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<mods:title id="AE8A6411F1807E9E90945D92EF439A3E">Jassa (Crustacea: Amphipoda): a new morphological and molecular assessment of the genus</mods:title>
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<mods:namePart id="3B1A87B238A2832D572ADE109311F61F">Conlan, Kathleen E.</mods:namePart>
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<treatment id="03B487DAFFFFD956C9C8193BFD3FF8B2" ID-DOI="http://doi.org/10.5281/zenodo.4580566" ID-GBIF-Taxon="181871891" ID-Zenodo-Dep="4580566" LSID="urn:lsid:plazi:treatment:03B487DAFFFFD956C9C8193BFD3FF8B2" httpUri="http://treatment.plazi.org/id/03B487DAFFFFD956C9C8193BFD3FF8B2" lastPageId="78" lastPageNumber="79" pageId="74" pageNumber="75">
<subSubSection id="C3076547FFFFD952C9C8193BFE61FD91" pageId="74" pageNumber="75" type="nomenclature">
<paragraph id="8BA236CCFFFFD952C9C8193BFE3AFDBA" blockId="74.[151,462,553,616]" box="[151,462,553,580]" pageId="74" pageNumber="75">
<heading id="D0EA81A0FFFFD952C9C8193BFE3AFDBA" bold="true" box="[151,462,553,580]" fontSize="11" level="2" pageId="74" pageNumber="75" reason="4">
<taxonomicName id="4C1D4D4FFFFFD952C9C8193BFE3AFDBA" ID-CoL="3QN24" authority="(Sars, 1894)" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[151,462,553,580]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFFD952C9C8193BFE3AFDBA" bold="true" box="[151,462,553,580]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9C8193BFEC7FDBD" bold="true" box="[151,307,553,580]" italics="true" pageId="74" pageNumber="75">Jassapusilla</emphasis>
(
<bibRefCitation id="EF8C4B3DFFFFD952C81C1938FE32FDBA" author="Sars, G. O." box="[323,454,553,579]" pageId="74" pageNumber="75" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA236CCFFFFD952C9C8195CFE61FD91" blockId="74.[151,462,553,616]" box="[151,405,589,616]" pageId="74" pageNumber="75">
(
<tableCitation id="C69F0377FFFFD952C9C0195CFEF6FD91" box="[159,258,589,616]" captionStart="TABLE 11" captionStartId="73.[151,239,225,249]" captionTargetPageId="73" captionText="TABLE 11. Key character states of locally distributed North Atlantic species. Trans-hemispheric species (Table 10) may occur there as well. X indicates the presence of the first character state described while — indicates the alternative character state given in italics. S and L indicate length.? indicates that the state is unknown. Character states apply to both sexes and juveniles unless indicated otherwise." httpUri="http://table.plazi.org/id/DF626644FFFCD951C9C81BF0FD79FE9C" pageId="74" pageNumber="75" tableUuid="DF626644FFFCD951C9C81BF0FD79FE9C">Table 11</tableCitation>
,
<figureCitation id="13262A49FFFFD952C851195CFE78FD91" box="[270,396,589,616]" captionStart-0="FIGURE 37" captionStart-1="FIGURE 38" captionStart-2="FIGURE 39" captionStart-3="FIGURE 40" captionStart-4="FIGURE 41" captionStartId-0="75.[151,250,1818,1842]" captionStartId-1="76.[151,250,1828,1852]" captionStartId-2="77.[151,250,1898,1922]" captionStartId-3="78.[151,250,1123,1147]" captionStartId-4="79.[151,250,1868,1892]" captionTargetBox-0="[173,1416,181,1786]" captionTargetBox-1="[228,1348,182,1794]" captionTargetBox-2="[163,1430,187,1869]" captionTargetBox-3="[159,1427,209,1092]" captionTargetBox-4="[159,1436,181,1844]" captionTargetId-0="figure-18@75.[170,1416,181,1794]" captionTargetId-1="figure-22@76.[219,1367,181,1804]" captionTargetId-2="figure-18@77.[151,1436,181,1874]" captionTargetId-3="figure-22@78.[151,1436,201,1100]" captionTargetId-4="figure-18@79.[151,1436,181,1844]" captionTargetPageId-0="75" captionTargetPageId-1="76" captionTargetPageId-2="77" captionTargetPageId-3="78" captionTargetPageId-4="79" captionText-0="FIGURE 37. Jassa pusilla (Sars, 1894). Paralectotype, adult female 1, about 4.5 mm, UiO F.10562 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 as Podocerus pusillus Sars, 1894. Adult female 2, body length unknown, UiO F.10564 Mp 41. Locality, date and collector unknown, illustrated in Sars (1894), plate 213 as Podocerus pusillus Sars, 1894. Adult female 3, 4.7 mm, UiO F.13359. Riŝr, Norway (approx. 59°N, 9°E), date unknown, G. O. Sars, coll. Lateral view: whole body, gnathopods 1 and 2 and pereopod 3; dorsal view: telson; medial view: uropod 3. Scale 0.1 mm. Illustration after Conlan (1990)." captionText-1="FIGURE 38. Jassa pusilla (Sars, 1894). Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus odontonyx Sars, 1894, submerged by Conlan (1990)). Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus pusillus Sars, 1894, transferred to Jassa pusilla by Stebbing, 1906). Medial views. Scale 0.1 mm." captionText-2="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." captionText-3="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." captionText-4="FIGURE 41. Jassa pusilla (Sars, 1894).Adult female 3 from FIGURE 37, 4.7 mm,, UiO F.13359. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll. Mouthparts. Frontal view: upper lip; lateral views: maxilla 1 and right and left mandibles; other views medial. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4580730" figureDoi-1="http://doi.org/10.5281/zenodo.4580732" figureDoi-2="http://doi.org/10.5281/zenodo.4580734" figureDoi-3="http://doi.org/10.5281/zenodo.4580736" figureDoi-4="http://doi.org/10.5281/zenodo.4580738" httpUri-0="https://zenodo.org/record/4580730/files/figure.png" httpUri-1="https://zenodo.org/record/4580732/files/figure.png" httpUri-2="https://zenodo.org/record/4580734/files/figure.png" httpUri-3="https://zenodo.org/record/4580736/files/figure.png" httpUri-4="https://zenodo.org/record/4580738/files/figure.png" pageId="74" pageNumber="75">Figs 3741</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3076547FFFFD952C9C81983FE24FD53" box="[151,464,658,682]" pageId="74" pageNumber="75" type="description">
<paragraph id="8BA236CCFFFFD952C9C81983FE24FD53" blockId="74.[151,464,658,682]" box="[151,464,658,682]" pageId="74" pageNumber="75">
Synonyms: see
<bibRefCitation id="EF8C4B3DFFFFD952C8691983FE39FD53" author="Conlan, K. E." box="[310,461,658,682]" pageId="74" pageNumber="75" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3076547FFFFD952C9C819C9FC33FA3A" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="8BA236CCFFFFD952C9C819C9FEE6FD0B" blockId="74.[151,1437,728,2014]" box="[151,274,728,754]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9C819C9FEE6FD0B" bold="true" box="[151,274,728,754]" pageId="74" pageNumber="75">Diagnosis.</emphasis>
</paragraph>
<paragraph id="8BA236CCFFFFD952C99819EDFEBAFCEF" blockId="74.[151,1437,728,2014]" box="[199,334,764,790]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C99819EDFEBAFCEF" bold="true" box="[199,334,764,790]" pageId="74" pageNumber="75">Both sexes:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981831FC42FCC2" blockId="74.[151,1437,728,2014]" box="[199,950,800,827]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981831FE72FCC3" box="[199,390,800,826]" italics="true" pageId="74" pageNumber="75">Mandibular palp</emphasis>
: article 2, dorsal margin without a fringe of setae.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981855FC19FCA7" blockId="74.[151,1437,728,2014]" box="[199,1005,836,863]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981855FEC7FCA7" box="[199,307,836,862]" italics="true" pageId="74" pageNumber="75">Maxilla 1</emphasis>
: without a seta or setal cluster at the base of the palp article 1.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981879FD4AFC5F" blockId="74.[151,1437,728,2014]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981879FEAEFC7B" box="[199,346,872,898]" italics="true" pageId="74" pageNumber="75">Gnathopod 1</emphasis>
: basis, anterolateral margin with a few very short setae; carpus without a single or small cluster of setae at the anterodistal junction of the propodus.
</paragraph>
<paragraph id="8BA236CCFFFFD952C99818A1FE7FFBEA" blockId="74.[151,1437,728,2014]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C99818A1FEAFFC33" box="[199,347,944,970]" italics="true" pageId="74" pageNumber="75">Gnathopod 2</emphasis>
: basis with a few minute setae along the anterolateral margin but without long filter setae (most setal lengths &lt;10% of the basis width); carpus and propodus, setae on the anterior margin short and simple (setal length &lt;basis width).
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981F0DFD09FBCF" blockId="74.[151,1437,728,2014]" box="[199,765,1052,1079]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981F0DFE9BFBCF" box="[199,367,1052,1078]" italics="true" pageId="74" pageNumber="75">Pereopods 57</emphasis>
: propodus not expanded anteriorly.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981F51FB59FBA2" blockId="74.[151,1437,728,2014]" box="[199,1197,1088,1115]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981F51FEC6FBA3" box="[199,306,1088,1114]" italics="true" pageId="74" pageNumber="75">Uropod 1</emphasis>
: ventral peduncular spinous process underlying about 1/4 of the longest ramus.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981F75FB64FB86" blockId="74.[151,1437,728,2014]" box="[199,1168,1124,1151]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981F75FEC6FB87" box="[199,306,1124,1150]" italics="true" pageId="74" pageNumber="75">Uropod 3</emphasis>
: inner ramus without spines mid-dorsally (with only the single apical spine).
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981F99FB99FB5A" blockId="74.[151,1437,728,2014]" box="[199,1133,1160,1187]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981F99FEFBFB5B" box="[199,271,1160,1186]" italics="true" pageId="74" pageNumber="75">Telson</emphasis>
: tip without apical setae, only the usual short setae at each dorsolateral cusp.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981FBDFE72FB3F" blockId="74.[151,1437,728,2014]" box="[199,390,1196,1222]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981FBDFE72FB3F" bold="true" box="[199,390,1196,1222]" pageId="74" pageNumber="75">Thumbed male:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981FC1FBEAFB12" blockId="74.[151,1437,728,2014]" box="[199,1054,1232,1259]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981FC1FECEFB13" box="[199,314,1232,1258]" italics="true" pageId="74" pageNumber="75">Antenna 2</emphasis>
: without plumose setae on the flagellum and peduncular article 5.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981FE5FEECFAAF" blockId="74.[151,1437,728,2014]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981FE5FEAEFAF7" box="[199,346,1268,1294]" italics="true" pageId="74" pageNumber="75">Gnathopod 2</emphasis>
: propodus, palmar defining spines absent except in small males. Thumb distally acute or squared in minor males and indented in major males. Dactyl expansion variable, centrally toothed or shallowly expanded proximally.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981E71FE9FFA83" blockId="74.[151,1437,728,2014]" box="[199,363,1376,1402]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981E71FE9FFA83" bold="true" box="[199,363,1376,1402]" pageId="74" pageNumber="75">Adult female:</emphasis>
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981E95FBEAFA66" blockId="74.[151,1437,728,2014]" box="[199,1054,1412,1439]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981E95FECEFA67" box="[199,314,1412,1438]" italics="true" pageId="74" pageNumber="75">Antenna 2</emphasis>
: without plumose setae on the flagellum and peduncular article 5.
</paragraph>
<paragraph id="8BA236CCFFFFD952C9981EB9FC33FA3A" blockId="74.[151,1437,728,2014]" box="[199,967,1448,1475]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981EB9FEAEFA3B" box="[199,346,1448,1474]" italics="true" pageId="74" pageNumber="75">Gnathopod 2</emphasis>
: propodus, palm concave, palmar defining angle acute.
</paragraph>
</subSubSection>
<subSubSection id="C3076547FFFFD956C9981EDDFD3FF8B2" lastPageId="78" lastPageNumber="79" pageId="74" pageNumber="75" type="discussion">
<paragraph id="8BA236CCFFFFD952C9981EDDFDF0F827" blockId="74.[151,1437,728,2014]" pageId="74" pageNumber="75">
<emphasis id="B969EADEFFFFD952C9981EDDFECFFA1F" bold="true" box="[199,315,1484,1510]" pageId="74" pageNumber="75">Remarks.</emphasis>
The shape of the male second gnathopod is highly variable but
<figureCitation id="13262A49FFFFD952CD5A1EDDFBA0FA1F" box="[1029,1108,1484,1510]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Fig. 39</figureCitation>
shows how shapes can grade within a series, this being a single collection made by G. O. Sars from Riŝr,
<collectingCountry id="F30A765CFFFFD952CAAC1EE0FBBBF9F2" box="[1011,1103,1521,1547]" name="Norway" pageId="74" pageNumber="75">Norway</collectingCountry>
. Juvenile males that approach the size of the thumbed males have pre-thumbs that are small relative to minor and major forms (
<figureCitation id="13262A49FFFFD952CDBE1D05FA68F9D7" box="[1249,1436,1556,1583]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Figs 39A, Band</figureCitation>
<figureCitation id="13262A49FFFFD952C9C81D29FF43F9AB" box="[151,183,1592,1618]" captionStart="FIGURE 40" captionStartId="78.[151,250,1123,1147]" captionTargetBox="[159,1427,209,1092]" captionTargetId="figure-22@78.[151,1436,201,1100]" captionTargetPageId="78" captionText="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="74" pageNumber="75">40</figureCitation>
). The pre-thumb is located in the distal half of the palm. Those juveniles that are about to molt into a thumbed stage (showing a thumbed cuticle under the juvenile cuticle) are termed subadult in
<figureCitation id="13262A49FFFFD952CD181D4DFB6EF98F" box="[1095,1178,1628,1654]" captionStart="FIGURE 40" captionStartId="78.[151,250,1123,1147]" captionTargetBox="[159,1427,209,1092]" captionTargetId="figure-22@78.[151,1436,201,1100]" captionTargetPageId="78" captionText="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="74" pageNumber="75">Fig. 40</figureCitation>
. These have small prethumbs as well. Minor forms are interpreted as being those that have thumbs that are also in the distal half of the palm (as in minor forms of other species) but that have a larger and longer thumb and never show a thumbed cuticle inside (i.e., will not molt again) (
<figureCitation id="13262A49FFFFD952C8A01DD9FDA6F91B" box="[511,594,1736,1762]" captionStart="FIGURE 38" captionStartId="76.[151,250,1828,1852]" captionTargetBox="[228,1348,182,1794]" captionTargetId="figure-22@76.[219,1367,181,1804]" captionTargetPageId="76" captionText="FIGURE 38. Jassa pusilla (Sars, 1894). Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus odontonyx Sars, 1894, submerged by Conlan (1990)). Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus pusillus Sars, 1894, transferred to Jassa pusilla by Stebbing, 1906). Medial views. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580732" httpUri="https://zenodo.org/record/4580732/files/figure.png" pageId="74" pageNumber="75">Figs 38</figureCitation>
and
<figureCitation id="13262A49FFFFD952CBD91DD9FD57F91B" box="[646,675,1736,1762]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">39</figureCitation>
CE). Minor forms also have centrally toothed dactyls (
<figureCitation id="13262A49FFFFD952CC581DD9FAAFF91B" box="[1287,1371,1736,1762]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Figs 39</figureCitation>
CE). Major form males have longer thumbs that are apically indented at the tip. Their thumbs originate in the proximal half of the propodus (
<figureCitation id="13262A49FFFFD952C8D61C01FE2AF8D3" box="[393,478,1808,1834]" captionStart="FIGURE 38" captionStartId="76.[151,250,1828,1852]" captionTargetBox="[228,1348,182,1794]" captionTargetId="figure-22@76.[219,1367,181,1804]" captionTargetPageId="76" captionText="FIGURE 38. Jassa pusilla (Sars, 1894). Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus odontonyx Sars, 1894, submerged by Conlan (1990)). Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus pusillus Sars, 1894, transferred to Jassa pusilla by Stebbing, 1906). Medial views. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580732" httpUri="https://zenodo.org/record/4580732/files/figure.png" pageId="74" pageNumber="75">Figs 38</figureCitation>
and
<figureCitation id="13262A49FFFFD952CB4A1C01FDC6F8D3" box="[533,562,1808,1834]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">39</figureCitation>
FJ). The dactyl expansion is shallow and proximal in most major forms (
<figureCitation id="13262A49FFFFD952CC301C01FF4BF8B7" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Fig. 39J</figureCitation>
) but is centrally toothed in some (
<figureCitation id="13262A49FFFFD952CB1E1C25FD5EF8B7" box="[577,682,1844,1870]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Fig. 39G</figureCitation>
). Sars originally named the major form
<taxonomicName id="4C1D4D4FFFFFD952CD331C25FAB7F8B7" authorityName="Sars" authorityYear="1894" box="[1132,1347,1844,1870]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B969EADEFFFFD952CD331C25FAB7F8B7" box="[1132,1347,1844,1870]" italics="true" pageId="74" pageNumber="75">Podocerus pusillus</emphasis>
</taxonomicName>
and the minor form
<taxonomicName id="4C1D4D4FFFFFD952C8411C49FDE5F88B" authorityName="Sars" authorityYear="1894" box="[286,529,1880,1906]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="odontonyx">
<emphasis id="B969EADEFFFFD952C8411C49FDE5F88B" box="[286,529,1880,1906]" italics="true" pageId="74" pageNumber="75">Podocerus odontonyx</emphasis>
</taxonomicName>
(
<figureCitation id="13262A49FFFFD952CB7F1C49FD85F88B" box="[544,625,1880,1906]" captionStart="FIGURE 38" captionStartId="76.[151,250,1828,1852]" captionTargetBox="[228,1348,182,1794]" captionTargetId="figure-22@76.[219,1367,181,1804]" captionTargetPageId="76" captionText="FIGURE 38. Jassa pusilla (Sars, 1894). Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus odontonyx Sars, 1894, submerged by Conlan (1990)). Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus pusillus Sars, 1894, transferred to Jassa pusilla by Stebbing, 1906). Medial views. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580732" httpUri="https://zenodo.org/record/4580732/files/figure.png" pageId="74" pageNumber="75">Fig. 38</figureCitation>
) (see
<bibRefCitation id="EF8C4B3DFFFFD952CBEB1C49FCBCF88B" author="Conlan, K. E." box="[692,840,1880,1906]" pageId="74" pageNumber="75" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan 1990</bibRefCitation>
).
<figureCitation id="13262A49FFFFD952CA041C49FC49F88B" box="[859,957,1880,1906]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Fig. 39B</figureCitation>
is interpreted as being juvenile because no thumbed cuticle was visible internally, its pre-thumb is short relative to body length (
<figureCitation id="13262A49FFFFD952CD031C6DFB5AF86F" box="[1116,1198,1916,1942]" captionStart="FIGURE 40" captionStartId="78.[151,250,1123,1147]" captionTargetBox="[159,1427,209,1092]" captionTargetId="figure-22@78.[151,1436,201,1100]" captionTargetPageId="78" captionText="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="74" pageNumber="75">Fig. 40</figureCitation>
) and its dactyl is not centrally toothed as in the minor forms (
<figureCitation id="13262A49FFFFD952CB021CB1FD47F843" box="[605,691,1952,1978]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="74" pageNumber="75">Figs 39</figureCitation>
CE). None of the specimens termed juvenile and subadult in
<figureCitation id="13262A49FFFFD952CC2F1CB1FF47F827" captionStart="FIGURE 40" captionStartId="78.[151,250,1123,1147]" captionTargetBox="[159,1427,209,1092]" captionTargetId="figure-22@78.[151,1436,201,1100]" captionTargetPageId="78" captionText="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="74" pageNumber="75">Fig. 40</figureCitation>
had centrally toothed dactyls.
</paragraph>
<caption id="DF626644FFFED953C9C81C0BFBBFF81F" ID-DOI="http://doi.org/10.5281/zenodo.4580730" ID-Zenodo-Dep="4580730" httpUri="https://zenodo.org/record/4580730/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,250,1818,1842]" targetBox="[173,1416,181,1786]" targetPageId="75">
<paragraph id="8BA236CCFFFED953C9C81C0BFBBFF81F" blockId="75.[151,1437,1818,2022]" pageId="75" pageNumber="76">
<emphasis id="B969EADEFFFED953C9C81C0BFED4F8CB" bold="true" box="[151,288,1818,1842]" pageId="75" pageNumber="76">FIGURE 37.</emphasis>
<taxonomicName id="4C1D4D4FFFFED953C8781C0AFDC5F8CB" authority="(Sars, 1894)" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[295,561,1818,1842]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFED953C8781C0AFE5EF8CB" box="[295,426,1818,1842]" italics="true" pageId="75" pageNumber="76">Jassa pusilla</emphasis>
(
<bibRefCitation id="EF8C4B3DFFFED953C8E71C0BFDDDF8CB" author="Sars, G. O." box="[440,553,1818,1842]" pageId="75" pageNumber="76" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
)
</taxonomicName>
. Paralectotype, adult female 1, about 4.5 mm, UiO F.10562 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in
<bibRefCitation id="EF8C4B3DFFFED953CC7E1C2FFA6DF8AF" author="Sars, G. O." box="[1313,1433,1854,1878]" pageId="75" pageNumber="76" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars (1894)</bibRefCitation>
, plate 213 as
<taxonomicName id="4C1D4D4FFFFED953C8491C73FDB8F883" authority="Sars, 1894" authorityName="Sars" authorityYear="1894" box="[278,588,1890,1914]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B969EADEFFFED953C8491C73FE2CF883" box="[278,472,1890,1914]" italics="true" pageId="75" pageNumber="76">Podocerus pusillus</emphasis>
<bibRefCitation id="EF8C4B3DFFFED953C8811C73FDB8F883" author="Sars, G. O." box="[478,588,1890,1914]" pageId="75" pageNumber="76" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
</taxonomicName>
. Adult female 2, body length unknown, UiO F.10564 Mp 41. Locality, date and collector unknown, illustrated in
<bibRefCitation id="EF8C4B3DFFFED953C8951C97FDB7F867" author="Sars, G. O." box="[458,579,1926,1950]" pageId="75" pageNumber="76" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars (1894)</bibRefCitation>
, plate 213 as
<taxonomicName id="4C1D4D4FFFFED953CB901C97FBF3F867" authority="Sars, 1894" authorityName="Sars" authorityYear="1894" box="[719,1031,1926,1950]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B969EADEFFFED953CB901C97FC65F867" box="[719,913,1926,1950]" italics="true" pageId="75" pageNumber="76">Podocerus pusillus</emphasis>
<bibRefCitation id="EF8C4B3DFFFED953CAC71C97FBF3F867" author="Sars, G. O." box="[920,1031,1926,1950]" pageId="75" pageNumber="76" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
</taxonomicName>
. Adult female 3, 4.7 mm, UiO F.13359. Riŝr, Norway (approx. 59°N, 9°E), date unknown, G. O. Sars, coll. Lateral view: whole body, gnathopods 1 and 2 and pereopod 3; dorsal view: telson; medial view: uropod 3. Scale 0.1 mm. Illustration after
<bibRefCitation id="EF8C4B3DFFFED953CAEE1CDFFBB3F81F" author="Conlan, K. E." box="[945,1095,1998,2022]" pageId="75" pageNumber="76" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990)</bibRefCitation>
.
</paragraph>
</caption>
<caption id="DF626644FFF9D954C9C81C35FD79F809" ID-DOI="http://doi.org/10.5281/zenodo.4580732" ID-Zenodo-Dep="4580732" httpUri="https://zenodo.org/record/4580732/files/figure.png" pageId="76" pageNumber="77" startId="76.[151,250,1828,1852]" targetBox="[228,1348,182,1794]" targetPageId="76">
<paragraph id="8BA236CCFFF9D954C9C81C35FD79F809" blockId="76.[151,1437,1828,2032]" pageId="76" pageNumber="77">
<emphasis id="B969EADEFFF9D954C9C81C35FED6F8C5" bold="true" box="[151,290,1828,1852]" pageId="76" pageNumber="77">FIGURE 38.</emphasis>
<taxonomicName id="4C1D4D4FFFF9D954C8761C34FDCCF8C5" authority="(Sars, 1894)" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[297,568,1828,1852]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFF9D954C8761C34FE5BF8C5" box="[297,431,1828,1852]" italics="true" pageId="76" pageNumber="77">Jassa pusilla</emphasis>
(
<bibRefCitation id="EF8C4B3DFFF9D954C8E11C35FDC4F8C5" author="Sars, G. O." box="[446,560,1828,1852]" pageId="76" pageNumber="77" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
)
</taxonomicName>
. Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in
<bibRefCitation id="EF8C4B3DFFF9D954CD681C59FB5BF899" author="Sars, G. O." box="[1079,1199,1864,1888]" pageId="76" pageNumber="77" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars (1894)</bibRefCitation>
, plate 213 (lectotype of
<taxonomicName id="4C1D4D4FFFF9D954C9C81C7DFE1DF87D" authority="Sars, 1894" authorityName="Sars" authorityYear="1894" box="[151,489,1900,1924]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="odontonyx">
<emphasis id="B969EADEFFF9D954C9C81C7DFE81F87D" box="[151,373,1900,1924]" italics="true" pageId="76" pageNumber="77">Podocerus odontonyx</emphasis>
<bibRefCitation id="EF8C4B3DFFF9D954C8241C7DFE1DF87D" author="Sars, G. O." box="[379,489,1900,1924]" pageId="76" pageNumber="77" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
</taxonomicName>
, submerged by
<bibRefCitation id="EF8C4B3DFFF9D954CBD51C7DFCDDF87D" author="Conlan, K. E." box="[650,809,1900,1924]" pageId="76" pageNumber="77" pagination="2031 - 2075" refId="ref90850" refString="Conlan, K. E. (1990) Revision of the crustacean amphipod genus Jassa Leach (Corophioidea: Ischyroceridae). Canadian Journal of Zoology, 68, 2031 - 2075. https: // doi. org / 10.1139 / z 90 - 288" type="journal article" year="1990">Conlan (1990))</bibRefCitation>
. Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in
<bibRefCitation id="EF8C4B3DFFF9D954C8821CA5FDA1F835" author="Sars, G. O." box="[477,597,1972,1996]" pageId="76" pageNumber="77" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars (1894)</bibRefCitation>
, plate 213 (lectotype of
<taxonomicName id="4C1D4D4FFFF9D954CA141CA5FB74F835" authority="Sars, 1894" authorityName="Sars" authorityYear="1894" box="[843,1152,1972,1996]" class="Malacostraca" family="Podoceridae" genus="Podocerus" kingdom="Animalia" order="Amphipoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B969EADEFFF9D954CA141CA5FBF8F835" box="[843,1036,1972,1996]" italics="true" pageId="76" pageNumber="77">Podocerus pusillus</emphasis>
<bibRefCitation id="EF8C4B3DFFF9D954CD4C1CA5FB74F835" author="Sars, G. O." box="[1043,1152,1972,1996]" pageId="76" pageNumber="77" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
</taxonomicName>
, transferred to
<taxonomicName id="4C1D4D4FFFF9D954CC461CA4FA68F835" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[1305,1436,1972,1996]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFF9D954CC461CA4FA68F835" box="[1305,1436,1972,1996]" italics="true" pageId="76" pageNumber="77">Jassa pusilla</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8C4B3DFFF9D954C9E81CC9FEA1F809" author="Stebbing, T. R. R." box="[183,341,2008,2032]" pageId="76" pageNumber="77" pagination="1 - 806" refId="ref105591" refString="Stebbing, T. R. R. (1906) Amphipoda. I. Gammaridea. Das Tierreich, 21, 1 - 806." type="journal article" year="1906">Stebbing, 1906</bibRefCitation>
). Medial views. Scale 0.1 mm.
</paragraph>
</caption>
<caption id="DF626644FFF8D955C9C81C7BFC0AF833" ID-DOI="http://doi.org/10.5281/zenodo.4580734" ID-Zenodo-Dep="4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="77" pageNumber="78" startId="77.[151,250,1898,1922]" targetBox="[163,1430,187,1869]" targetPageId="77">
<paragraph id="8BA236CCFFF8D955C9C81C7BFC0AF833" blockId="77.[151,1437,1898,1995]" pageId="77" pageNumber="78">
<emphasis id="B969EADEFFF8D955C9C81C7BFED5F87B" bold="true" box="[151,289,1898,1922]" pageId="77" pageNumber="78">FIGURE 39.</emphasis>
<taxonomicName id="4C1D4D4FFFF8D955C8771C7AFDC0F87A" authority="(Sars, 1894)" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[296,564,1898,1923]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFF8D955C8771C7AFE59F87B" box="[296,429,1898,1922]" italics="true" pageId="77" pageNumber="78">Jassa pusilla</emphasis>
(
<bibRefCitation id="EF8C4B3DFFF8D955C8E41C7AFDD9F87A" author="Sars, G. O." box="[443,557,1899,1923]" pageId="77" pageNumber="78" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
)
</taxonomicName>
. Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in
<emphasis id="B969EADEFFF8D955CB7A1CA3FD59F833" bold="true" box="[549,685,1970,1994]" pageId="77" pageNumber="78">FIGURE 40.</emphasis>
All views medial. Scale 0.1 mm.
</paragraph>
</caption>
<caption id="DF626644FFFBD956C9C81F72FC9AFAF2" ID-DOI="http://doi.org/10.5281/zenodo.4580736" ID-Zenodo-Dep="4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,250,1123,1147]" targetBox="[159,1427,209,1092]" targetPageId="78">
<paragraph id="8BA236CCFFFBD956C9C81F72FC9AFAF2" blockId="78.[151,1437,1123,1291]" pageId="78" pageNumber="79">
<emphasis id="B969EADEFFFBD956C9C81F72FED4FB82" bold="true" box="[151,288,1123,1147]" pageId="78" pageNumber="79">FIGURE 40.</emphasis>
<taxonomicName id="4C1D4D4FFFFBD956C8791F75FDDBFB82" authority="(Sars, 1894)" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[294,559,1123,1147]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956C8791F75FE5DFB82" box="[294,425,1123,1147]" italics="true" pageId="78" pageNumber="79">Jassa pusilla</emphasis>
(
<bibRefCitation id="EF8C4B3DFFFBD956C8E81F72FDD3FB82" author="Sars, G. O." box="[439,551,1123,1147]" pageId="78" pageNumber="79" pagination="589 - 628" refId="ref104187" refString="Sars, G. O. (1894) Amphipoda. Part XXVII &amp; XXVIII. Podoceridae (concluded), Corophiidae, Cheluridae. An account of the Crustacea of Norway, with short descriptions and figures of all the species, 1, 589 - 628. https: // doi. org / 10.5962 / bhl. title. 1164" type="journal article" year="1894">Sars, 1894</bibRefCitation>
)
</taxonomicName>
. Variation in male gnathopod 2 propodus length relative to body length in four populations. Väderârna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiOF. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiOF. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael
<superScript id="7C689B84FFFBD956CBE61FDEFD18FB25" attach="left" box="[697,748,1230,1254]" fontSize="6" pageId="78" pageNumber="79">Parsn</superScript>
Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in
<emphasis id="B969EADEFFFBD956C9B61FE2FEB8FAF2" bold="true" box="[233,332,1267,1291]" pageId="78" pageNumber="79">FIGURE</emphasis>
39. Linear regression assumptions failed for all plots.
</paragraph>
</caption>
<paragraph id="8BA236CCFFFBD956C9981E2BFC20FA46" blockId="78.[151,1437,1337,1868]" pageId="78" pageNumber="79">
Four populations sampled in Väderârna,
<collectingCountry id="F30A765CFFFBD956CBF41E28FCF2FAAA" box="[683,774,1337,1363]" name="Sweden" pageId="78" pageNumber="79">Sweden</collectingCountry>
, Trondheimsfjord,
<collectingCountry id="F30A765CFFFBD956CABE1E2BFBC8FAAD" box="[993,1084,1338,1364]" name="Norway" pageId="78" pageNumber="79">Norway</collectingCountry>
, Riŝr,
<collectingCountry id="F30A765CFFFBD956CDC91E2BFB04FAAD" box="[1174,1264,1338,1364]" name="Norway" pageId="78" pageNumber="79">Norway</collectingCountry>
and Tórshavn,
<collectingCountry id="F30A765CFFFBD956C9C81E4FFED9FA8E" box="[151,301,1373,1400]" name="Faroe Islands" pageId="78" pageNumber="79">Farøe Islands</collectingCountry>
showed a similar relationship between thumb length and body length, with the juveniles having prethumbs and the minor forms being rare (
<figureCitation id="13262A49FFFBD956CB071E90FD58FA62" box="[600,684,1409,1435]" captionStart="FIGURE 40" captionStartId="78.[151,250,1123,1147]" captionTargetBox="[159,1427,209,1092]" captionTargetId="figure-22@78.[151,1436,201,1100]" captionTargetPageId="78" captionText="FIGURE 40. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus length relative to body length in four populations. Väder̂arna, Sweden (~ 58.57°N, 11.06°E), NRM 5427. Trondheimsfjord, Norway (~ 63.43°N, 10.39°E), date unknown, G. O. Sars, coll, UiO F. 13352. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Tórshavn, Farøe Islands (60°55ʹN, 8°56ʹW) station 69L.9,33, Michael Parsn Sd. Jenson (UiO). Letters on the Riŝr graph pertain to the illustrations in FIGURE 39. Linear regression assumptions failed for all plots." figureDoi="http://doi.org/10.5281/zenodo.4580736" httpUri="https://zenodo.org/record/4580736/files/figure.png" pageId="78" pageNumber="79">Fig. 40</figureCitation>
).
<taxonomicName id="4C1D4D4FFFFBD956CBE01E93FCA4FA62" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[703,848,1409,1435]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956CBE01E93FCA4FA62" box="[703,848,1409,1435]" italics="true" pageId="78" pageNumber="79">Jassa pusilla</emphasis>
</taxonomicName>
is a small species, with the minor forms
<quantity id="4CE59B29FFFBD956CC491E90FA69FA65" box="[1302,1437,1409,1436]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.8" metricValueMin="2.8" pageId="78" pageNumber="79" unit="mm" value="3.3" valueMax="3.8" valueMin="2.8">2.83.8 mm</quantity>
in body length and the major forms 3.0
<quantity id="4CE59B29FFFBD956CB041EB4FD44FA39" box="[603,688,1445,1472]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="78" pageNumber="79" unit="mm" value="4.5">4.5 mm</quantity>
in these four populations.
</paragraph>
<paragraph id="8BA236CCFFFBD956C9981ED8FD3FF8B2" blockId="78.[151,1437,1337,1868]" pageId="78" pageNumber="79">
The record of
<taxonomicName id="4C1D4D4FFFFBD956C83C1EDBFE3CFA1A" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[355,456,1481,1507]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956C83C1EDBFE3CFA1A" box="[355,456,1481,1507]" italics="true" pageId="78" pageNumber="79">J. pusilla</emphasis>
</taxonomicName>
from samples along the coast of
<collectingCountry id="F30A765CFFFBD956CA6B1ED8FC67FA1A" box="[820,915,1481,1507]" name="Portugal" pageId="78" pageNumber="79">Portugal</collectingCountry>
by
<bibRefCitation id="EF8C4B3DFFFBD956CAE51ED8FB77FA1A" author="Lobo, J. &amp; Ferreira, M. S. &amp; Antunes, I. C. &amp; Teixeira, M. A. L. &amp; Borges, L. M. S. &amp; Sousa, R. &amp; Gomes, P. A. &amp; Costa, M. H. &amp; Cunha, M. R. &amp; Costa, F. O." box="[954,1155,1481,1507]" pageId="78" pageNumber="79" pagination="147 - 157" refId="ref99132" refString="Lobo, J., Ferreira, M. S., Antunes, I. C., Teixeira, M. A. L., Borges, L. M. S., Sousa, R., Gomes, P. A., Costa, M. H., Cunha, M. R. &amp; Costa, F. O. (2017) Contrasting morphological and DNA barcode-suggested species boundaries among shallow-water amphipod fauna from the southern European Atlantic coast. Genome, 60, 147 - 157. https: // doi. org / 10.1139 / gen- 2016 - 0009" type="journal article" year="2017">
Lobo
<emphasis id="B969EADEFFFBD956CAA41EDBFBC6FA1A" box="[1019,1074,1481,1507]" italics="true" pageId="78" pageNumber="79">et al.</emphasis>
(2017)
</bibRefCitation>
is in error, and is the new species
<taxonomicName id="4C1D4D4FFFFBD956C9AF1EFFFE93F9FE" authorityName="Conlan &amp; Desiderato &amp; Beermann" authorityYear="2021" box="[240,359,1517,1543]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="laurieae">
<emphasis id="B969EADEFFFBD956C9AF1EFFFE93F9FE" box="[240,359,1517,1543]" italics="true" pageId="78" pageNumber="79">J. laurieae</emphasis>
</taxonomicName>
. The three collections of
<taxonomicName id="4C1D4D4FFFFBD956CBD71EFFFD1BF9FE" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[648,751,1517,1543]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956CBD71EFFFD1BF9FE" box="[648,751,1517,1543]" italics="true" pageId="78" pageNumber="79">J. pusilla</emphasis>
</taxonomicName>
from deep water offshore of the Atlantic
<collectingCountry id="F30A765CFFFBD956CD9C1EFCFB01F9FE" box="[1219,1269,1517,1543]" name="United States of America" pageId="78" pageNumber="79">U.S.</collectingCountry>
coast (NMNH 6335, 33530, and 106781) (
<figureCitation id="13262A49FFFBD956C8931D00FDFBF9D2" box="[460,527,1553,1579]" captionStart="FIGURE 9" captionStartId="20.[151,250,1907,1931]" captionTargetBox="[224,1363,181,1883]" captionTargetId="figure-22@20.[224,1363,181,1883]" captionTargetPageId="20" captionText="FIGURE 9. Records of Northern Hemisphere species of Jassa that were found in the eastern and/or western North Atlantic and seas, along with J. kimi in the western Pacific. Red dot: identification confirmed by specimen examination; green triangle: literature record (Supplementary Table S1) with identification not confirmed but judged likely to be this species; blue x: suspected or known to be on the hull or interior of a boat or ship." figureDoi="http://doi.org/10.5281/zenodo.4580652" httpUri="https://zenodo.org/record/4580652/files/figure.png" pageId="78" pageNumber="79">Fig. 9</figureCitation>
) morphologically resembled this species, not either of the other two species from this coast (
<taxonomicName id="4C1D4D4FFFFBD956C8511D27FE99F9B6" baseAuthorityName="Dana" baseAuthorityYear="1853" box="[270,365,1589,1615]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="valida">
<emphasis id="B969EADEFFFBD956C8511D27FE99F9B6" box="[270,365,1589,1615]" italics="true" pageId="78" pageNumber="79">J. valida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D4FFFFBD956C8FE1D27FDCCF9B6" authorityName="Holmes" authorityYear="1905" box="[417,568,1590,1615]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis id="B969EADEFFFBD956C8FE1D27FDCCF9B6" box="[417,568,1590,1615]" italics="true" pageId="78" pageNumber="79">J. marmorata</emphasis>
</taxonomicName>
), which occur shallower.
<taxonomicName id="4C1D4D4FFFFBD956CA0B1D27FC10F9B6" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[852,996,1589,1615]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956CA0B1D27FC10F9B6" box="[852,996,1589,1615]" italics="true" pageId="78" pageNumber="79">Jassa pusilla</emphasis>
</taxonomicName>
is a smaller species with delicate, never plumose antennae, lacking the typical long setae along the basis of gnathopod 2 that the other two possess, and also lacking the diagnostic antero-medial seta on the gnathopod 1 carpus at the junction of the propodus. The minor form thumbed male of
<taxonomicName id="4C1D4D4FFFFBD956C8001DB3FE05F942" baseAuthorityName="Sars" baseAuthorityYear="1894" box="[351,497,1697,1723]" class="Malacostraca" family="Ischyroceridae" genus="Jassa" kingdom="Animalia" order="Amphipoda" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="pusilla">
<emphasis id="B969EADEFFFBD956C8001DB3FE05F942" box="[351,497,1697,1723]" italics="true" pageId="78" pageNumber="79">Jassa pusilla</emphasis>
</taxonomicName>
has a strong tooth on the inner margin of the dactyl, which the minor forms of the other two species lack. One of the specimens from NMNH 6335 was a
<quantity id="4CE59B29FFFBD956CAF21DD4FC1AF919" box="[941,1006,1733,1760]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="78" pageNumber="79" unit="mm" value="3.0">3 mm</quantity>
long minor form thumbed male with a strong tooth on the dactyl, its propodus closely resembling that in
<figureCitation id="13262A49FFFBD956CA2A1DF8FC3CF8FA" box="[885,968,1769,1795]" captionStart="FIGURE 38" captionStartId="76.[151,250,1828,1852]" captionTargetBox="[228,1348,182,1794]" captionTargetId="figure-22@76.[219,1367,181,1804]" captionTargetPageId="76" captionText="FIGURE 38. Jassa pusilla (Sars, 1894). Adult male gnathopod 2, minor form, body length unknown, UiO F. 10565 Mp 41. Same location and collector, drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus odontonyx Sars, 1894, submerged by Conlan (1990)). Adult male gnathopod 2, major form, body length unknown, UiO F. 10563 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 (lectotype of Podocerus pusillus Sars, 1894, transferred to Jassa pusilla by Stebbing, 1906). Medial views. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580732" httpUri="https://zenodo.org/record/4580732/files/figure.png" pageId="78" pageNumber="79">Figs 38</figureCitation>
and
<figureCitation id="13262A49FFFBD956CAA21DF8FBDAF8FA" box="[1021,1070,1769,1795]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="78" pageNumber="79">39E</figureCitation>
. Another thumbed male (NMNH 33530) closely resembled
<figureCitation id="13262A49FFFBD956C89D1C1CFDDCF8DE" box="[450,552,1805,1831]" captionStart="FIGURE 39" captionStartId="77.[151,250,1898,1922]" captionTargetBox="[163,1430,187,1869]" captionTargetId="figure-18@77.[151,1436,181,1874]" captionTargetPageId="77" captionText="FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4580734" httpUri="https://zenodo.org/record/4580734/files/figure.png" pageId="78" pageNumber="79">Fig. 39B</figureCitation>
. The other specimens were female or juvenile. These three collections are the only ones known from the western North Atlantic.
</paragraph>
</subSubSection>
</treatment>
</document>