328 lines
40 KiB
XML
328 lines
40 KiB
XML
<document id="EA29E5ABDC516B81E0FC656097B6613E" ID-ISSN="2336-9744" ID-ZooBank="urn:lsid:zoobank.org:pub:EAEB7103-1655-4464-B258-481D881A45DC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702514279291" checkinUser="felipe" docAuthor="- & Grosser, Clemens" docDate="2015" docId="03A387EEA3369071FF3EFA3DB272FB1A" docLanguage="en" docName="EcolMontenegrina.2.1.29-41.pdf" docOrigin="Ecologica Montenegrina 2 (1)" docStyle="DocumentStyle:17621BF4B6DAF9FC3B559F0A069E6C5F.2:EcolMontenegrina.2014-2016.journal_article" docStyleId="17621BF4B6DAF9FC3B559F0A069E6C5F" docStyleName="EcolMontenegrina.2014-2016.journal_article" docStyleVersion="2" docTitle="Trocheta taunensis - & Grosser 2015, n. sp." docType="treatment" docVersion="1" lastPageNumber="40" masterDocId="FF9AFF96A33E907AFFA9FF98B124FF95" masterDocTitle="Differentiation of some similar species of the subfamily Trochetinae (Hirudinida: Erpobdellidae)" masterLastPageNumber="41" masterPageNumber="29" pageNumber="37" updateTime="1702514848032" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="600DF8B59E58C56878279479832C84A3">Differentiation of some similar species of the subfamily Trochetinae (Hirudinida: Erpobdellidae)</mods:title>
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<mods:namePart id="C458385A6856E025DFAB19D874903A31">Grosser, Clemens</mods:namePart>
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<mods:typeOfResource id="3633E8A81C442583818A38D1B5D26186">text</mods:typeOfResource>
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<mods:title id="F56BF5D471D4935E7C58A679F3DC3F60">Ecologica Montenegrina</mods:title>
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<mods:part id="442B6600805EFC621003D6AC85DBB326">
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<mods:date id="B6D6340DD1702AA25CD1BFB79A673E96">2015</mods:date>
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<mods:number id="27C70E10901F5A72073231DE214A463E">2015-01-08</mods:number>
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<mods:number id="CDF51FD44CC45C26BC38423EFB478D59">2</mods:number>
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<mods:identifier id="29DF850A188EF005BBDEDBBBB66B5448" type="ISSN">2336-9744</mods:identifier>
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<mods:identifier id="27BCC608407A9E110D2B02135A921F9F" type="ZooBank">urn:lsid:zoobank.org:pub:EAEB7103-1655-4464-B258-481D881A45DC</mods:identifier>
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<treatment id="03A387EEA3369071FF3EFA3DB272FB1A" LSID="urn:lsid:plazi:treatment:03A387EEA3369071FF3EFA3DB272FB1A" httpUri="http://treatment.plazi.org/id/03A387EEA3369071FF3EFA3DB272FB1A" lastPageId="11" lastPageNumber="40" pageId="8" pageNumber="37">
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<subSubSection id="C3106573A3369072FF3EFA3DB0EDFA2A" box="[151,457,1445,1471]" pageId="8" pageNumber="37" type="nomenclature">
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<paragraph id="8BB536F8A3369072FF3EFA3DB0EDFA2A" blockId="8.[151,457,1445,1505]" box="[151,457,1445,1471]" pageId="8" pageNumber="37">
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<heading id="D0FD8194A3369072FF3EFA3DB0EDFA2A" box="[151,457,1445,1471]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="37" reason="2">
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<emphasis id="B97EEAEAA3369072FF3EFA3DB0EDFA2A" bold="true" box="[151,457,1445,1471]" pageId="8" pageNumber="37">
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<taxonomicName id="4C0A4D7BA3369072FF3EFA3DB0A4FA2A" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[151,384,1445,1471]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="8" pageNumber="37" phylum="Annelida" rank="species" species="taunensis" status="sp. nov.">
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<emphasis id="B97EEAEAA3369072FF3EFA3DB0A4FA2A" bold="true" box="[151,384,1445,1471]" italics="true" pageId="8" pageNumber="37">Trocheta taunensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D5791A3369072FE2EFA3DB0EDFA2A" box="[391,457,1445,1471]" pageId="8" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3106573A3369072FF3EFA5FB008FA74" box="[151,300,1479,1505]" pageId="8" pageNumber="37" type="description">
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<paragraph id="8BB536F8A3369072FF3EFA5FB008FA74" blockId="8.[151,457,1445,1505]" box="[151,300,1479,1505]" pageId="8" pageNumber="37">
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(
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<figureCitation id="13312A7DA3369072FF09FA5FB1D5FA74" box="[160,241,1479,1505]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="8" pageNumber="37">Figs. 8</figureCitation>
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AE)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3106573A3369072FF3EF991B401F93E" pageId="8" pageNumber="37" type="materials_examined">
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<paragraph id="8BB536F8A3369072FF3EF991B53DF9F2" blockId="8.[151,1444,1545,1639]" pageId="8" pageNumber="37">
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<emphasis id="B97EEAEAA3369072FF3EF991B0A1F9B6" bold="true" box="[151,389,1545,1571]" pageId="8" pageNumber="37">Material examined</emphasis>
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:
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<materialsCitation id="3B623CA5A3369072FE3EF992B5C8F9D3" collectingDate="2000-04-19" collectionCode="SMF" collectorName="C. Grosser." country="Germany" location="river Wisper" municipality="Taunus Mt." pageId="8" pageNumber="37" specimenCode="SMF 19957" specimenCount="1" stateProvince="Hessen" typeStatus="holotype">
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<typeStatus id="54B1885AA3369072FE3EF992B320F9B1" box="[407,516,1546,1572]" pageId="8" pageNumber="37" type="holotype">Holotype</typeStatus>
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(
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<specimenCode id="DBAC9E83A3369072FDBCF992B38DF9B1" box="[533,681,1546,1572]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="37">SMF 19957</specimenCode>
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): body length
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<quantity id="4CF29B1DA3369072FCFDF992B29EF9B1" box="[852,954,1546,1572]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="37" unit="mm" value="101.0">101 mm</quantity>
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, width
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<quantity id="4CF29B1DA3369072FBBAF992B578F9B1" box="[1043,1116,1546,1572]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="37" unit="mm" value="6.0">6 mm</quantity>
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;
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<collectingCountry id="F31D7668A3369072FBDBF992B5C6F9B1" box="[1138,1250,1546,1572]" name="Germany" pageId="8" pageNumber="37">Germany</collectingCountry>
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,
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<collectingRegion id="49CEF81AA3369072FB59F992B41DF9B1" box="[1264,1337,1546,1572]" country="Germany" name="Hessen" pageId="8" pageNumber="37">Hesse</collectingRegion>
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,
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<collectingMunicipality id="6BD1AC82A3369072FAEFF992B1E7F9D3" pageId="8" pageNumber="37">Taunus Mt.</collectingMunicipality>
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,
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<location id="8ED56023A3369072FF7CF9B4B04AF9D3" LSID="urn:lsid:plazi:treatment:03A387EEA3369071FF3EFA3DB272FB1A:8ED56023A3369072FF7CF9B4B04AF9D3" box="[213,366,1580,1606]" country="Germany" municipality="Taunus Mt." name="river Wisper" pageId="8" pageNumber="37" stateProvince="Hessen">river Wisper</location>
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<locationDeviation id="9A072FDAA3369072FED3F9B4B271F9D3" box="[378,853,1580,1606]" firstLocation="Geroldstein" pageId="8" pageNumber="37" secondLocation="Kammerburg">between Geroldstein and Kammerburg</locationDeviation>
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,
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<date id="FFB41038A3369072FCCFF9B4B53DF9D3" box="[870,1049,1580,1606]" pageId="8" pageNumber="37" value="2000-04-19">
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<collectingDate id="EFF0E9D0A3369072FCCFF9B4B53DF9D3" box="[870,1049,1580,1606]" pageId="8" pageNumber="37" value="2000-04-19">19 April 2000</collectingDate>
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</date>
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, leg.
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<collectorName id="26FF532EA3369072FBC8F9B4B5C8F9D3" box="[1121,1260,1580,1606]" pageId="8" pageNumber="37">C. Grosser.</collectorName>
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</materialsCitation>
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<typeStatus id="54B1885AA3369072FB51F9B4B44BF9D3" box="[1272,1391,1580,1606]" pageId="8" pageNumber="37" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D0CFD71A3369072FAD6F9B4B030F9F2" count="14" pageId="8" pageNumber="37" type="generic">14 specimens</specimenCount>
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,
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||
<materialsCitation id="3B623CA5A3369072FE89F9D5B532F9F2" box="[288,1046,1613,1639]" collectingDate="2000-04-19" collectionCode="SMF" collectorName="C. Grosser." country="Germany" location="river Wisper" municipality="Taunus Mt." pageId="8" pageNumber="37" specimenCode="SMF 19958" specimenCount="4" stateProvince="Hessen" typeStatus="paratype">
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||
same locality and data as
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<typeStatus id="54B1885AA3369072FDE4F9D5B390F9F2" box="[589,692,1613,1639]" pageId="8" pageNumber="37" type="holotype">holotype</typeStatus>
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||
(
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||
<specimenCount id="9D0CFD71A3369072FD6BF9D5B256F9F2" box="[706,882,1613,1639]" count="4" pageId="8" pageNumber="37" type="generic" typeStatus="paratype">four paratypes</specimenCount>
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||
:
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<specimenCode id="DBAC9E83A3369072FCD5F9D5B528F9F2" box="[892,1036,1613,1639]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="37">SMF 19958</specimenCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BB536F8A3369072FF3EF909B401F93E" blockId="8.[151,1317,1681,1707]" box="[151,1317,1681,1707]" pageId="8" pageNumber="37">
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<emphasis id="B97EEAEAA3369072FF3EF909B069F93E" bold="true" box="[151,333,1681,1707]" pageId="8" pageNumber="37">Locus typicus:</emphasis>
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||
<collectingCountry id="F31D7668A3369072FEFDF909B0E0F93E" box="[340,452,1681,1707]" name="Germany" pageId="8" pageNumber="37">Germany</collectingCountry>
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||
,
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<collectingRegion id="49CEF81AA3369072FE79F909B33DF93E" box="[464,537,1681,1707]" country="Germany" name="Hessen" pageId="8" pageNumber="37">Hesse</collectingRegion>
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, Taunus Mt., river Wisper between Geroldstein and Kammerburg.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3106573A3369071FF3EF94CB273FE3C" lastPageId="11" lastPageNumber="40" pageId="8" pageNumber="37" type="description">
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||
<paragraph id="8BB536F8A3369072FF3EF94CB00CF97B" blockId="8.[151,1444,1748,2044]" box="[151,296,1748,1774]" pageId="8" pageNumber="37">
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<emphasis id="B97EEAEAA3369072FF3EF94CB00CF97B" bold="true" box="[151,296,1748,1774]" pageId="8" pageNumber="37">Description</emphasis>
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</paragraph>
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<paragraph id="8BB536F8A3369072FF3EF96EB061F869" blockId="8.[151,1444,1748,2044]" pageId="8" pageNumber="37">
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<emphasis id="B97EEAEAA3369072FF3EF96EB03DF89A" box="[151,281,1782,1807]" italics="true" pageId="8" pageNumber="37">Annulation</emphasis>
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: Typical
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<taxonomicName id="4C0A4D7BA3369072FE39F96EB0DEF89A" authorityName="Dutrochet" authorityYear="1817" box="[400,506,1782,1807]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="8" pageNumber="37" phylum="Annelida" rank="genus">
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<emphasis id="B97EEAEAA3369072FE39F96EB0DEF89A" box="[400,506,1782,1807]" italics="true" pageId="8" pageNumber="37">Trocheta</emphasis>
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</taxonomicName>
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like annulation. The somites of the
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<typeStatus id="54B1885AA3369072FC1CF96EB538F885" box="[949,1052,1782,1808]" pageId="8" pageNumber="37" type="holotype">holotype</typeStatus>
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are heteronomously subdivided into three broadened (b2, a2, b5) and five narrow (c1, c2, c11, d23, d24) annuli. This annulation correspond to the diagnosis of
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<taxonomicName id="4C0A4D7BA3369072FED5F8A2B0C2F8C6" authorityName="Dutrochet" authorityYear="1817" box="[380,486,1850,1875]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="8" pageNumber="37" phylum="Annelida" rank="genus">
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<emphasis id="B97EEAEAA3369072FED5F8A2B0C2F8C6" box="[380,486,1850,1875]" italics="true" pageId="8" pageNumber="37">Trocheta</emphasis>
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</taxonomicName>
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and justify the classification to this genus (Sawyer 1886,
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<bibRefCitation id="EF9B4B09A3369072FB33F8A2B1F2F8E0" author="Nesemann, H. & Neubert, E." pageId="8" pageNumber="37" pagination="2" refId="ref5124" refString="Nesemann, H. & Neubert, E. (1999) Annelida, Clitellata: Branchiobdellida, Acanthobdellea, Hirudinea. - Susswasserfauna von Mitteleuropa 6 / 2, Heidelberg." type="journal article" year="1999">Nesemann & Neubert 1999</bibRefCitation>
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). The male genital pore is in furrow b2/a2 and the female is in furrow b5/c11. The genital pores are separated by two full annuli. Annulus b5 is slightly subdivided (
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<figureCitation id="13312A7DA3369072FC15F8E5B53AF802" box="[956,1054,1917,1943]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="8" pageNumber="37">Fig. 8C</figureCitation>
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||
). Some
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<typeStatus id="54B1885AA3369072FB2EF8E5B5D3F802" box="[1159,1271,1917,1943]" pageId="8" pageNumber="37" type="paratype">paratypes</typeStatus>
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show a high tendency to split annulus b5 into annuli c9 and c10. In these specimens the genital pores are separated by three annuli. Sometimes, annulus a2 is also subdivided. In this case, the male and female pores are separated by four annuli.
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</paragraph>
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<caption id="DF756670A3379073FF3EF832B0DAF86A" pageId="9" pageNumber="38" startId="9.[151,227,1962,1986]" targetBox="[168,1392,188,1940]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8BB536F8A3379073FF3EF832B0DAF86A" blockId="9.[151,1443,1962,2047]" pageId="9" pageNumber="38">
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||
<emphasis id="B97EEAEAA3379073FF3EF832B312F857" bold="true" box="[151,566,1962,1986]" pageId="9" pageNumber="38">
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Figure 8.
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<taxonomicName id="4C0A4D7BA3379073FEA1F832B0F9F857" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[264,477,1962,1986]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="9" pageNumber="38" phylum="Annelida" rank="species" species="taunensis" status="sp. nov.">
|
||
<emphasis id="B97EEAEAA3379073FEA1F832B0F9F857" bold="true" box="[264,477,1962,1986]" italics="true" pageId="9" pageNumber="38">Trocheta taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3379073FE4FF832B301F857" box="[486,549,1962,1986]" pageId="9" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
</emphasis>
|
||
A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium.
|
||
</paragraph>
|
||
</caption>
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||
<caption id="DF756670A3349070FF3EFA6BB451F99E" box="[151,1397,1523,1547]" pageId="10" pageNumber="39" startId="10.[151,227,1523,1547]" targetBox="[301,1366,216,1472]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BB536F8A3349070FF3EFA6BB451F99E" blockId="10.[151,1397,1523,1547]" box="[151,1397,1523,1547]" pageId="10" pageNumber="39">
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||
<emphasis id="B97EEAEAA3349070FF3EFA6BB1DAF99E" bold="true" box="[151,254,1523,1547]" pageId="10" pageNumber="39">Figure 9.</emphasis>
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<taxonomicName id="4C0A4D7BA3349070FEADFA6BB368F99E" authority=", Bystrica" authorityName="Bystrica" box="[260,588,1523,1547]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="10" pageNumber="39" phylum="Annelida" rank="species" species="cylindrical">
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||
<emphasis id="B97EEAEAA3349070FEADFA6BB0CEF99E" bold="true" box="[260,490,1523,1547]" pageId="10" pageNumber="39">
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||
<emphasis id="B97EEAEAA3349070FEADFA6BB0C7F99E" bold="true" box="[260,483,1523,1547]" italics="true" pageId="10" pageNumber="39">Trocheta cylindrical</emphasis>
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,
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</emphasis>
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Bystrica
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</taxonomicName>
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, Slovakia: A shape and extension of the sexual organs, B – genital atrium.
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</paragraph>
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</caption>
|
||
<paragraph id="8BB536F8A3349070FF3EF9CFB057F907" blockId="10.[151,1443,1623,1682]" pageId="10" pageNumber="39">
|
||
<emphasis id="B97EEAEAA3349070FF3EF9CFB1E2F9E5" box="[151,198,1623,1648]" italics="true" pageId="10" pageNumber="39">Size</emphasis>
|
||
: Large leeches, preserved specimens up to
|
||
<quantity id="4CF29B1DA3349070FD76F9CFB267F9E4" box="[735,835,1623,1649]" metricMagnitude="-1" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="39" unit="mm" value="114.0">114 mm</quantity>
|
||
(the largest preserved
|
||
<typeStatus id="54B1885AA3349070FBF7F9CFB5ECF9E4" box="[1118,1224,1623,1649]" pageId="10" pageNumber="39" type="paratype">paratype</typeStatus>
|
||
: body length
|
||
<quantity id="4CF29B1DA3349070FAD9F9CFB1E3F907" metricMagnitude="-1" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="39" unit="mm" value="114.0">114 mm</quantity>
|
||
, width
|
||
<quantity id="4CF29B1DA3349070FEB7F9E0B043F907" box="[286,359,1656,1682]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="39" unit="mm" value="6.0">6 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3349070FF3EF924B096F8C8" blockId="10.[151,1444,1724,1885]" pageId="10" pageNumber="39">
|
||
<emphasis id="B97EEAEAA3349070FF3EF924B1D0F940" box="[151,244,1724,1749]" italics="true" pageId="10" pageNumber="39">Habitus</emphasis>
|
||
: The anterior part of the body is cylindrical, the posterior part is slightly flattened. The dorsal surface is convex in their total run. Lateral keels exist, special distinct in the last third of the body. The keels are small and not curved dorsally. The upper lip of the cranial sucker is elongated (
|
||
<figureCitation id="13312A7DA3349070FB28F967B5FDF88C" box="[1153,1241,1791,1817]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="10" pageNumber="39">Figs. 8</figureCitation>
|
||
AB). The body surface is smooth, only few very small and inconspicuous papillae are visible. Eyes are not visible on the preserved
|
||
<typeStatus id="54B1885AA3349070FEB9F8DBB067F8C8" box="[272,323,1859,1885]" pageId="10" pageNumber="39">type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3349070FF3EF81EB2A9F857" blockId="10.[151,1443,1926,1986]" pageId="10" pageNumber="39">
|
||
<emphasis id="B97EEAEAA3349070FF3EF81EB1CEF80A" box="[151,234,1926,1951]" italics="true" pageId="10" pageNumber="39">Colour</emphasis>
|
||
: The colour of dorsal surface is unicolour greyish to bright reddish brown. Preserved specimens are white to pale grey. Dark lines, stripes or yellow spots are absent.
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3359071FF3EFF5DB273FE3C" blockId="11.[151,1444,197,425]" pageId="11" pageNumber="40">
|
||
<emphasis id="B97EEAEAA3359071FF3EFF5DB01AFF4B" box="[151,318,197,222]" italics="true" pageId="11" pageNumber="40">Sexual organs</emphasis>
|
||
: The genital atrium is typical
|
||
<taxonomicName id="4C0A4D7BA3359071FD09FF5DB22EFF4B" authorityName="Dutrochet" authorityYear="1817" box="[672,778,197,222]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="genus">
|
||
<emphasis id="B97EEAEAA3359071FD09FF5DB22EFF4B" box="[672,778,197,222]" italics="true" pageId="11" pageNumber="40">Trocheta</emphasis>
|
||
</taxonomicName>
|
||
like with cornua which extends distinct on the previous somite. The genital atrium has long and straight or only horizontally slightly curved cornua. The apical part of the cornua is coiled (
|
||
<figureCitation id="13312A7DA3359071FE11FE90B332FEB7" box="[440,534,264,290]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="11" pageNumber="40">Fig. 8E</figureCitation>
|
||
). The cornua extends to annulus b5 of the previous somite. The broadened part of the vasa deferentia extends to the sixth ganglion after the female genital pore. Here is the start of the testicles. The ovarian sacks extends to the end of the third or slightly on the fourth somite after the female genital pore. They are only few winded in their total run and not curled in posterior parts. The ovisacs run only dorsally and do not cross the vasa deferentia (
|
||
<figureCitation id="13312A7DA3359071FD42FE17B26EFE3C" box="[747,842,399,425]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="11" pageNumber="40">Fig. 8D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106573A3359071FF3EFE4AB555FE78" box="[151,1137,466,493]" pageId="11" pageNumber="40" type="etymology">
|
||
<paragraph id="8BB536F8A3359071FF3EFE4AB555FE78" blockId="11.[151,1137,466,493]" box="[151,1137,466,493]" pageId="11" pageNumber="40">
|
||
<emphasis id="B97EEAEAA3359071FF3EFE4AB03AFE79" bold="true" box="[151,286,466,492]" pageId="11" pageNumber="40">Etymology</emphasis>
|
||
: The species is named after the mountains where the species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106573A3359071FF3EFD8EB08CFB9D" pageId="11" pageNumber="40" type="diagnosis">
|
||
<paragraph id="8BB536F8A3359071FF3EFD8EB433FD00" blockId="11.[151,1444,534,897]" pageId="11" pageNumber="40">
|
||
<emphasis id="B97EEAEAA3359071FF3EFD8EB08FFDA5" bold="true" box="[151,427,534,560]" pageId="11" pageNumber="40">Differential diagnosis</emphasis>
|
||
: The unicolour species
|
||
<taxonomicName id="4C0A4D7BA3359071FD76FD8EB2E3FDBA" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[735,967,534,559]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="sp. nov.">
|
||
<emphasis id="B97EEAEAA3359071FD76FD8EB2E3FDBA" box="[735,967,534,559]" italics="true" pageId="11" pageNumber="40">Trocheta taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FC7CFD8EB53FFDA5" box="[981,1051,534,560]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="4C0A4D7BA3359071FB78FD8EB44BFDBA" baseAuthorityName="Kosel" baseAuthorityYear="2004" box="[1233,1391,534,559]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="cylindrica">
|
||
<emphasis id="B97EEAEAA3359071FB78FD8EB44BFDBA" box="[1233,1391,534,559]" italics="true" pageId="11" pageNumber="40">T. cylindrica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D7BA3359071FA2DFD8EB08BFDC7" authority="Grosser, 2000" authorityName="Grosser" authorityYear="2000" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="haskonis">
|
||
<emphasis id="B97EEAEAA3359071FA2DFD8EB1DAFDC4" italics="true" pageId="11" pageNumber="40">T. haskonis</emphasis>
|
||
<bibRefCitation id="EF9B4B09A3359071FEA1FDA0B08BFDC7" author="Grosser, C." box="[264,431,568,594]" pageId="11" pageNumber="40" pagination="29 - 36" refId="ref4701" refString="Grosser, C. (2000) Beschreibung von Trocheta haskonis n. sp. (Hirudinea, Erpobdellidae) aus Sachsen- Anhalt. Lauterbornia, 38, 29 - 36." type="journal article" year="2000">Grosser, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D7BA3359071FE45FDA0B25BFDC7" authority="Stschegolew, 1938" authorityName="Stschegolew" authorityYear="1938" box="[492,895,568,594]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="danastrica">
|
||
<emphasis id="B97EEAEAA3359071FE45FDA0B3AFFDC4" box="[492,651,568,593]" italics="true" pageId="11" pageNumber="40">T. danastrica</emphasis>
|
||
Stschegolew, 1938
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C0A4D7BA3359071FC2FFDA0B53AFDC4" baseAuthorityName="Kosel" baseAuthorityYear="2004" box="[902,1054,568,593]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="cylindrica">
|
||
<emphasis id="B97EEAEAA3359071FC2FFDA0B53AFDC4" box="[902,1054,568,593]" italics="true" pageId="11" pageNumber="40">T. cylindrica</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0A4D7BA3359071FB68FDA0B475FDC4" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[1217,1361,568,593]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="n. sp.">
|
||
<emphasis id="B97EEAEAA3359071FB68FDA0B475FDC4" box="[1217,1361,568,593]" italics="true" pageId="11" pageNumber="40">T. taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FAF2FDA0B480FDC7" box="[1371,1444,568,594]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
by the characteristics of the reproductive system.
|
||
<taxonomicName id="4C0A4D7BA3359071FCABFDC2B2BAFDE6" baseAuthorityName="Kosel" baseAuthorityYear="2004" box="[770,926,602,627]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="cylindrica">
|
||
<emphasis id="B97EEAEAA3359071FCABFDC2B2BAFDE6" box="[770,926,602,627]" italics="true" pageId="11" pageNumber="40">T. cylindrica</emphasis>
|
||
</taxonomicName>
|
||
has a genital atrium with strong angular curved cornua; the ovarian sacks are one somite shorter and curled in posterior parts (
|
||
<figureCitation id="13312A7DA3359071FB2AFDE3B5F3FD00" box="[1155,1239,635,661]" captionStart="Figure 9" captionStartId="10.[151,227,1523,1547]" captionTargetBox="[301,1366,216,1472]" captionTargetPageId="10" captionText="Figure 9. Trocheta cylindrical, Bystrica, Slovakia: A shape and extension of the sexual organs, B – genital atrium." pageId="11" pageNumber="40">Figs. 9</figureCitation>
|
||
AB).
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3359071FF5FFD05B243FC89" blockId="11.[151,1444,534,897]" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C0A4D7BA3359071FF5FFD05B0F5FD23" authorityName="Grosser" authorityYear="2000" box="[246,465,669,694]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="haskonis">
|
||
<emphasis id="B97EEAEAA3359071FF5FFD05B0F5FD23" box="[246,465,669,694]" italics="true" pageId="11" pageNumber="40">Trocheta haskonis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0A4D7BA3359071FDDEFD05B22CFD23" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[631,776,669,694]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="n. sp.">
|
||
<emphasis id="B97EEAEAA3359071FDDEFD05B22CFD23" box="[631,776,669,694]" italics="true" pageId="11" pageNumber="40">T. taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FCBAFD05B271FD22" box="[787,853,669,695]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
in the shape of the papillae and position of the accessory genital pores.
|
||
<taxonomicName id="4C0A4D7BA3359071FE6BFD27B368FD4D" authorityName="Grosser" authorityYear="2000" box="[450,588,703,728]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="haskonis">
|
||
<emphasis id="B97EEAEAA3359071FE6BFD27B368FD4D" box="[450,588,703,728]" italics="true" pageId="11" pageNumber="40">T. haskonis</emphasis>
|
||
</taxonomicName>
|
||
has numerous small but well visible papillae roughen the dorsal and ventral surface, and accessory pores are located left and right of the male genital pore (
|
||
<figureCitation id="13312A7DA3359071FB62FD78B430FD6F" box="[1227,1300,736,762]" captionStart="Figure 2" captionStartId="4.[151,227,1713,1737]" captionTargetBox="[274,1310,210,1687]" captionTargetId="figure-8@4.[283,806,210,1002]" captionTargetPageId="4" captionText="Figure 2. AB Genital atrium: A – Dina stschegolewi, B – Dina pseudotrocheta. CD Colour of the anterior part: C – Dina stschegolewi, D – Dina pseudotrocheta (A, C: Crimean Peninsula; B: Germany, Saxony; D: Germany, Iterbach, holotype)" pageId="11" pageNumber="40">
|
||
Fig.
|
||
<quantity id="4CF29B1DA3359071FAACFD78B430FD6F" box="[1285,1300,736,762]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="40" unit="in" value="2.0">2</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="EF9B4B09A3359071FAE8FD78B1F3FC89" author="Grosser, C." pageId="11" pageNumber="40" pagination="111 - 113" refId="ref4733" refString="Grosser, C. (2013) First record of Trocheta haskonis Grosser, 2000 (Hirudinea: Erpobdellidae) in Serbia. Lauterbornia, 76, 111 - 113." type="journal article" year="2013">Grosser 2013</bibRefCitation>
|
||
). These accessory pores are absent in
|
||
<taxonomicName id="4C0A4D7BA3359071FD3DFC9AB206FC8E" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[660,802,770,795]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="n. sp.">
|
||
<emphasis id="B97EEAEAA3359071FD3DFC9AB206FC8E" box="[660,802,770,795]" italics="true" pageId="11" pageNumber="40">T. taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FC81FC9AB243FC89" box="[808,871,770,796]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3359071FF5FFCBCB4B0FC14" blockId="11.[151,1444,534,897]" pageId="11" pageNumber="40">
|
||
<taxonomicName id="4C0A4D7BA3359071FF5FFCBCB0C2FCA8" authorityName="Stschegolew" authorityYear="1938" box="[246,486,804,829]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="danastrica">
|
||
<emphasis id="B97EEAEAA3359071FF5FFCBCB0C2FCA8" box="[246,486,804,829]" italics="true" pageId="11" pageNumber="40">Trocheta danastrica</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0A4D7BA3359071FD2EFCBCB233FCA8" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[647,791,804,829]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="n. sp.">
|
||
<emphasis id="B97EEAEAA3359071FD2EFCBCB233FCA8" box="[647,791,804,829]" italics="true" pageId="11" pageNumber="40">T. taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FC89FCBCB245FCAB" box="[800,865,804,830]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
in the body form, and the shape of the papillae and the genital atrium. The body of
|
||
<taxonomicName id="4C0A4D7BA3359071FD97FCDEB3FFFCCA" authorityName="Stschegolew" authorityYear="1938" box="[574,731,838,863]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="danastrica">
|
||
<emphasis id="B97EEAEAA3359071FD97FCDEB3FFFCCA" box="[574,731,838,863]" italics="true" pageId="11" pageNumber="40">T. danastrica</emphasis>
|
||
</taxonomicName>
|
||
is longer and more slender, the genital atrium has very long and curved cornua, and the papillae are numerous and well visible (
|
||
<figureCitation id="13312A7DA3359071FC06FCFFB2DEFC14" box="[943,1018,871,897]" captionStart="Figure 1" captionStartId="3.[151,227,1508,1532]" captionTargetBox="[207,1403,183,1488]" captionTargetId="figure-15@3.[627,975,509,1488]" captionTargetPageId="3" captionText="Figure 1. Shape and extension of the sexual organs in the yellow spotted Erpobdellids: A – Dina stschegolewi (Crimean Peninsula); B – Dina apathyi (Germany, Altwarp); C – Dina orientalis (Lebanon, holotype)." pageId="11" pageNumber="40">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13312A7DA3359071FBA1FCFFB53DFC14" box="[1032,1049,871,897]" captionStart="Figure 6" captionStartId="7.[151,227,747,771]" captionTargetBox="[292,1233,178,722]" captionTargetPageId="7" captionText="Figure 6. Position of genital pores:A – Trocheta intermedia, paratype, B – Dina p. mauchi Germany, locus typicus; m = male, f = female." pageId="11" pageNumber="40">6</figureCitation>
|
||
,
|
||
<quantity id="4CF29B1DA3359071FB8CFCFFB576FC14" box="[1061,1106,871,897]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="11" pageNumber="40" unit="in" value="8.0">
|
||
<figureCitation id="13312A7DA3359071FB8CFCFFB510FC14" box="[1061,1076,871,897]" captionStart="Figure 8" captionStartId="9.[151,227,1962,1986]" captionTargetBox="[168,1392,188,1940]" captionTargetId="figure-7@9.[168,859,629,1332]" captionTargetPageId="9" captionText="Figure 8. Trocheta taunensis n. sp. (A – C holotype, D – E paratype), Wisper, Taunus Mt., Germany: A – habitus lateral view, B – habitus ventral view, C – annulation and position of genital pores, D shape and extension of the sexual organs, E – genital atrium." pageId="11" pageNumber="40">8</figureCitation>
|
||
in
|
||
</quantity>
|
||
<bibRefCitation id="EF9B4B09A3359071FBF0FCFFB4ACFC14" author="Grosser, C. & Epshtein, V. M." box="[1113,1416,871,897]" pageId="11" pageNumber="40" pagination="77 - 91" refId="ref4764" refString="Grosser, C. & Epshtein, V. M. (2009) Zum Artstatus des Egels Trocheta danastrica Stschegolew, 1938 (Annelida, Hirudinea: Erpobdellidae). Lauterbornia, 67, 77 - 91." type="journal article" year="2009">Grosser & Epshtein 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB536F8A3359071FF3EFC32B08CFB9D" blockId="11.[151,1444,938,1032]" pageId="11" pageNumber="40">
|
||
<emphasis id="B97EEAEAA3359071FF3EFC32B1D3FC51" bold="true" box="[151,247,938,964]" pageId="11" pageNumber="40">Habitat</emphasis>
|
||
: The Wisper is a small stream, shaded by riparian vegetation, with a width of ca. four to five metres. The fast running water is cold, shallow and stony. The most leeches were found under stones in the moist bank outside the water.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3106573A3359071FF3EFBA9B272FB1A" pageId="11" pageNumber="40" type="distribution">
|
||
<paragraph id="8BB536F8A3359071FF3EFBA9B272FB1A" blockId="11.[151,1444,1073,1167]" pageId="11" pageNumber="40">
|
||
<emphasis id="B97EEAEAA3359071FF3EFBA9B014FBDE" bold="true" box="[151,304,1073,1099]" pageId="11" pageNumber="40">Distribution</emphasis>
|
||
: The species is only known from the locus typicus. The examination of further finds of “
|
||
<taxonomicName id="4C0A4D7BA3359071FF0DFBCCB0AFFBF8" authorityName="sensu Dresscher & Engel" authorityYear="1955" box="[164,395,1108,1133]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="bykowskii">
|
||
<emphasis id="B97EEAEAA3359071FF0DFBCCB0AFFBF8" box="[164,395,1108,1133]" italics="true" pageId="11" pageNumber="40">Trocheta bykowskii</emphasis>
|
||
</taxonomicName>
|
||
” from Central Europe (e.g.
|
||
<collectingCountry id="F31D7668A3359071FD4BFBCBB270FBF8" box="[738,852,1107,1133]" name="Germany" pageId="11" pageNumber="40">Germany</collectingCountry>
|
||
: Eifel or
|
||
<collectingCountry id="F31D7668A3359071FC6CFBCBB504FBF8" box="[965,1056,1107,1133]" name="Austria" pageId="11" pageNumber="40">Austria</collectingCountry>
|
||
:
|
||
<collectingRegion id="49CEF81AA3359071FB87FBCBB5A7FBF8" box="[1070,1155,1107,1133]" country="Austria" name="Wien" pageId="11" pageNumber="40">Vienna</collectingRegion>
|
||
Woods; see
|
||
<bibRefCitation id="EF9B4B09A3359071FA89FBCBB1F0FB1A" author="Nesemann, H." pageId="11" pageNumber="40" pagination="37 - 60" refId="ref5031" refString="Nesemann, H. (1993) Bestimmungsschlussel fur mitteleuropaische Egel der Familie Erpobdelidae Blanchard 1894 (Hirudinea). Lauterbornia, 13, 37 - 60." type="journal article" year="1993">Nesemann 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9B4B09A3359071FF48FBEDB039FB1A" author="Nesemann, H." box="[225,285,1141,1167]" pageId="11" pageNumber="40" refId="ref5091" refString="Nesemann, H. (1997) Egel und Krebsegel (Clitellata: Hirudinea, Branchiobdellida) Osterreichs. Sonderheft der Ersten Vorarlberger Malakologischen Gesellschaft, Rankweil, Osterreich, 104 pp." type="book" year="1997">1997</bibRefCitation>
|
||
) will lead to a new records of
|
||
<taxonomicName id="4C0A4D7BA3359071FD2BFBEDB234FB1B" authority="- & Grosser, 2015" authorityName="- & Grosser" authorityYear="2015" box="[642,784,1141,1166]" class="Clitellata" family="Erpobdellidae" genus="Trocheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arhynchobdellida" pageId="11" pageNumber="40" phylum="Annelida" rank="species" species="taunensis" status="n. sp.">
|
||
<emphasis id="B97EEAEAA3359071FD2BFBEDB234FB1B" box="[642,784,1141,1166]" italics="true" pageId="11" pageNumber="40">T. taunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D5791A3359071FCBEFBEDB272FB1A" box="[791,854,1141,1167]" pageId="11" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |