treatments-xml/data/03/A3/87/03A387EAFF99FFF3FF7CFA65B98DFEF5.xml
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<document id="105FD3B0BF6B5FBAD954C7FF16834C1C" ID-DOI="10.5281/zenodo.5433967" ID-ISSN="0253-116X" ID-Zenodo-Dep="5433967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630432108880" checkinUser="valdenar" docAuthor="Abak &amp; D. W" docDate="2006" docId="03A387EAFF99FFF3FF7CFA65B98DFEF5" docLanguage="en" docName="LinzerbiolBeitr.2006.38.1.717-729.pdf" docOrigin="Linzer biologische Beiträge 38 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Dicheirotrichus (Trichocellus) latimanus Abak &amp; D.W 2006, spec. nova" docType="treatment" docVersion="4" lastPageNumber="723" masterDocId="FF9AFF92FF9DFFF4FFEBFF82BB1DFF8E" masterDocTitle="Three new species of the subgenus Trichocellus GANGLBAUER 1891 of the genus Dicheirotrichus JACQUELIN DU VAL 1857 from the East Palaearctic, with description of the male of D. stenothorax (KABAK &amp; KATAEV 1994) (Coleoptera, Carabidae)" masterLastPageNumber="729" masterPageNumber="717" pageNumber="721" updateTime="1699268493081" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BDE0B3D43E95E3BCB89CEC04B68BF328">Three new species of the subgenus Trichocellus GANGLBAUER 1891 of the genus Dicheirotrichus JACQUELIN DU VAL 1857 from the East Palaearctic, with description of the male of D. stenothorax (KABAK &amp; KATAEV 1994) (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="6E1BFA81925B77B2EED067EA14315F0D" type="personal">
<mods:role id="8AF9998F8EC79F8571EA783340E3C31A">
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<mods:namePart id="1CCBDCBF86F1DBB7B4BC9532135F72E0">Abak</mods:namePart>
</mods:name>
<mods:name id="202AC24F4B4594698A2A156B11242DD3" type="personal">
<mods:role id="3B442AFBB1DB95696E3A09269CB469D0">
<mods:roleTerm id="5CEC95E53FCFF4D2EB7B2AEDA758BA9A">Author</mods:roleTerm>
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<mods:namePart id="C0E1C1C92F9BACCD0C711F74D6033DFE">D. W</mods:namePart>
</mods:name>
<mods:typeOfResource id="066025D75B6F493CDFCC8FE41F8ADA48">text</mods:typeOfResource>
<mods:relatedItem id="33BDFBFE08FF8067A02C3D1DDDF30F37" type="host">
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<mods:title id="FE422FEA6533F72E74BB7F2CD656CC2A">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="34DC0FB7FEEBD7A1D9907B17BD346041">
<mods:date id="7882BB61BAAC9EA65E9B1035AD4956C3">2006</mods:date>
<mods:detail id="B600418FE69DD9D43A9AB82EB5B20AB7" type="pubDate">
<mods:number id="332F5A93545CC1ED41FD21EEBAE5BCC4">2006-07-21</mods:number>
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<mods:number id="353926B7C4249EBF923BB7D75C66F984">38</mods:number>
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<mods:start id="568932B403C8B0AD2536B6B80374BBB2">717</mods:start>
<mods:end id="E36517466A7B7B48CD3DE142181162B2">729</mods:end>
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<mods:classification id="D66E9BD10DE642AF05BE12B42308109B">journal article</mods:classification>
<mods:identifier id="01C712EA8D927F44D7029F5BBD94A526" type="DOI">10.5281/zenodo.5433967</mods:identifier>
<mods:identifier id="0CA4688DD46853B4EBD075CC3F89EF35" type="ISSN">0253-116X</mods:identifier>
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<treatment id="03A387EAFF99FFF3FF7CFA65B98DFEF5" ID-DOI="http://doi.org/10.5281/zenodo.5489002" ID-Zenodo-Dep="5489002" LSID="urn:lsid:plazi:treatment:03A387EAFF99FFF3FF7CFA65B98DFEF5" httpUri="http://treatment.plazi.org/id/03A387EAFF99FFF3FF7CFA65B98DFEF5" lastPageId="7" lastPageNumber="723" pageId="4" pageNumber="721">
<subSubSection id="C3106577FF99FFF1FF7CFA65B9B6FEA6" lastPageId="5" lastPageNumber="722" pageId="4" pageNumber="721" type="nomenclature">
<paragraph id="8BB536FCFF99FFF0FF7CFA65B822FA73" blockId="4.[151,831,1511,1533]" box="[151,831,1511,1533]" pageId="4" pageNumber="721">
<heading id="D0FD8190FF99FFF0FF7CFA65B822FA73" box="[151,831,1511,1533]" fontSize="8" level="2" pageId="4" pageNumber="721" reason="2">
<emphasis id="B97EEAEEFF99FFF0FF7CFA65B822FA73" bold="true" box="[151,831,1511,1533]" pageId="4" pageNumber="721">
<taxonomicName id="4C0A4D7FFF99FFF0FF7CFA65B959FA73" ID-CoL="35L86" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[151,580,1511,1533]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="721" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov." subGenus="Trichocellus">
<emphasis id="B97EEAEEFF99FFF0FF7CFA65BA26FA73" bold="true" box="[151,315,1511,1533]" italics="true" pageId="4" pageNumber="721">Dicheirotrichus</emphasis>
(
<emphasis id="B97EEAEEFF99FFF0FEA1FA65BAD0FA73" bold="true" box="[330,461,1511,1533]" italics="true" pageId="4" pageNumber="721">Trichocellus</emphasis>
)
<emphasis id="B97EEAEEFF99FFF0FE37FA65B959FA73" bold="true" box="[476,580,1511,1533]" italics="true" pageId="4" pageNumber="721">latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF99FFF0FDA1FA65B9A4FA73" box="[586,697,1511,1533]" pageId="4" pageNumber="721" rank="species">spec. nova</taxonomicNameLabel>
(
<figureCitation id="13312A79FF99FFF0FD2DFA65B825FA73" box="[710,824,1511,1533]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="4" pageNumber="721">Figs 11-18</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536FCFF99FFF1FF7CF98FB9B6FEA6" blockId="4.[151,1059,1549,1592]" lastBlockId="5.[151,1059,207,296]" lastPageId="5" lastPageNumber="722" pageId="4" pageNumber="721">
<materialsCitation id="3B623CA1FF99FFF0FF7CF98FB9CEF9B6" collectingDate="1998-07-21" collectingDateMax="1998-07-24" collectingDateMin="1998-07-21" collectionCode="ZISP" collectorName="A. Brinev" country="Russia" county="Tashtynskij Distr." location="Bolshoy On" municipality="Bolshoy On Valley" pageId="4" pageNumber="721" specimenCount="1" stateProvince="Khakasiya" typeStatus="holotype">
T y p e m a t e r i a l:
<typeStatus id="54B1885EFF99FFF0FE79F98FBAC3F9AF" box="[402,478,1549,1569]" pageId="4" pageNumber="721" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F31D766CFF99FFF0FDE7F98FB951F9AF" box="[524,588,1549,1569]" name="Russia" pageId="4" pageNumber="721">Russia</collectingCountry>
:
<collectingRegion id="49CEF81EFF99FFF0FDBEF98FB9A8F9AF" box="[597,693,1549,1569]" country="Russia" name="Khakasiya" pageId="4" pageNumber="721">Khakassia</collectingRegion>
,
<collectingCounty id="62D44E70FF99FFF0FD55F98FB879F9AF" box="[702,868,1549,1569]" pageId="4" pageNumber="721">Tashtynskij Distr.</collectingCounty>
,
<collectingMunicipality id="6BD1AC86FF99FFF0FC86F98FBF02F9AF" box="[877,1055,1549,1569]" pageId="4" pageNumber="721">Bolshoy On Valley</collectingMunicipality>
, env.
<location id="8ED56027FF99FFF0FF2EF9A6BA29F9B6" LSID="urn:lsid:plazi:treatment:03A387EAFF99FFF3FF7CFA65B98DFEF5:8ED56027FF99FFF0FF2EF9A6BA29F9B6" box="[197,308,1572,1592]" country="Russia" county="Tashtynskij Distr." municipality="Bolshoy On Valley" name="Bolshoy On" pageId="4" pageNumber="721" stateProvince="Khakasiya">Bolshoy On</location>
vill.,
<date id="FFB4103CFF99FFF0FE87F9A6BAE6F9B6" box="[364,507,1572,1592]" pageId="4" pageNumber="721" value="1998-07-21" valueMax="1998-07-24" valueMin="1998-07-21">
<collectingDate id="EFF0E9D4FF99FFF0FE87F9A6BAE6F9B6" box="[364,507,1572,1592]" pageId="4" pageNumber="721" value="1998-07-21" valueMax="1998-07-24" valueMin="1998-07-21">21-24.VII.1998</collectingDate>
</date>
,
<collectorName id="26FF532AFF99FFF0FDECF9A6B97FF9B6" box="[519,610,1572,1592]" pageId="4" pageNumber="721">A. Brinev</collectorName>
leg. (
<collectionCode id="ED1BAE39FF99FFF0FD72F9A6B9D0F9B6" box="[665,717,1572,1592]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="4" pageNumber="721">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B623CA1FF99FFF1FD36F9A6BA70FF6D" collectingDate="1998-07-21" collectingDateMax="1998-07-24" collectingDateMin="1998-07-21" collectionCode="MPU, ZISP" collectorName="A. Brinev" country="Russia" county="Tashtynskij Distr." lastPageId="5" lastPageNumber="722" location="Bolshoy On" municipality="Bolshoy On Valley" pageId="4" pageNumber="721" specimenCount="1" stateProvince="Khakasiya" typeStatus="paratype">
<typeStatus id="54B1885EFF99FFF0FD36F9A6B824F9B6" box="[733,825,1572,1592]" pageId="4" pageNumber="721" type="paratype">Paratypes</typeStatus>
: 2, 1, same data as holotype (
<collectionCode id="ED1BAE39FF98FFF1FF1EFF4DBA37FF6D" box="[245,298,207,227]" pageId="5" pageNumber="722">MPU</collectionCode>
,
<collectionCode id="ED1BAE39FF98FFF1FEDEFF4DBA75FF6D" box="[309,360,207,227]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="722">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B623CA1FF98FFF1FE92FF4DBAEBFF74" collectingDate="1979-07-13" collectorName="V. Eryshov" country="Russia" location="Gavrilovka" municipality="Tisul Distr." pageId="5" pageNumber="722" specimenCount="1" stateProvince="Kemerovo" typeStatus="paratype">
1,
<collectingRegion id="49CEF81EFF98FFF1FE4EFF4DB95CFF6D" box="[421,577,207,227]" country="Russia" name="Kemerovo" pageId="5" pageNumber="722">Kemerovo Prov.</collectingRegion>
,
<collectingMunicipality id="6BD1AC86FF98FFF1FDA0FF4DB9A5FF6D" box="[587,696,207,227]" pageId="5" pageNumber="722">Tisul Distr.</collectingMunicipality>
,
<location id="8ED56027FF98FFF1FD28FF4DB836FF6D" LSID="urn:lsid:plazi:treatment:03A387EAFF99FFF3FF7CFA65B98DFEF5:8ED56027FF98FFF1FD28FF4DB836FF6D" box="[707,811,207,227]" country="Russia" municipality="Tisul Distr." name="Gavrilovka" pageId="5" pageNumber="722" stateProvince="Kemerovo">Gavrilovka</location>
, shore of mountain brook,
<date id="FFB4103CFF98FFF1FF7CFF64BA14FF74" box="[151,265,230,250]" pageId="5" pageNumber="722" value="1979-07-13">
<collectingDate id="EFF0E9D4FF98FFF1FF7CFF64BA14FF74" box="[151,265,230,250]" pageId="5" pageNumber="722" value="1979-07-13">13.VII.1979</collectingDate>
</date>
,
<collectorName id="26FF532AFF98FFF1FEF9FF64BA66FF74" box="[274,379,230,250]" pageId="5" pageNumber="722">V. Eryshov</collectorName>
leg. (cSHIL)
</materialsCitation>
;
<materialsCitation id="3B623CA1FF98FFF1FDEAFF64BBC5FE9F" collectorName="Gornoaltaiskaya AO &amp; Snizek" country="Russia" location="Altai" pageId="5" pageNumber="722" specimenCount="1" stateProvince="Altay" typeStatus="paratype">
1,
<collectingRegion id="49CEF81EFF98FFF1FDC0FF64B946FF74" box="[555,603,230,250]" country="Russia" name="Altay" pageId="5" pageNumber="722">Altai</collectingRegion>
,
<collectorName id="26FF532AFF98FFF1FD8FFF64B83FFF74" box="[612,802,230,250]" pageId="5" pageNumber="722">Gornoaltaiskaya AO</collectorName>
, near Turochak,
<collectorName id="26FF532AFF98FFF1FC56FF64B8E6FF74" box="[957,1019,230,250]" pageId="5" pageNumber="722">Snižek</collectorName>
leg. (cWR)
</materialsCitation>
;
<materialsCitation id="3B623CA1FF98FFF1FF0FFF7FB9BBFEA6" collectingDate="2000-08-17" collectionCode="ISEAN" collectorName="A. Vorontsov" country="Russia" elevation="1700" location="Shumak River" municipality="Tunkinskie Goltsi" pageId="5" pageNumber="722" specimenCount="1" stateProvince="Buryatiya" typeStatus="paratype">
2,
<collectingRegion id="49CEF81EFF98FFF1FEC8FF7FBA69FE9F" box="[291,372,253,273]" country="Russia" name="Buryatiya" pageId="5" pageNumber="722">Buryatia</collectingRegion>
,
<collectingMunicipality id="6BD1AC86FF98FFF1FE94FF7FB93AFE9F" box="[383,551,253,273]" pageId="5" pageNumber="722">Tunkinskie Goltsi</collectingMunicipality>
,
<locationDeviation id="9A072FDEFF98FFF1FDD8FF7FB81EFE9F" box="[563,771,253,273]" location="Turan" pageId="5" pageNumber="722">
<quantity id="4CF29B19FF98FFF1FDD8FF7FB972FE9F" box="[563,623,253,273]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="722" unit="km" value="30.0">30 km</quantity>
NNE of Turan
</locationDeviation>
, upp. of
<location id="8ED56027FF98FFF1FCB1FF7FB8FFFE9F" LSID="urn:lsid:plazi:treatment:03A387EAFF99FFF3FF7CFA65B98DFEF5:8ED56027FF98FFF1FCB1FF7FB8FFFE9F" box="[858,994,253,273]" country="Russia" municipality="Tunkinskie Goltsi" name="Shumak River" pageId="5" pageNumber="722" stateProvince="Buryatiya">Shumak River</location>
,
<quantity id="4CF29B19FF98FFF1FC06FF7FBBC3FEA6" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="5" pageNumber="722" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<elevation id="0027D1CFFF98FFF1FC06FF7FBBC3FEA6" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="5" pageNumber="722" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">1600- 1800 m</elevation>
</quantity>
, forest,
<date id="FFB4103CFF98FFF1FECCFE96BA82FEA6" box="[295,415,276,296]" pageId="5" pageNumber="722" value="2000-08-17">
<collectingDate id="EFF0E9D4FF98FFF1FECCFE96BA82FEA6" box="[295,415,276,296]" pageId="5" pageNumber="722" value="2000-08-17">17.VIII.2000</collectingDate>
</date>
,
<collectorName id="26FF532AFF98FFF1FE42FE96B938FEA6" box="[425,549,276,296]" pageId="5" pageNumber="722">A. Vorontsov</collectorName>
leg. (
<collectionCode id="ED1BAE39FF98FFF1FDB3FE96B9BDFEA6" box="[600,672,276,296]" pageId="5" pageNumber="722">ISEAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106577FF98FFF3FF7CFEB5B98DFEF5" lastPageId="7" lastPageNumber="724" pageId="5" pageNumber="722" type="description">
<paragraph id="8BB536FCFF98FFF1FF7CFEB5B828FEC2" blockId="5.[151,821,310,332]" box="[151,821,310,332]" pageId="5" pageNumber="722">
D e s c r i p t i o n: Body length
<quantity id="4CF29B19FF98FFF1FE1CFEB4B973FEC2" box="[503,622,310,332]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="5" pageNumber="722" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">4.6-5.1 mm</quantity>
, width
<quantity id="4CF29B19FF98FFF1FD52FEB4B82CFEC2" box="[697,817,310,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="5" pageNumber="722" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8-2.1 mm</quantity>
.
</paragraph>
<paragraph id="8BB536FCFF98FFF1FF7CFED9B981FDB0" blockId="5.[151,1059,347,574]" pageId="5" pageNumber="722">
Colour: Dorsum largely reddish brown, shiny. Head usually strongly infuscate, almost throughout black, or with bases of mandibles, apical margin of clypeus and lateral areas before and behind eyes pale. Pronotum with large central blurred transverse dark macula sometimes reaching lateral margins. Elytra rather strongly infuscate in apical two-thirds, with pale sutural intervals. Epipleurae of pronotum pale. Epipleurae of elytra infuscate anteriorly. Underside black. Appendages reddish brown but apical palpomeres, antennae from 4
<superScript id="7C7F9BB4FF98FFF1FF35FD87BBF7FD9D" attach="left" box="[222,234,517,531]" fontSize="6" pageId="5" pageNumber="722">th</superScript>
(sometimes from 3
<superScript id="7C7F9BB4FF98FFF1FE53FD87BAD8FD9D" attach="left" box="[440,453,517,531]" fontSize="6" pageId="5" pageNumber="722">rd</superScript>
or even 2
<superScript id="7C7F9BB4FF98FFF1FDDFFD87B959FD9D" attach="none" box="[564,580,517,531]" fontSize="6" pageId="5" pageNumber="722">nd</superScript>
) antennomere, and metatarsi (sometimes also mesotarsi) distinctly infuscate. Pubescence yellow.
</paragraph>
<paragraph id="8BB536FCFF98FFF1FF7CFDCFB920FC28" blockId="5.[151,1059,589,934]" pageId="5" pageNumber="722">Head: Relatively large (WHmax/WP = 0.77-0.81; WHmin/WP = 0.59-0.62), with very coarse and dense puncturation areas under and particularly behind eyes, also on vertex, and with more fine puncturation around frontal foveae, sometimes also on frons and along apical margin of clypeus, but usually clypeus, central part of frons and anterior part of vertex unpunctured. Each puncture bearing a short seta. Eyes large, moderately or strongly convex, separated from buccal fissure by distance much lesser than width of first antennomere. Tempora rather short, convex. Clypeus depressed along apical margin. Labrum with nearly straight apical margin. Dorsal microsculpture in male usually not visible, in female (and rarely in male) obliterate isodiametric and weakly transverse meshes recognizable on clypeus and on areas behind eyes. Mandibles acute at apices. Antennae rather long, extending approximately to a quarter of elytra, with middle antennomeres about two times as long as wide.</paragraph>
<paragraph id="8BB536FCFF98FFF1FF7CFC34B852FAEB" blockId="5.[151,1059,949,1381]" pageId="5" pageNumber="722">
Pronotum (
<figureCitation id="13312A79FF98FFF1FEE0FC37BA45FC45" box="[267,344,949,971]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="5" pageNumber="722">Fig. 11</figureCitation>
): Weakly convex, slightly flattened basally, 1.34-1.41 times as wide as long, widest in apical third, with sides basally more or less rectilinearly converging posteriad. Apical margin weakly arcuately emarginate, bordered only laterally. Basal margin nearly straight medially, clearly oblique laterally, with more or less distinct bead along oblique portion on each side, approximately equal to apical margin, notably narrower than elytral base, and ciliate on basal ridge. Apical angles not protruding anteriad, rounded at apices. Basal angles distinct, obtuse, usually with sharp apices, each bearing a moderately long seta. Lateral furrow narrow throughout and lateral depressions not developed. Basal foveae large and narrow, usually reaching basal pronotal margin and curved slightly to sides apically, separated from each other and in most cases also from sides by convexities; sometimes areas near basal angles slightly flattened. Pronotal surface rather coarsely and densely punctured along margins (punctures in basal foveae confluent) and more finely and sparsely on disc; central part of pronotum often almost unpunctured. Each puncture bearing a short seta. Dorsal microsculpture usually invisible, sometimes obliterate meshes recognizable at apical and basal angles.
</paragraph>
<paragraph id="8BB536FCFF98FFF2FF7CFAF6BA4EFE5D" blockId="5.[151,1060,1396,1594]" lastBlockId="6.[151,1059,211,467]" lastPageId="6" lastPageNumber="723" pageId="5" pageNumber="722">
Elytra: Moderately convex, elongated, 1.50-1.60 times as long as wide, 3.00-3.20 times as long and 1.40-1.49 times as wide as pronotum, weakly widened posteriad and broadly rounded at sides; widest behind middle. Humeri slightly prominent, evenly and rather broadly rounded at apices. Subapical sinuation weak. Sutural angle narrowly rounded at apex. Basal bead slightly sinuate, arcuately curving inside humerus up to lateral margin. Parascutellar setigerous pore puncture present, large. Third interval in apical third or just before with a discal setigerous pore near second stria. Posterior group of umbilicate series consisting of seven or eight setigerous pores. Striae unpunctured, slightly crenulate, weakly impressed on disc, superficial along elytral sides, particularly basally; 7
<superScript id="7C7F9BB4FF9BFFF2FBFDFF68BF3FFF76" attach="none" box="[1046,1058,234,248]" fontSize="6" pageId="6" pageNumber="723">th</superScript>
stria obliterate at basal bead of elytra. Intervals rather flat, at most weakly convex in basal half, weakly narrowed posteriad. Puncturation of intervals on elytral disc somewhat coarse, with usually two (rarely one or three) punctures in a transverse row across each interval at middle. Each puncture bearing a short, posteriorly inclined seta. Microsculpture consisting of isodiametric meshes, in male clearly visible only in apical quarter, along basal bead and on 1-2 lateral intervals; in female evanescent meshes hardly recognized also on disc.
</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFE61BA8FFE77" blockId="6.[151,402,483,505]" box="[151,402,483,505]" pageId="6" pageNumber="723">Wings: Fully developed.</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFD8ABA28FDD6" blockId="6.[151,1059,520,784]" pageId="6" pageNumber="723">
Ventral surface: Metepisterna (
<figureCitation id="13312A79FF9BFFF2FE37FD8AB934FD90" box="[476,553,520,542]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Fig. 14</figureCitation>
) markedly longer than wide, strongly narrowed posteriad. Anal sternum rounded at apex, with one pair of marginal setae in male and two pairs in female.
</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFDEAB808FC9E" blockId="6.[151,1059,520,784]" pageId="6" pageNumber="723">
Legs: 5
<superScript id="7C7F9BB4FF9BFFF2FF0CFDE0BBEEFDFE" attach="left" box="[231,243,610,624]" fontSize="6" pageId="6" pageNumber="723">th</superScript>
tarsomere with three pairs of ventro-lateral setae. Metatarsus comparatively stout, a little shorter than width of head across eyes, with almost triangular tarsomeres; 1
<superScript id="7C7F9BB4FF9BFFF2FBF3FDFDBF3EFD03" attach="none" box="[1048,1059,639,653]" fontSize="6" pageId="6" pageNumber="723">st</superScript>
metatarsomere much longer than second and slightly shorter than second and third together. In male protarsi (
<figureCitation id="13312A79FF9BFFF2FE5FFD42B91CFD58" box="[436,513,704,726]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Fig. 12</figureCitation>
) rather strongly dilated, with spongy vestiture ventrally; mesotarsi (
<figureCitation id="13312A79FF9BFFF2FEBAFD5FBA82FD7D" box="[337,415,733,755]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Fig. 17</figureCitation>
) distinctly, even if rather weakly, dilated and with biseriate vestiture ventrally (for comparison female mesotarsi -
<figureCitation id="13312A79FF9BFFF2FD55FD78B814FC9E" box="[702,777,762,784]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFC9DBBD3FC03" blockId="6.[151,1059,799,909]" pageId="6" pageNumber="723">
Female genitalia (
<figureCitation id="13312A79FF9BFFF2FEBDFC9DBABBFCBB" box="[342,422,799,821]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Fig. 13</figureCitation>
): Hemisternite asymmetrical, without spines or setae. Basal stylomere rather strongly widened apicad, also without spines or setae. Apical stylomere weakly arcuate, somewhat long, with a peg-like seta on outer ventral margin in basal third.
</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFC1FBA2CFB84" blockId="6.[151,1059,925,1034]" pageId="6" pageNumber="723">
Median lobe of aedeagus (
<figureCitation id="13312A79FF9BFFF2FE4FFC1FB90AFC3D" box="[420,535,925,947]" captionStart="Figs 11-18" captionStartId="11.[151,191,1395,1415]" captionTargetBox="[163,1036,217,1366]" captionTargetId="figure-107@11.[151,1058,204,1383]" captionTargetPageId="11" captionText="Figs 11-18: Dicheirotrichus (Trichocellus) latimanus spec. nova. (11) Pronotum. (12) Male protarsus. (13) Hemisternite and stylus. (14) Left metepisternum. (15-16) Median lobe of aedeagus (holotype), (15) dorsal view, (16) lateral view. (17) Male mesotarsus. (18) Female mesotarsus. Figs 19-20: D. (T.) angularis (REITTER) (Amur Prov.). Median lobe of aedeagus, (19) dorsal view, (20) lateral view. Scales = 0.5 mm (A: Fig. 11; B: Figs 12, 14, 17, 18; C: Figs 13, 15-16, 19-20)." figureDoi="http://doi.org/10.5281/zenodo.5433971" httpUri="https://zenodo.org/record/5433971/files/figure.png" pageId="6" pageNumber="723">Figs 15-16</figureCitation>
): Arcuate, with apex distinctly curved dorsad (lateral aspect), markedly dilated apically, its sides rather strongly sinuate just behind middle (dorsal aspect). Internal sac with asymmetrical pattern of tiny spines, without larger spines or teeth.
</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFB98BBE2FBC3" blockId="6.[151,1058,1050,1101]" pageId="6" pageNumber="723">D i s t r i b u t i o n: This new species seems to be confined to the Altai-Sayan Mountain Land.</paragraph>
<paragraph id="8BB536FCFF9BFFF2FF7CFBDFB95DFB23" blockId="6.[151,1059,1116,1197]" pageId="6" pageNumber="723">
E t y m o l o g y: The species epithet is based on two Latin words,
<emphasis id="B97EEAEEFF9BFFF2FC66FBDEB8A3FBFC" box="[909,958,1116,1138]" italics="true" pageId="6" pageNumber="723">latus</emphasis>
meaning wide and
<emphasis id="B97EEAEEFF9BFFF2FEE7FBF8BA53FB01" box="[268,334,1146,1167]" italics="true" pageId="6" pageNumber="723">manus</emphasis>
meaning hand, referring to the dilated mesotarsi and strongly dilated protarsi, characteristic of the new species.
</paragraph>
<paragraph id="8BB536FCFF9BFFF3FF7CFB3FB98DFEF5" blockId="6.[151,1059,1212,1585]" lastBlockId="7.[151,1059,211,379]" lastPageId="7" lastPageNumber="724" pageId="6" pageNumber="723">
R e m a r k s:
<taxonomicName id="4C0A4D7FFF9BFFF2FED3FB3FBAA1FB5C" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[312,444,1212,1234]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov.">
<emphasis id="B97EEAEEFF9BFFF2FED3FB3FBAA1FB5C" box="[312,444,1212,1234]" italics="true" pageId="6" pageNumber="723">D. latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF9BFFF2FE28FB3FB933FB5C" box="[451,558,1213,1234]" pageId="6" pageNumber="723" rank="species">spec. nova</taxonomicNameLabel>
is easily distinguished from all the other species of the genus by the mesotarsomeres 1-4 of males dilated and with adhesive vestiture ventrally. Another prominent distinctive feature of this new species is the median lobe of the aedeagus notably dilated apically. In all other species of
<taxonomicName id="4C0A4D7FFF9BFFF2FCFCFA96B8AAFAA4" authorityName="JACQUELIN DU VAL" authorityYear="1857" box="[791,951,1300,1322]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAEEFF9BFFF2FCFCFA96B8AAFAA4" box="[791,951,1300,1322]" italics="true" pageId="6" pageNumber="723">Dicheirotrichus</emphasis>
</taxonomicName>
known to us, the male mesotarsi are simple (not dilated) and the median lobe of the aedeagus is much narrower apically. At the same time,
<taxonomicName id="4C0A4D7FFF9BFFF2FD8EFACDB9F6FAEA" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[613,747,1358,1380]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov.">
<emphasis id="B97EEAEEFF9BFFF2FD8EFACDB9F6FAEA" box="[613,747,1358,1380]" italics="true" pageId="6" pageNumber="723">D. latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF9BFFF2FD1EFACDB87EFAEA" box="[757,867,1359,1380]" pageId="6" pageNumber="723" rank="species">spec. nova</taxonomicNameLabel>
is very similar in other character states, including the shape of pronotum with distinct basal angles and the details of the internal sac of the median lobe, to
<taxonomicName id="4C0A4D7FFF9BFFF2FD67FA08B8ABFA11" authority="(REITTER 1899)" baseAuthorityName="REITTER" baseAuthorityYear="1899" box="[652,950,1417,1439]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9BFFF2FD67FA08B813FA11" box="[652,782,1417,1439]" italics="true" pageId="6" pageNumber="723">D. angularis</emphasis>
(REITTER 1899)
</taxonomicName>
from East Siberia; females of both species are difficult to separate. In addition to the two distinctive characters listed above,
<taxonomicName id="4C0A4D7FFF9BFFF2FE66FA46B90DFA54" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[397,528,1476,1498]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov.">
<emphasis id="B97EEAEEFF9BFFF2FE66FA46B90DFA54" box="[397,528,1476,1498]" italics="true" pageId="6" pageNumber="723">D. latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF9BFFF2FDFCFA46B99EFA57" box="[535,643,1476,1497]" pageId="6" pageNumber="723" rank="species">spec. nova</taxonomicNameLabel>
may be distinguished from
<taxonomicName id="4C0A4D7FFF9BFFF2FC49FA46BF3EFA54" baseAuthorityName="REITTER" baseAuthorityYear="1899" box="[930,1059,1476,1498]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9BFFF2FC49FA46BF3EFA54" box="[930,1059,1476,1498]" italics="true" pageId="6" pageNumber="723">D. angularis</emphasis>
</taxonomicName>
by having the male protarsi more strongly dilated, the apical portion of the median lobe of the aedeagus clearly curved dorsad (the median lobe of
<taxonomicName id="4C0A4D7FFF9BFFF2FD16FA7DB862F99A" baseAuthorityName="REITTER" baseAuthorityYear="1899" box="[765,895,1534,1556]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="723" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9BFFF2FD16FA7DB862F99A" box="[765,895,1534,1556]" italics="true" pageId="6" pageNumber="723">D. angularis</emphasis>
</taxonomicName>
is illustrated in Figs 19-20), the pronotum relatively wider, the elytra slightly longer, and the elytral microsculpture more distinct and more widely distributed (in both sexes of
<taxonomicName id="4C0A4D7FFF9AFFF3FC70FF56BF01FF67" baseAuthorityName="REITTER" baseAuthorityYear="1899" box="[923,1052,211,233]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9AFFF3FC70FF56BF01FF67" box="[923,1052,211,233]" italics="true" pageId="7" pageNumber="724">D. angularis</emphasis>
</taxonomicName>
, meshes recognizable only apically). In our opinion,
<taxonomicName id="4C0A4D7FFF9AFFF3FD21FF73B84EFE88" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[714,851,240,262]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov.">
<emphasis id="B97EEAEEFF9AFFF3FD21FF73B84EFE88" box="[714,851,240,262]" italics="true" pageId="7" pageNumber="724">D. latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF9AFFF3FCB5FF73B8D3FE88" box="[862,974,241,262]" pageId="7" pageNumber="724" rank="species">spec. nova</taxonomicNameLabel>
and
<taxonomicName id="4C0A4D7FFF9AFFF3FBE1FF73BBE5FEAA" baseAuthorityName="REITTER" baseAuthorityYear="1899" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9AFFF3FBE1FF73BBE5FEAA" italics="true" pageId="7" pageNumber="724">D. angularis</emphasis>
</taxonomicName>
seems to be two closely related and probably vicariant species. It is most likely also that the dilated male mesotarsi and the very wide apical portion of the apical lobe characteristic of
<taxonomicName id="4C0A4D7FFF9AFFF3FEAAFECBBAD8FED0" authority="Abak &amp; D.W, 2006" authorityName="Abak &amp; D.W" authorityYear="2006" box="[321,453,328,350]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="latimanus" status="sp. nov.">
<emphasis id="B97EEAEEFF9AFFF3FEAAFECBBAD8FED0" box="[321,453,328,350]" italics="true" pageId="7" pageNumber="724">D. latimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D5795FF9AFFF3FE27FECBB92AFED0" box="[460,567,329,350]" pageId="7" pageNumber="724" rank="species">spec. nova</taxonomicNameLabel>
should be treated as autapomorphies evolved in this species after its separation from
<taxonomicName id="4C0A4D7FFF9AFFF3FDE1FEE4B997FEF5" baseAuthorityName="REITTER" baseAuthorityYear="1899" box="[522,650,357,379]" class="Insecta" family="Carabidae" genus="Dicheirotrichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="724" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B97EEAEEFF9AFFF3FDE1FEE4B997FEF5" box="[522,650,357,379]" italics="true" pageId="7" pageNumber="724">D. angularis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>