236 lines
28 KiB
XML
236 lines
28 KiB
XML
<document id="DF56F1069BF1AD068F77228E7F94F519" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. & Mathewes, Rolf W." docDate="2021" docId="03B487C2007FFFFAFF5BFE53FE781460" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Whetwhetaksidae Archibald & Cannings 2021, new family" docType="treatment" docVersion="8" lastPageNumber="110" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="110" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:mods id="E4234488FF7AD74D0B0A9D50FD7B06B2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="23A79FB7B28551E7A34CC31BD811F79F">
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<mods:title id="50772220D5635F45D8B5A0D8C5BE1725">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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</mods:titleInfo>
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<mods:name id="573E38938355BA9256D3807D9F712D6D" type="personal">
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</mods:role>
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<mods:namePart id="EFD43D06182003CD4B7A5CCC0CAB79BF">Archibald, Bruce</mods:namePart>
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</mods:name>
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<mods:role id="E0BE1A046CCB9187587FCE6D5000EC45">
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<mods:roleTerm id="DDD6B46079226E0AB9C49955920AEAAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="68CFD72388CD067DCB658B4EDAC54626">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="855548B24CDAA5672A150D672F061C56" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:nameIdentifier id="A99A510C60A36A236F19A9A2B39E4DB7" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name id="0FA6A519BF1B687EBD7805A7BC04824C" type="personal">
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<mods:roleTerm id="93401038F6293D496B70C33B7A1D9745">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E23F13440DEEE79999681D8796AB025D">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="CE13A38C3AE2EE3CD04073D206DDE0A9" type="ORCID">0000-0003-1162-0355</mods:nameIdentifier>
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<mods:nameIdentifier id="CB760B22F8980C0271AC6FE5F2B4E8C6" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="9EC7A6D1226D189519C2A176C00D4935" type="personal">
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<mods:roleTerm id="8A2ED58FC13A2B6B0FAC8ACEF215B4AF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2873E5D52B02B6CD90BDFE0EE64068F9">Bybee, Seth M.</mods:namePart>
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<mods:nameIdentifier id="C1EA5FCCE45B9085FC4B44BB9DF1158C" type="ORCID">0000-0001-6224-6303</mods:nameIdentifier>
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<mods:nameIdentifier id="4EE0F6A5F61E72C2B69C311A392694B8" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="6144B6DD8E2FF7F97AB3C662E3B68DA9" type="personal">
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<mods:roleTerm id="D543775F3360FF846F63408F5EAF6304">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4BC23D16D95EFFD1BDCC6E905F92CA08">Mathewes, Rolf W.</mods:namePart>
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<mods:nameIdentifier id="A4E501557537AEB3B05C6187000DD172" type="ORCID">0000-0001-7637-199X</mods:nameIdentifier>
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<mods:nameIdentifier id="80BF88471C48EE96222E294D5ED6FB11" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
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<mods:typeOfResource id="0E9A8F67563F80C321A96636352C0263">text</mods:typeOfResource>
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<mods:title id="4B57CF0D7EA4C745950F0ABABA8AC535">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AC68B52BA2A6E0C6AE27249A1B895627">
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<mods:date id="90EE9C8B0941856953A5E67F5D192703">2021</mods:date>
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<mods:number id="705F68F1FBEF46ED453DDCC35C31EBDF">2021-02-24</mods:number>
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<mods:detail id="2E60701E11CD3EFF9382812CD75E3B3D" type="volume">
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<mods:number id="37716060785D15A9335E557646AF914F">4934</mods:number>
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</mods:detail>
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<mods:detail id="CF75483E2671872E782FE7F5A4A9F8A8" type="issue">
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<mods:number id="8457E74F9B20E45D57245D4A75729049">1</mods:number>
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<mods:classification id="9035CE2C4847A6F5901AB56DD8D68EBF">journal article</mods:classification>
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<mods:identifier id="5219BDC5BD84B2C02B511F81FB6D4C5B" type="CLB-Dataset">7439</mods:identifier>
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<mods:identifier id="B0A6EF538A71E6F56675C5CFB545F1E9" type="DOI">10.11646/zootaxa.4934.1.1</mods:identifier>
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<mods:identifier id="E35C3A1CB551B792BFC0B8521358FD23" type="GBIF-Dataset">300bac18-55f5-4e34-90e1-7fccd02d68c1</mods:identifier>
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<mods:identifier id="F1B18FB5B6F062F7B1486B2367C4EE05" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EC4EC50CFD9501D413807AB5142A529D" type="Zenodo-Dep">4558796</mods:identifier>
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<mods:identifier id="8818F5EDE054D1D7B81EABC99FD067E3" type="ZooBank">79895443-4597-42A5-AF8A-023EACB20E10</mods:identifier>
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<treatment id="03B487C2007FFFFAFF5BFE53FE781460" ID-DOI="http://doi.org/10.5281/zenodo.4672726" ID-GBIF-Taxon="180881783" ID-Zenodo-Dep="4672726" LSID="urn:lsid:plazi:treatment:03B487C2007FFFFAFF5BFE53FE781460" httpUri="http://treatment.plazi.org/id/03B487C2007FFFFAFF5BFE53FE781460" lastPageNumber="110" pageId="109" pageNumber="110">
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<subSubSection id="C307655F007FFFFAFF5BFE53FC9F13C5" box="[151,817,438,465]" pageId="109" pageNumber="110" type="nomenclature">
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<paragraph id="8BA236D4007FFFFAFF5BFE53FC9F13C5" blockId="109.[151,817,438,501]" box="[151,817,438,465]" pageId="109" pageNumber="110">
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<heading id="D0EA81B8007FFFFAFF5BFE53FC9F13C5" bold="true" box="[151,817,438,465]" fontSize="11" level="2" pageId="109" pageNumber="110" reason="3">
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<emphasis id="B969EAC6007FFFFAFF5BFE53FC9F13C5" bold="true" box="[151,817,438,465]" pageId="109" pageNumber="110">
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<taxonomicName id="4C1D4D57007FFFFAFF5BFE53FD3513C5" authority="Archibald & Cannings" authorityName="Archibald & Cannings" authorityYear="2021" box="[151,667,438,464]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae Archibald & Cannings</taxonomicName>
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,
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<taxonomicNameLabel id="A25A57BD007FFFFAFD64FE52FC9F13C5" box="[680,817,438,465]" pageId="109" pageNumber="110" rank="family">new family</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F007FFFFAFF5BFE3EFE8D13E0" box="[151,291,475,501]" pageId="109" pageNumber="110" type="description">
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<paragraph id="8BA236D4007FFFFAFF5BFE3EFE8D13E0" blockId="109.[151,817,438,501]" box="[151,291,475,501]" pageId="109" pageNumber="110">
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<figureCitation id="13262A51007FFFFAFF5BFE3EFEB113E0" box="[151,287,475,501]" captionStart-0="FIGURE 66" captionStart-1="FIGURE 67" captionStart-2="FIGURE 68" captionStart-3="FIGURE 69" captionStart-4="FIGURE 70" captionStartId-0="112.[151,250,1985,2009]" captionStartId-1="113.[151,250,1984,2008]" captionStartId-2="114.[151,250,1961,1985]" captionStartId-3="115.[151,250,1805,1829]" captionStartId-4="116.[151,250,1955,1979]" captionTargetBox-0="[343,1248,184,1961]" captionTargetBox-1="[269,1319,187,1957]" captionTargetBox-2="[399,1189,182,1937]" captionTargetBox-3="[151,1423,181,1769]" captionTargetBox-4="[272,1316,181,1931]" captionTargetId-0="figure-19@112.[326,1261,181,1961]" captionTargetId-1="figure-17@113.[257,1330,181,1960]" captionTargetId-2="figure-19@114.[386,1202,181,1937]" captionTargetId-3="figure-17@115.[151,1436,181,1781]" captionTargetId-4="figure-19@116.[257,1331,181,1931]" captionTargetPageId-0="112" captionTargetPageId-1="113" captionTargetPageId-2="114" captionTargetPageId-3="115" captionTargetPageId-4="116" captionText-0="FIGURE 66. Whetwhetaksa millerae holotype, SR 06-01-38. A, photograph; B, drawing. Both to scale, 5 mm." captionText-1="FIGURE 67. Whetwhetaksa millerae paratype 1, SR 10-36-10. A, photograph; B, drawing. Both to scale." captionText-2="FIGURE 68. Whetwhetaksa millerae paratype 2, SRUI 99-98-34. A, photograph; B, drawing. Both to scale; aa, accessory antenodal crossvein." captionText-3="FIGURE 69. Whetwhetaksa millerae additional material. SR 93-11-01: A, photograph; B, drawing; SR 06-01-36: C, photograph; D, drawing.All to scale, 5 mm." captionText-4="FIGURE 70. Whetwhetaksa millerae, additional material. SR 10-14-19: A, photograph; B, drawing; SR 05-11-01: C, photograph; D, drawing.All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558960" figureDoi-1="http://doi.org/10.5281/zenodo.4558964" figureDoi-2="http://doi.org/10.5281/zenodo.4558968" figureDoi-3="http://doi.org/10.5281/zenodo.4558970" figureDoi-4="http://doi.org/10.5281/zenodo.4558972" httpUri-0="https://zenodo.org/record/4558960/files/figure.png" httpUri-1="https://zenodo.org/record/4558964/files/figure.png" httpUri-2="https://zenodo.org/record/4558968/files/figure.png" httpUri-3="https://zenodo.org/record/4558970/files/figure.png" httpUri-4="https://zenodo.org/record/4558972/files/figure.png" pageId="109" pageNumber="110">Figs. 66–70</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F007FFFFAFF5BFDC7FAAE11F8" pageId="109" pageNumber="110" type="diagnosis">
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<paragraph id="8BA236D4007FFFFAFF5BFDC7FAAE11F8" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
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<emphasis id="B969EAC6007FFFFAFF5BFDC7FEBC1029" bold="true" box="[151,274,546,572]" pageId="109" pageNumber="110">Diagnosis.</emphasis>
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Most easily separated from all other
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<taxonomicName id="4C1D4D57007FFFFAFD74FDC6FC2A1028" box="[696,900,547,573]" kingdom="Animalia" pageId="109" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
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and
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<taxonomicName id="4C1D4D57007FFFFAFC77FDC6FB801028" box="[955,1070,547,573]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
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families by 1, distinctively long pterostigma: length about ten times width (roughly, where known), nodus to base of pterostigma 55–60% of nodus to apex; and from all
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<taxonomicName id="4C1D4D57007FFFFAFE41FD8EFDF71090" box="[397,601,619,645]" kingdom="Animalia" pageId="109" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
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and all
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<taxonomicName id="4C1D4D57007FFFFAFD7AFD8EFC871090" box="[694,809,619,645]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
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except the extant
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<taxonomicName id="4C1D4D57007FFFFAFC35FD8EFA321093" box="[1017,1436,619,646]" class="Insecta" family="Dicteriadidae" genus="Heliocharis" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="amazona">
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<emphasis id="B969EAC6007FFFFAFC35FD8EFB451091" box="[1017,1259,619,645]" italics="true" pageId="109" pageNumber="110">Heliocharis amazona</emphasis>
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(Dicteriadidae)
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</taxonomicName>
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and some Paleogene
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<taxonomicName id="4C1D4D57007FFFFAFE41FD6AFDBE10BC" baseAuthorityName="Jacobson & Bianchi" baseAuthorityYear="1905" box="[397,528,655,681]" class="Insecta" family="Euphaeidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Euphaeidae</taxonomicName>
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by 2, arculus just distal to Ax1, closer to it than to Ax2. Differs further from
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<taxonomicName id="4C1D4D57007FFFFAFF5BFD56FE8E10D8" box="[151,288,691,717]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
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by: 3, oblique vein O absent [present]; 4, nodal, subnodal crossveins oblique [usually nearly vertical, sometimes slightly oblique or reversed obliquity]; 5, base of RP2 close to nodus [more distal]; 6, CuA terminates at margin over half wing length [notably shorter, longest
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<taxonomicName id="4C1D4D57007FFFFAFCD6FD1EFAB01100" authority="Nel & Paicheler" authorityName="Nel & Paicheler" authorityYear="1994" box="[794,1310,763,789]" class="Insecta" family="Sieblosiidae" genus="Parastenolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="oligocenicus">
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<emphasis id="B969EAC6007FFFFAFCD6FD1EFBCE1100" box="[794,1120,763,789]" italics="true" pageId="109" pageNumber="110">Parastenolestes oligocenicus</emphasis>
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Nel & Paicheler
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</taxonomicName>
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, under half wing length]; 7, MA close to, subparallel with RP3-4 to margin [subparallel with MP]; 8, MA, MP diverge to margin [remain close]; 9, MP, CuA remain subparallel to margin [widely diverge]; 10, base to nodus about 40% wing length, minimum about 38% [maximum about 35% where measurable]. Differs further from
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<taxonomicName id="4C1D4D57007FFFFAFB7BFC82FAF41194" box="[1207,1370,871,897]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
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by: 4 [usually like
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<taxonomicName id="4C1D4D57007FFFFAFEE5FC6EFE5D11B0" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[297,499,907,933]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae</taxonomicName>
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, but
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<taxonomicName id="4C1D4D57007FFFFAFDE4FC6EFD3011B1" box="[552,670,907,932]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAC6007FFFFAFDE4FC6EFD3011B1" box="[552,670,907,932]" italics="true" pageId="109" pageNumber="110">Dysagrion</emphasis>
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</taxonomicName>
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subnodal crossvein reverse obliquity]; 7–9, [varies], 10, [
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<taxonomicName id="4C1D4D57007FFFFAFAEAFC6EFEAF11DC" authorityName="Scudder Lutetian Grube Messel" authorityYear="1885" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="packardii">
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<emphasis id="B969EAC6007FFFFAFAEAFC6EFEAF11DC" italics="true" pageId="109" pageNumber="110">Dysagrion packardii</emphasis>
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</taxonomicName>
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, about 42%,
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<taxonomicName id="4C1D4D57007FFFFAFE58FC4AFD3C11DC" authorityName="Cockerell" authorityYear="1927" box="[404,658,943,969]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="B969EAC6007FFFFAFE58FC4AFD3C11DC" box="[404,658,943,969]" italics="true" pageId="109" pageNumber="110">Petrolestes hendersoni</emphasis>
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</taxonomicName>
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, about 42%, the rest about 36–25%]; 11, origin of IR
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<quantity id="4CE59B31007FFFFAFB2FFC4AFAA311DC" box="[1251,1293,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="109" pageNumber="110" unit="in" value="2.0">2 in</quantity>
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middle third between arculus, nodus [at subnodus in
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<taxonomicName id="4C1D4D57007FFFFAFD9AFC36FD5211F8" box="[598,764,979,1005]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
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, but as in
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<taxonomicName id="4C1D4D57007FFFFAFCA1FC36FB9911F8" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[877,1079,979,1005]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae</taxonomicName>
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, in Petrolestinae].
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F007FFFFAFF0BFC12FD1C1604" box="[199,690,1014,1041]" pageId="109" pageNumber="110" type="materials_examined">
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<paragraph id="8BA236D4007FFFFAFF0BFC12FD1C1604" blockId="109.[151,1437,546,1653]" box="[199,690,1014,1041]" pageId="109" pageNumber="110">
|
||
<emphasis id="B969EAC6007FFFFAFF0BFC12FDAD1604" bold="true" box="[199,515,1014,1041]" pageId="109" pageNumber="110">
|
||
<typeStatus id="54A68876007FFFFAFF0BFC12FEAC1604" box="[199,258,1015,1041]" pageId="109" pageNumber="110">Type</typeStatus>
|
||
and included genera.
|
||
</emphasis>
|
||
<taxonomicName id="4C1D4D57007FFFFAFDC5FC12FD051604" authorityName="Archibald & Cannings" authorityYear="2021" box="[521,683,1015,1041]" class="Insecta" family="Sieblosiidae" genus="Whetwhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6007FFFFAFDC5FC12FD051604" box="[521,683,1015,1041]" italics="true" pageId="109" pageNumber="110">Whetwhetaksa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F007FFFFAFF0BFBFFFD641620" box="[199,714,1050,1077]" pageId="109" pageNumber="110" type="description">
|
||
<paragraph id="8BA236D4007FFFFAFF0BFBFFFD641620" blockId="109.[151,1437,546,1653]" box="[199,714,1050,1077]" pageId="109" pageNumber="110">
|
||
<emphasis id="B969EAC6007FFFFAFF0BFBFFFEF71621" bold="true" box="[199,345,1050,1076]" pageId="109" pageNumber="110">Description.</emphasis>
|
||
As for its only genus, see below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F007FFFFAFF0BFBDAFD9416B4" pageId="109" pageNumber="110" type="etymology">
|
||
<paragraph id="8BA236D4007FFFFAFF0BFBDAFB7C164C" blockId="109.[151,1437,546,1653]" box="[199,1234,1087,1113]" pageId="109" pageNumber="110">
|
||
<emphasis id="B969EAC6007FFFFAFF0BFBDAFEE0164C" bold="true" box="[199,334,1087,1113]" pageId="109" pageNumber="110">Etymology.</emphasis>
|
||
The family name is formed from that of its
|
||
<typeStatus id="54A68876007FFFFAFCF6FBDAFCC4164C" box="[826,874,1087,1113]" pageId="109" pageNumber="110">type</typeStatus>
|
||
and only genus,
|
||
<taxonomicName id="4C1D4D57007FFFFAFBE5FBDAFB65164C" authorityName="Archibald & Cannings" authorityYear="2021" box="[1065,1227,1087,1113]" class="Insecta" family="Sieblosiidae" genus="Whetwhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6007FFFFAFBE5FBDAFB65164C" box="[1065,1227,1087,1113]" italics="true" pageId="109" pageNumber="110">Whetwhetaksa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4007FFFFAFF0BFB86FD9416B4" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
|
||
<emphasis id="B969EAC6007FFFFAFF0BFB86FE2A1668" bold="true" box="[199,388,1123,1149]" pageId="109" pageNumber="110">Range and age.</emphasis>
|
||
Republic localities B4131 and A0307B of the Tom Thumb Tuff Member of the Klondike Mountain Formation; latest Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F007FFFFAFF0BFB4FFE781460" pageId="109" pageNumber="110" type="discussion">
|
||
<paragraph id="8BA236D4007FFFFAFF0BFB4FFDD216FF" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
|
||
<emphasis id="B969EAC6007FFFFAFF0BFB4FFEE216D1" bold="true" box="[199,332,1194,1220]" pageId="109" pageNumber="110">Discussion.</emphasis>
|
||
Such an extremely long pterostigma is never found in the
|
||
<taxonomicName id="4C1D4D57007FFFFAFBCEFB4EFBDB16D0" box="[1026,1141,1195,1221]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
to our knowledge, but is matched by some
|
||
<taxonomicName id="4C1D4D57007FFFFAFEAEFB2AFDD716FF" authority="(Anisoptera)" baseAuthorityName="Anisoptera" box="[354,633,1231,1258]" class="Insecta" family="Petaluridae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Petaluridae (Anisoptera)</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4007FFFFAFF0BFB16FAC1176C" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
|
||
The position of the arculus closer to, immediately distal to Ax1 is remarkable. This condition is found in the
|
||
<taxonomicName id="4C1D4D57007FFFFAFABDFB16FE881724" authorityName="Handlirsch" authorityYear="1906" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Anisozygoptera">Anisozygoptera</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D57007FFFFAFE96FAF2FE761724" box="[346,472,1303,1329]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="4" phylum="Arthropoda" rank="subOrder" subOrder="Anisoptera">Anisoptera</taxonomicName>
|
||
, but in the
|
||
<taxonomicName id="4C1D4D57007FFFFAFD9DFAF2FD6A1724" box="[593,708,1303,1329]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
only in the South American
|
||
<taxonomicName id="4C1D4D57007FFFFAFBCEFAF2FB261725" box="[1026,1160,1303,1328]" class="Insecta" family="Dicteriadidae" genus="Heliocharis" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="amazona">
|
||
<emphasis id="B969EAC6007FFFFAFBCEFAF2FB261725" box="[1026,1160,1303,1328]" italics="true" pageId="109" pageNumber="110">H. amazona</emphasis>
|
||
</taxonomicName>
|
||
, which otherwise differs strongly in many ways and a few Paleogene
|
||
<taxonomicName id="4C1D4D57007FFFFAFD46FADEFCA31740" baseAuthorityName="Jacobson & Bianchi" baseAuthorityYear="1905" box="[650,781,1339,1365]" class="Insecta" family="Euphaeidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Euphaeidae</taxonomicName>
|
||
(see above) which also strongly differ. In all other
|
||
<taxonomicName id="4C1D4D57007FFFFAFA86FADEFF6F176C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
it is aligned with, slightly proximal to, or slightly to clearly distal to Ax2 (
|
||
<bibRefCitation id="EF8C4B25007FFFFAFBCFFABAFB39176C" author="Bechly, G." box="[1027,1175,1375,1401]" pageId="109" pageNumber="110" pagination="1 - 402" refId="ref56079" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata) unter besonderer Beruchsichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Boblingen, Special Volume, 2, 1 - 402." type="journal article" year="1996">Bechly 1996</bibRefCitation>
|
||
; Fleck
|
||
<emphasis id="B969EAC6007FFFFAFB2BFABAFAB7176C" box="[1255,1305,1375,1401]" italics="true" pageId="109" pageNumber="110">et al</emphasis>
|
||
. 2004).
|
||
</paragraph>
|
||
<paragraph id="8BA236D4007FFFFAFF0BFA66FAB717F0" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
|
||
In SR 06-01-36, the subcosta appears to cross through the nodus, which is characteristic of the
|
||
<taxonomicName id="4C1D4D57007FFFFAFB16FA66FACD1788" box="[1242,1379,1411,1437]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
(see,
|
||
<emphasis id="B969EAC6007FFFFAFF5BFA42FF1617D5" box="[151,184,1447,1472]" italics="true" pageId="109" pageNumber="110">e.g</emphasis>
|
||
.,
|
||
<bibRefCitation id="EF8C4B25007FFFFAFF02FA42FE3217D4" author="Nel, A. & Fleck, G." box="[206,412,1447,1473]" pageId="109" pageNumber="110" pagination="64 - 68" refId="ref59893" refString="Nel, A. & Fleck, G. (2012) A new genus and species of Sieblosiidae from the middle Miocene of Germany (Odonata: Epiproctophora). Zootaxa, 3582 (1), 64 - 68. https: // doi. org / 10.11646 / zootaxa. 3582.1.7" type="journal article" year="2012">Nel & Fleck 2012</bibRefCitation>
|
||
, their
|
||
<figureCitation id="13262A51007FFFFAFE2EFA42FD8817D4" box="[482,550,1447,1473]" captionStart="FIGURE 3" captionStartId="10.[151,250,1805,1829]" captionTargetBox="[151,1436,181,1781]" captionTargetId="figure-19@10.[151,1436,181,1781]" captionTargetPageId="10" captionText="FIGURE 3. Phenacolestes parallelus. A, specimen YPMIP-220974 of the Yale Peabody Museum; B, drawing from A, photograph provided by Peabody Museum personnel; C, holotype, UCM-4503 of the University of Colorado (Boulder) paleontology collections; D, drawing of C, photograph provided by University of Colorado personnel; E, UCM-4545, University of Colorado (Boulder). A–D to scale, E to scale, both scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558803" httpUri="https://zenodo.org/record/4558803/files/figure.png" pageId="109" pageNumber="110">Fig. 3</figureCitation>
|
||
). This is not present in any other specimen of
|
||
<taxonomicName id="4C1D4D57007FFFFAFBFDFA42FB7D17D4" authorityName="Archibald & Cannings" authorityYear="2021" box="[1073,1235,1447,1473]" class="Insecta" family="Sieblosiidae" genus="Whetwhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC6007FFFFAFBFDFA42FB7D17D4" box="[1073,1235,1447,1473]" italics="true" pageId="109" pageNumber="110">Whetwhetaksa</emphasis>
|
||
</taxonomicName>
|
||
, and it could be a crossvein adventitiously positioned between the costa and nodus in this specimen and not the subcosta.
|
||
</paragraph>
|
||
<paragraph id="8BA236D4007FFFFAFF0BFA0AFE781460" blockId="109.[151,1437,546,1653]" pageId="109" pageNumber="110">
|
||
We tentatively associate the
|
||
<taxonomicName id="4C1D4D57007FFFFAFDDFFA0AFD74141C" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[531,730,1519,1545]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae</taxonomicName>
|
||
with the
|
||
<taxonomicName id="4C1D4D57007FFFFAFC82FA0AFBB4141C" box="[846,1050,1519,1545]" kingdom="Animalia" pageId="109" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
|
||
by the many similarities of their wings (see description of the suborder, above). The wings of
|
||
<taxonomicName id="4C1D4D57007FFFFAFC9CF9F6FBB91438" authority="Archibald & Cannings & Erickson & Bybee & Mathewes, 2021" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[848,1047,1555,1581]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae</taxonomicName>
|
||
are more like those of the
|
||
<taxonomicName id="4C1D4D57007FFFFAFA87F9F6FEB51444" kingdom="Animalia" pageId="109" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
|
||
than any
|
||
<taxonomicName id="4C1D4D57007FFFFAFE42F9D2FDAA1444" box="[398,516,1591,1617]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="109" pageNumber="124" phylum="Arthropoda" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
|
||
, but await confirmation of this relationship by the discovery of a more complete fossil that includes the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |