB5" box="[277,325,1000,1024]" pageId="10" pageNumber="327">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="4C124D45FFC30725FDED80F6D411FBB5" ID-CoL="9G24W" box="[564,625,999,1024]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FDED80F6D411FBB5" box="[564,625,999,1024]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D45FFC30725FDA680F6D54DFBB5" ID-CoL="7B73R" box="[639,813,999,1024]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="species" species="monilaris">
|
||
<emphasis id="B966EAD4FFC30725FDA680F6D54DFBB5" box="[639,813,999,1024]" italics="true" pageId="10" pageNumber="327">Syllis monilaris</emphasis>
|
||
</taxonomicName>
|
||
, does not have fused chaetae, contrary to what is stated in the diagnosis of the genus by
|
||
<bibRefCitation id="EF834B37FFC30725FDD9871DD4CEFB91" author="Savigny" box="[512,686,1036,1061]" pageId="10" pageNumber="327" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny (1822)</bibRefCitation>
|
||
(see above redescription of
|
||
<typeStatus id="54A98864FFC30725FC33871DD228FB91" box="[1002,1096,1036,1060]" pageId="10" pageNumber="327" type="neotype">neotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFC30725FF1E8721D717FBD8" blockId="10.[151,1437,151,1384]" pageId="10" pageNumber="327">
|
||
2. There are several older genera synonymized with
|
||
<taxonomicName id="4C124D45FFC30725FCC1873ED534FBFD" ID-CoL="9G24W" box="[792,852,1071,1096]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FCC1873ED534FBFD" box="[792,852,1071,1096]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
that would have priority over
|
||
<taxonomicName id="4C124D45FFC30725FB77873ED37DFBFD" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[1198,1309,1071,1096]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FB77873ED37DFBFD" box="[1198,1309,1071,1096]" italics="true" pageId="10" pageNumber="327">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
(see list of synonymies above).
|
||
</paragraph>
|
||
<paragraph id="8BAD36C6FFC30725FF1E8768D6AFFADD" blockId="10.[151,1437,151,1384]" pageId="10" pageNumber="327">
|
||
Two of the authors (P.A.-C. and G.S.M.) are currently revising the available
|
||
<typeStatus id="54A98864FFC30725FB878768D2F9FB24" box="[1118,1177,1145,1169]" pageId="10" pageNumber="327">types</typeStatus>
|
||
of each one of these synonymies, and preparing redescriptions of some older species, in the framework of a global revision of the systematics of the genus, supported by molecular data whenever available (Álvarez-Campos
|
||
<emphasis id="B966EAD4FFC30725FB3C87D0D343FB6D" box="[1253,1315,1215,1240]" italics="true" pageId="10" pageNumber="327">et al.</emphasis>
|
||
in press). Considering the possible future division of
|
||
<taxonomicName id="4C124D45FFC30725FD5F87F5D4A2FB48" ID-CoL="9G24W" box="[646,706,1252,1277]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FD5F87F5D4A2FB48" box="[646,706,1252,1277]" italics="true" pageId="10" pageNumber="327">Syllis</emphasis>
|
||
</taxonomicName>
|
||
into several genera as a result of the ongoing studies, the name
|
||
<taxonomicName id="4C124D45FFC30725FF4E8616D767FA95" ID-CoL="ad03e8cb-d5d7-47dd-88ed-5c19b5f233a1" box="[151,263,1287,1312]" class="Polychaeta" family="Syllidae" genus="Typosyllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="genus">
|
||
<emphasis id="B966EAD4FFC30725FF4E8616D767FA95" box="[151,263,1287,1312]" italics="true" pageId="10" pageNumber="327">Typosyllis</emphasis>
|
||
</taxonomicName>
|
||
could only be applied for its
|
||
<typeStatus id="54A98864FFC30725FD838618D4EAFA94" box="[602,650,1289,1313]" pageId="10" pageNumber="327">type</typeStatus>
|
||
species (
|
||
<taxonomicName id="4C124D45FFC30725FD2F8616D244FA95" ID-CoL="53MTT" authority="Ehlers, 1864" authorityName="Ehlers" authorityYear="1864" box="[758,1060,1287,1312]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="327" phylum="Annelida" rank="species" species="krohnii">
|
||
<emphasis id="B966EAD4FFC30725FD2F8616D5EBFA95" box="[758,907,1287,1312]" italics="true" pageId="10" pageNumber="327">Syllis krohnii</emphasis>
|
||
<bibRefCitation id="EF834B37FFC30725FC4D8616D244FA95" author="Ehlers" box="[916,1060,1287,1312]" pageId="10" pageNumber="327" refString="Ehlers, E. (1864) Die Borstenwurmer (Annelida Chaetopoda) nach Systematischen und anatomischen Untersuchungen. Wilhelm Engelmann, Leipzig, 270 pp. http: // dx. doi. org / 10.5962 / bhl. title. 2081" type="book" year="1864">Ehlers, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
) and closely related species, and only in the case there would be no available previous name to designate the group, according to the ICZN article 23.3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |