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<document id="3C2D588DDBBC548C4C8A0026D6FA9B1D" ID-CLB-Dataset="37320" ID-DOI="10.5281/zenodo.272951" ID-GBIF-Dataset="e2c88f4c-3ac2-45f9-95e4-99b75561a081" ID-ISSN="1175-5326" ID-Zenodo-Dep="272951" ID-ZooBank="6D68A019-6F63-4AA4-A8B3-92D351F1F69B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484409208030" checkinUser="plazi" docAuthor="Van, Rob W. M." docDate="2017" docId="03A8001077EFFF15FF14A7F094B8FE4A" docLanguage="en" docName="zootaxa.4217.1.1.pdf.imf" docOrigin="Zootaxa 1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymeniacidon heliophila Wilson 1911" docType="treatment" docVersion="21" lastPageNumber="186" masterDocId="FF9178687757FFACFF83A36B907DFF9A" masterDocTitle="Sponges of the Guyana Shelf" masterLastPageNumber="225" masterPageNumber="1" pageNumber="185" updateTime="1698435695899" updateUser="ExternalLinkService">
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<mods:title id="C30F9F6A3430A8EE5B992E3A1288114A">Sponges of the Guyana Shelf</mods:title>
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<subSubSection id="C31BE28D77EFFF14FF14A7F0913FFB4C" pageId="184" pageNumber="185" type="nomenclature">
<paragraph id="8BBEB10677EFFF14FF14A7F092FEFB2F" blockId="184.[151,643,1179,1238]" box="[151,643,1179,1205]" pageId="184" pageNumber="185">
<heading id="D0F6066A77EFFF14FF14A7F092FEFB2F" bold="true" box="[151,643,1179,1205]" fontSize="11" level="1" pageId="184" pageNumber="185" reason="1">
<taxonomicName id="4C01CA8577EFFF14FF14A7F092FEFB2F" authority="Wilson, 1911" authorityName="Wilson" authorityYear="1911" box="[151,643,1179,1205]" class="Demospongiae" family="Halichondriidae" genus="Hymeniacidon" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="heliophila">
<emphasis id="B9756D1477EFFF14FF14A7F092FEFB2F" bold="true" box="[151,643,1179,1205]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF14A7F091B7FB2F" bold="true" box="[151,458,1179,1205]" italics="true" pageId="184" pageNumber="185">Hymeniacidon heliophila</emphasis>
(
<bibRefCitation id="EF90CCF777EFFF14FE5AA7F09201FB2F" author="Wilson" box="[473,636,1179,1205]" pageId="184" pageNumber="185" refString="Wilson, H. V. (1911) Development of sponges from dissociated tissue cells. Bulletin of the Bureau of Fisheries, 30, 1 - 30." type="journal article" year="1911">Wilson, 1911</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBEB10677EFFF14FF14A7D6913FFB4C" blockId="184.[151,643,1179,1238]" box="[151,322,1213,1238]" pageId="184" pageNumber="185">
<figureCitation id="133AAD8377EFFF14FF14A7D69166FB4C" box="[151,283,1213,1238]" captionStart="FIGURE 116" captionStartId="185.[151,250,1684,1706]" captionTargetBox="[147,1432,513,1662]" captionTargetId="figure@185.[147,1432,513,1662]" captionTargetPageId="185" captionText="FIGURE 116. Hymeniacidon heliophila (Wilson, 1911), a, habitus of RMNH Por. 9923 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c c 1, SEM images of the spicules, c, various shapes of styles, c 1, details of c." httpUri="https://zenodo.org/record/273066/files/figure.png" pageId="184" pageNumber="185">Figures 116</figureCitation>
ac
</paragraph>
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<subSubSection id="C31BE28D77EFFF14FF14A6699254FA09" pageId="184" pageNumber="185" type="reference_group">
<paragraph id="8BBEB10677EFFF14FF14A6699324FA82" blockId="184.[151,1436,1282,1427]" box="[151,857,1282,1304]" pageId="184" pageNumber="185">
<treatmentCitationGroup id="AB11962877EFFF14FF14A6699324FA82" box="[151,857,1282,1304]" pageId="184" pageNumber="185">
<taxonomicName id="4C01CA8577EFFF14FF14A6699210FA82" ID-CoL="53B85" authority="Wilson in Parker, 1910: 2" authorityName="Wilson in Parker" authorityPageNumber="2" authorityYear="1910" box="[151,621,1282,1304]" class="Demospongiae" family="Halichondriidae" genus="Stylotella" higherTaxonomySource="CoL" kingdom="Animalia" order="Suberitida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="heliophila">
<emphasis id="B9756D1477EFFF14FF14A669911EFA82" box="[151,355,1282,1304]" italics="true" pageId="184" pageNumber="185">Stylotella heliophila</emphasis>
Wilson in
<treatmentCitation id="0AA0971777EFFF14FE52A6699210FA82" author="Parker" box="[465,621,1282,1304]" page="2" pageId="184" pageNumber="185" year="1910">
<bibRefCitation id="EF90CCF777EFFF14FE52A6699228FA82" author="Parker" box="[465,597,1282,1304]" pageId="184" pageNumber="185" refString="Parker, G. H. (1910) The reactions of sponges, with a consideration of the origin of the nervous system. Journal of Experimental Zoology, 8, 1 - 41." type="journal article" year="1910">Parker, 1910</bibRefCitation>
: 2
</treatmentCitation>
</taxonomicName>
, fig. 1 (
<emphasis id="B9756D1477EFFF14FD3FA6689336FA82" box="[700,843,1282,1304]" italics="true" pageId="184" pageNumber="185">nomen nudum</emphasis>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBEB10677EFFF14FF14A64A9110FACC" blockId="184.[151,1436,1282,1427]" pageId="184" pageNumber="185">
<treatmentCitationGroup id="AB11962877EFFF14FF14A64A9110FACC" pageId="184" pageNumber="185">
<taxonomicName id="4C01CA8577EFFF14FF14A64A9262FAAD" ID-CoL="53B85" authority="Wilson, 1911: 13" authorityName="Wilson" authorityPageNumber="13" authorityYear="1911" box="[151,543,1313,1335]" class="Demospongiae" family="Halichondriidae" genus="Stylotella" higherTaxonomySource="CoL" kingdom="Animalia" order="Suberitida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="heliophila">
<emphasis id="B9756D1477EFFF14FF14A64A9118FAAD" box="[151,357,1313,1335]" italics="true" pageId="184" pageNumber="185">Stylotella heliophila</emphasis>
<treatmentCitation id="0AA0971777EFFF14FEEEA64A9262FAAD" author="Wilson" box="[365,543,1313,1335]" page="13" pageId="184" pageNumber="185" year="1911">
<bibRefCitation id="EF90CCF777EFFF14FEEEA64A9184FAAD" author="Wilson" box="[365,505,1313,1335]" pageId="184" pageNumber="185" refString="Wilson, H. V. (1911) Development of sponges from dissociated tissue cells. Bulletin of the Bureau of Fisheries, 30, 1 - 30." type="journal article" year="1911">Wilson, 1911</bibRefCitation>
: 13
</treatmentCitation>
</taxonomicName>
(proper description);
<treatmentCitation id="0AA0971777EFFF14FC80A64A9450FAAD" author="George" box="[771,1069,1313,1335]" page="147" pageId="184" pageNumber="185" year="1919">
<bibRefCitation id="EF90CCF777EFFF14FC80A64A9387FAAD" author="George" box="[771,1018,1313,1335]" pageId="184" pageNumber="185" refString="George, W. C. &amp; Wilson, H. V. (1919) Sponges of Beaufort (N. C.) harbor and vicinity. Bulletin of the Bureau of Fisheries, Washington, 36, 129 - 179." type="journal article" year="1919">George &amp; Wilson 1919</bibRefCitation>
: 147
</treatmentCitation>
, pls LVIII fig. 13, LIX figs 1819, LXVI fis 53ac.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBEB10677EFFF14FF14A6359254FA09" blockId="184.[151,1436,1282,1427]" pageId="184" pageNumber="185">
<treatmentCitationGroup id="AB11962877EFFF14FF14A6359254FA09" pageId="184" pageNumber="185">
<taxonomicName id="4C01CA8577EFFF14FF14A63591EAFAEE" box="[151,407,1374,1396]" class="Demospongiae" family="Halichondriidae" genus="Hymeniacidon" kingdom="Animalia" order="Halichondrida" pageId="184" pageNumber="185" phylum="Porifera" rank="species" species="heliophila">
<emphasis id="B9756D1477EFFF14FF14A63591EAFAEE" box="[151,407,1374,1396]" italics="true" pageId="184" pageNumber="185">Hymeniacidon heliophila</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AA0971777EFFF14FE26A63492D7FAEE" author="De" box="[421,682,1374,1396]" page="138" pageId="184" pageNumber="185" year="1936">
<bibRefCitation id="EF90CCF777EFFF14FE26A634920AFAEE" author="De" box="[421,631,1374,1396]" pageId="184" pageNumber="185" refString="De Laubenfels, M. W. (1936) A discussion of the sponge fauna of the Dry Tortugas, in particular, and the West Indies in general, with material for a revision of the families and orders of the Porifera. Carnegie Institution of Washington Publication, 467 (Tortugas Laboratory Paper, 30, 1 - 225." type="journal article" year="1936">De Laubenfels 1936</bibRefCitation>
: 138
</treatmentCitation>
;
<treatmentCitation id="0AA0971777EFFF14FD36A63593CCFAEE" author="Wiedenmayer" box="[693,945,1374,1396]" page="150" pageId="184" pageNumber="185" year="1977">
<bibRefCitation id="EF90CCF777EFFF14FD36A6359302FAEE" author="Wiedenmayer" box="[693,895,1374,1396]" pageId="184" pageNumber="185" refString="Wiedenmayer, F. (1977) Shallow-water sponges of the western Bahamas. Experientia Supplementum, 28, 1 - 287." type="journal article" year="1977">Wiedenmayer 1977</bibRefCitation>
: 150
</treatmentCitation>
, pl. 30 fig.8, text-fig.152;
<treatmentCitation id="0AA0971777EFFF14FB3DA63595E5FAEE" author="Diaz" box="[1214,1432,1374,1397]" page="298" pageId="184" pageNumber="185" year="1993">
<bibRefCitation id="EF90CCF777EFFF14FB3DA6359518FAEE" author="Diaz" box="[1214,1381,1374,1397]" pageId="184" pageNumber="185" refString="Diaz, M. C., Pomponi, S. A. &amp; Van Soest, R. W. M. (1993) A systematic revision of the central West Atlantic Halichondrida (Demospongiae, Porifera). Part III: Description of valid species. In: Uriz, M. - J. &amp; Rutzler, K. (Eds.), Recent Advances in Ecology and Systematics of Sponges. Scientia Marina, 57 (4), 283 - 306." type="journal article" year="1993">
Díaz
<emphasis id="B9756D1477EFFF14FB76A60B9554FAEE" box="[1269,1321,1374,1397]" italics="true" pageId="184" pageNumber="185">et al.</emphasis>
1993
</bibRefCitation>
: 298
</treatmentCitation>
, figs 26, 33; Muricy
<emphasis id="B9756D1477EFFF14FE12A61591B9FA09" box="[401,452,1405,1427]" italics="true" pageId="184" pageNumber="185">et al.</emphasis>
2011: 94.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77EFFF14FF14A6D49165F9DF" pageId="184" pageNumber="185" type="materials_examined">
<paragraph id="8BBEB10677EFFF14FF14A6D49165F9DF" blockId="184.[151,1436,1471,2037]" pageId="184" pageNumber="185">
<materialsCitation id="3B69BB5B77EFFF14FF14A6D493CAFA66" ID-GBIF-Occurrence="1424726572" collectingDate="1970-09-02" collectionCode="RMNH" country="Guyana" elevation="59" latitude="7.5667" location="Guyana" longLatPrecision="7" longitude="-57.2667" pageId="184" pageNumber="185" specimenCount="1">
<emphasis id="B9756D1477EFFF14FF14A6D491F5FA42" bold="true" box="[151,392,1471,1496]" pageId="184" pageNumber="185">Material examined.</emphasis>
<collectionCode id="ED1029C377EFFF14FE16A6AB9194FA4D" box="[405,489,1472,1495]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="184" pageNumber="185">RMNH</collectionCode>
Por. 9834,
<collectingCountry id="F316F19677EFFF14FDFCA6AB92A4FA42" box="[639,729,1472,1496]" name="Guyana" pageId="184" pageNumber="185">Guyana</collectingCountry>
, Luymes
<collectingCountry id="F316F19677EFFF14FCE4A6AB93C3FA42" box="[871,958,1472,1496]" name="Guyana" pageId="184" pageNumber="185">Guyana</collectingCountry>
Shelf Expedition, station 65,
<geoCoordinate id="EE35D7C177EFFF14FAB3A6D495E1FA42" box="[1328,1436,1471,1496]" direction="north" orientation="latitude" pageId="184" pageNumber="185" precision="5" value="7.5667">7.5667°N</geoCoordinate>
<geoCoordinate id="EE35D7C177EFFF14FF14A68F916AFA67" box="[151,279,1508,1533]" direction="west" orientation="longitude" pageId="184" pageNumber="185" precision="5" value="-57.2667">57.2667°W</geoCoordinate>
, depth
<quantity id="4CF91CE377EFFF14FEFBA68F91C1FA66" box="[376,444,1508,1533]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="184" pageNumber="185" unit="m" value="59.0">
<elevation id="002C563577EFFF14FEFBA68F91C1FA66" box="[376,444,1508,1533]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="184" pageNumber="185" unit="m" value="59.0">59 m</elevation>
</quantity>
, sandy shelly bottom,
<date id="FFBF97C677EFFF14FD5BA68F93CAFA66" box="[728,951,1508,1533]" pageId="184" pageNumber="185" value="1970-09-02">
<collectingDate id="EFFB6E2E77EFFF14FD5BA68F93CAFA66" box="[728,951,1508,1533]" pageId="184" pageNumber="185" value="1970-09-02">2 September 1970</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B69BB5B77EFFF14FC49A68E9169F9DF" ID-GBIF-Occurrence="1424726560" collectingDate="1969-04-16" collectionCode="RMNH" collectorName="Luymes O. C. P. S." country="Suriname" elevation="130" latitude="7.3083" location="Suriname" longLatPrecision="7" longitude="-54.1667" pageId="184" pageNumber="185" specimenCount="1">
<collectionCode id="ED1029C377EFFF14FC49A68E9463FA66" box="[970,1054,1509,1532]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="184" pageNumber="185">RMNH</collectionCode>
Por. 9923,
<collectingCountry id="F316F19677EFFF14FB34A68F9558FA67" box="[1207,1317,1508,1533]" name="Suriname" pageId="184" pageNumber="185">Suriname</collectingCountry>
,
<collectorName id="26F4D4D077EFFF14FABEA68F9088F9BA" pageId="184" pageNumber="185">Luymes O.C.P.S.</collectorName>
II
<collectingCountry id="F316F19677EFFF14FE9FA563910EF9BA" box="[284,371,1544,1568]" name="Guyana" pageId="184" pageNumber="185">Guyana</collectingCountry>
Shelf Expedition, station M97,
<geoCoordinate id="EE35D7C177EFFF14FD6BA5639328F9BA" box="[744,853,1544,1568]" direction="north" orientation="latitude" pageId="184" pageNumber="185" precision="5" value="7.3083">7.3083°N</geoCoordinate>
<geoCoordinate id="EE35D7C177EFFF14FCDCA56C939DF9BA" box="[863,992,1543,1568]" direction="west" orientation="longitude" pageId="184" pageNumber="185" precision="5" value="-54.1667">54.1667°W</geoCoordinate>
, depth
<quantity id="4CF91CE377EFFF14FBB5A56394FFF985" box="[1078,1154,1544,1568]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="184" pageNumber="185" unit="m" value="130.0">
<elevation id="002C563577EFFF14FBB5A56394FFF985" box="[1078,1154,1544,1568]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="184" pageNumber="185" unit="m" value="130.0">130 m</elevation>
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, bottom coarse sand,
<date id="FFBF97C677EFFF14FA03A56C9169F9DF" pageId="184" pageNumber="185" value="1969-04-16">
<collectingDate id="EFFB6E2E77EFFF14FA03A56C9169F9DF" pageId="184" pageNumber="185" value="1969-04-16">16 April 1969</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77EFFF14FF44A52494F6F962" pageId="184" pageNumber="185" type="description">
<paragraph id="8BBEB10677EFFF14FF44A52494E5F917" blockId="184.[151,1436,1471,2037]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF44A5249124F9F2" bold="true" box="[199,345,1615,1640]" pageId="184" pageNumber="185">Description.</emphasis>
(
<figureCitation id="133AAD8377EFFF14FEEAA52491B4F9F2" box="[361,457,1615,1640]" captionStart="FIGURE 116" captionStartId="185.[151,250,1684,1706]" captionTargetBox="[147,1432,513,1662]" captionTargetId="figure@185.[147,1432,513,1662]" captionTargetPageId="185" captionText="FIGURE 116. Hymeniacidon heliophila (Wilson, 1911), a, habitus of RMNH Por. 9923 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c c 1, SEM images of the spicules, c, various shapes of styles, c 1, details of c." httpUri="https://zenodo.org/record/273066/files/figure.png" pageId="184" pageNumber="185">Fig. 116</figureCitation>
a) Irregular stringy lobes and thin branches, presumably detached from a buried main body, which was not collected. Also thin encrustations on dead shells and carbonate rubble.
</paragraph>
<paragraph id="8BBEB10677EFFF14FF44A5FC956FF92A" blockId="184.[151,1436,1471,2037]" box="[199,1298,1687,1712]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF44A5FC9148F92A" bold="true" box="[199,309,1687,1712]" pageId="184" pageNumber="185">Skeleton.</emphasis>
Loosely confused arrangement in the choanosome, tangential at the surface (
<figureCitation id="133AAD8377EFFF14FB15A5FC948BF92A" box="[1174,1270,1687,1712]" captionStart="FIGURE 116" captionStartId="185.[151,250,1684,1706]" captionTargetBox="[147,1432,513,1662]" captionTargetId="figure@185.[147,1432,513,1662]" captionTargetPageId="185" captionText="FIGURE 116. Hymeniacidon heliophila (Wilson, 1911), a, habitus of RMNH Por. 9923 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c c 1, SEM images of the spicules, c, various shapes of styles, c 1, details of c." httpUri="https://zenodo.org/record/273066/files/figure.png" pageId="184" pageNumber="185">Fig. 116</figureCitation>
b).
</paragraph>
<paragraph id="8BBEB10677EFFF14FF44A5D7921CF94F" blockId="184.[151,1436,1471,2037]" box="[199,609,1724,1749]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF44A5D7914DF94F" bold="true" box="[199,304,1724,1749]" pageId="184" pageNumber="185">Spicules.</emphasis>
(
<figureCitation id="133AAD8377EFFF14FEBCA5D791D8F94F" box="[319,421,1724,1749]" captionStart="FIGURE 116" captionStartId="185.[151,250,1684,1706]" captionTargetBox="[147,1432,513,1662]" captionTargetId="figure@185.[147,1432,513,1662]" captionTargetPageId="185" captionText="FIGURE 116. Hymeniacidon heliophila (Wilson, 1911), a, habitus of RMNH Por. 9923 (scale bar = 1 cm), b, light microscopic image of surface skeleton, c c 1, SEM images of the spicules, c, various shapes of styles, c 1, details of c." httpUri="https://zenodo.org/record/273066/files/figure.png" pageId="184" pageNumber="185">Figs 116</figureCitation>
c,c1) Styles only.
</paragraph>
<paragraph id="8BBEB10677EFFF14FF44A5B494F6F962" blockId="184.[151,1436,1471,2037]" box="[199,1163,1759,1784]" pageId="184" pageNumber="185">
Styles, straight or slightly curved, often with faint tyle, 320
<emphasis id="B9756D1477EFFF14FCE8A58B93EEF962" box="[875,915,1760,1784]" italics="true" pageId="184" pageNumber="185">411</emphasis>
479
<date id="FFBF97C677EFFF14FC52A58B938FF962" box="[977,1010,1760,1784]" pageId="184" pageNumber="185">x 3</date>
<emphasis id="B9756D1477EFFF14FB80A5B4945BF962" box="[1027,1062,1759,1784]" italics="true" pageId="184" pageNumber="185">7.6</emphasis>
9.5 µm.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77EFFF14FF44A46F9268F812" pageId="184" pageNumber="185" type="distribution">
<paragraph id="8BBEB10677EFFF14FF44A46F9268F812" blockId="184.[151,1436,1471,2037]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF44A46F918FF887" bold="true" box="[199,498,1796,1821]" pageId="184" pageNumber="185">Distribution and ecology.</emphasis>
Originally described from
<collectingRegion id="49C57FE477EFFF14FCA3A46F93B6F887" box="[800,971,1796,1821]" country="United States of America" name="North Carolina" pageId="184" pageNumber="185">
North
<collectingRegion id="49C57FE477EFFF14FCEBA46F93B6F887" box="[872,971,1796,1821]" country="Puerto Rico" name="Carolina" pageId="184" pageNumber="185">Carolina</collectingRegion>
</collectingRegion>
, the species has been reported throughout the Caribbean region and also from
<collectingCountry id="F316F19677EFFF14FDCDA44C92EFF8DA" box="[590,658,1831,1856]" name="Brazil" pageId="184" pageNumber="185">Brazil</collectingCountry>
(Muricy
<emphasis id="B9756D1477EFFF14FC8AA442933AF8DA" box="[777,839,1831,1856]" italics="true" pageId="184" pageNumber="185">et al.</emphasis>
2011). Predominantly known from shallow-water, intertidal and mangrove habitats. The present records from sandy bottoms at
<quantity id="4CF91CE377EFFF14FC72A427941FF8FE" box="[1009,1122,1868,1893]" metricMagnitude="1" metricUnit="m" metricValue="9.45" metricValueMax="13.0" metricValueMin="5.9" pageId="184" pageNumber="185" unit="m" value="94.5" valueMax="130.0" valueMin="59.0">59130 m</quantity>
depth are by far the deepest occurrence known for the species.
</paragraph>
</subSubSection>
<subSubSection id="C31BE28D77EFFF15FF44A4FF94B8FE4A" lastPageId="185" lastPageNumber="186" pageId="184" pageNumber="185" type="discussion">
<paragraph id="8BBEB10677EFFF14FF44A4FF9196F86F" blockId="184.[151,1436,1471,2037]" pageId="184" pageNumber="185">
<emphasis id="B9756D1477EFFF14FF44A4FF9146F837" bold="true" box="[199,315,1940,1965]" pageId="184" pageNumber="185">Remarks.</emphasis>
In view of the deep occurrence there is some doubt about the identification. However, similarity with the original description and subsequent morphological information does not warrant erecting a new species for the
<collectingCountry id="F316F19677EFFF14FF6BA4B79142F86E" box="[232,319,2012,2036]" name="Guyana" pageId="184" pageNumber="185">Guyana</collectingCountry>
Shelf material.
</paragraph>
<paragraph id="8BBEB10677EEFF15FF44A3FC94B8FE4A" blockId="185.[151,1437,151,464]" pageId="185" pageNumber="186">
<materialsCitation id="3B69BB5B77EEFF15FF44A3FC9405FEDA" ID-GBIF-Occurrence="1424726557" collectionCode="ICZN" collectorName="Authorship &amp; Parker &amp; De Laubenfels &amp; However &amp; H. V. Wilson &amp; Apart" location="Central West" pageId="185" pageNumber="186" specimenCount="1" stateProvince="Central West" typeStatus="holotype">
<collectorName id="26F4D4D077EEFF15FF44A3FC9138FF2A" box="[199,325,151,176]" pageId="185" pageNumber="186">Authorship</collectorName>
of this common
<collectingRegion id="49C57FE477EEFF15FDA1A3FC92C6FF35" box="[546,699,151,176]" isEnumeration="true" pageId="185" pageNumber="186">Central West</collectingRegion>
Atlantic species has been assigned to
<bibRefCitation id="EF90CCF777EEFF15FB1CA3FC953CFF2A" author="Parker" box="[1183,1345,151,176]" pageId="185" pageNumber="186" refString="Parker, G. H. (1910) The reactions of sponges, with a consideration of the origin of the nervous system. Journal of Experimental Zoology, 8, 1 - 41." type="journal article" year="1910">
<collectorName id="26F4D4D077EEFF15FB1CA3FC9495FF2A" box="[1183,1256,151,176]" pageId="185" pageNumber="186">Parker</collectorName>
(1910)
</bibRefCitation>
by
<bibRefCitation id="EF90CCF777EEFF15FAF8A3F39119FF4F" pageId="185" pageNumber="186" refString="De Laubenfels, M. W. (1936) A discussion of the sponge fauna of the Dry Tortugas, in particular, and the West Indies in general, with material for a revision of the families and orders of the Porifera. Carnegie Institution of Washington Publication, 467 (Tortugas Laboratory Paper, 30, 1 - 225." type="journal article">
<collectorName id="26F4D4D077EEFF15FAF8A3F39169FF4F" pageId="185" pageNumber="186">De Laubenfels</collectorName>
(1936)
</bibRefCitation>
(p.138).
<collectorName id="26F4D4D077EEFF15FE49A3D6924FFF4E" box="[458,562,189,212]" pageId="185" pageNumber="186">However</collectorName>
,
<collectorName id="26F4D4D077EEFF15FDBEA3D792FBFF4F" box="[573,646,188,213]" pageId="185" pageNumber="186">Parker</collectorName>
himself (1910) (p. 2) made it abundantly clear that
<collectorName id="26F4D4D077EEFF15FB46A3D69533FF4F" box="[1221,1358,188,213]" pageId="185" pageNumber="186">H.V. Wilson</collectorName>
was in the process of describing this species and he only borrowed the name for his physiological study.
<collectorName id="26F4D4D077EEFF15FA8BA38B9535FF62" box="[1288,1352,224,248]" pageId="185" pageNumber="186">Apart</collectorName>
from a figure of the species, there is no taxonomic description, nor a
<typeStatus id="54BA0FA477EEFF15FC0EA26E93C0FE87" box="[909,957,261,285]" pageId="185" pageNumber="186">type</typeStatus>
specimen assignment in Parkers paper. Effectively, Parkers use of the name is a
<emphasis id="B9756D1477EEFF15FD16A2429345FEDA" box="[661,824,295,320]" italics="true" pageId="185" pageNumber="186">nomen nudum</emphasis>
in the sense of the
<collectionCode id="ED1029C377EEFF15FBB0A2439405FEDA" box="[1075,1144,296,320]" pageId="185" pageNumber="186">ICZN</collectionCode>
</materialsCitation>
. Authorship and year of publication should go to
<bibRefCitation id="EF90CCF777EEFF15FE32A2279228FEFE" author="Wilson" box="[433,597,332,357]" pageId="185" pageNumber="186" refString="Wilson, H. V. (1911) Development of sponges from dissociated tissue cells. Bulletin of the Bureau of Fisheries, 30, 1 - 30." type="journal article" year="1911">Wilson (1911)</bibRefCitation>
, who gave a taxonomic description including information on the spicules. ICZN Art. 50.1 clearly stipulates the case that Parkers (1910) referral to Wilson as the author of a forthcoming publication, in which the species is to be properly described, is a valid nomenclatorial procedure. In conclusion
<taxonomicName id="4C01CA8577EEFF15FF14A2DC921CFE4A" authority="Wilson, 1911" authorityName="Wilson" authorityYear="1911" box="[151,609,439,464]" class="Demospongiae" family="Halichondriidae" genus="Hymeniacidon" kingdom="Animalia" order="Halichondrida" pageId="185" pageNumber="186" phylum="Porifera" rank="species" species="heliophila">
<emphasis id="B9756D1477EEFF15FF14A2DC91CEFE4A" box="[151,435,439,464]" italics="true" pageId="185" pageNumber="186">Hymeniacidon heliophila</emphasis>
(
<bibRefCitation id="EF90CCF777EEFF15FE42A2DC9224FE4A" author="Wilson" box="[449,601,439,464]" pageId="185" pageNumber="186" refString="Wilson, H. V. (1911) Development of sponges from dissociated tissue cells. Bulletin of the Bureau of Fisheries, 30, 1 - 30." type="journal article" year="1911">Wilson, 1911</bibRefCitation>
)
</taxonomicName>
is the proper author-year combination for this species.
</paragraph>
</subSubSection>
</treatment>
</document>