313 lines
36 KiB
XML
313 lines
36 KiB
XML
<document id="9B40D8E9EB202505B24EDF7115899B23" ID-DOI="10.11646/zootaxa.5406.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10611440" ID-ZooBank="897EDC9B-860D-4A9C-BBB0-43B4B359AC99" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="felipe" checkinTime="1706895390540" checkinUser="plazi" docAuthor="Mendes, Gabriela Caroline, Nascimento, Jeane Marcelle Cavalcante Do, Fusari, Lívia Maria & Hamada, Neusa" docDate="2024" docId="03A387D3FFC1FFF71C87DC67E966AEE2" docLanguage="en" docName="zootaxa.5406.1.6.pdf" docOrigin="Zootaxa 5406 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ilyobius brasiliensis" docType="treatment" docVersion="3" lastPageNumber="139" masterDocId="FF9AFFABFFCEFFE71C10DD25EA21AA27" masterDocTitle="Updates on the taxonomy of Ilyobius nubilus (Navás, 1933) (Megaloptera: Sialidae: Sialinae) and Ilyobius brasiliensis (Navás, 1936) from southeastern Brazil" masterLastPageNumber="140" masterPageNumber="123" pageNumber="138" updateTime="1707225908592" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="C6B1AF712F54465539FEA2AE61C469C3">Updates on the taxonomy of Ilyobius nubilus (Navás, 1933) (Megaloptera: Sialidae: Sialinae) and Ilyobius brasiliensis (Navás, 1936) from southeastern Brazil</mods:title>
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<mods:namePart id="E473BCCA067CD55E96D1012E468100E0">Mendes, Gabriela Caroline</mods:namePart>
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<mods:namePart id="A36D80263FC59E6CFB67B97365B266B1">Nascimento, Jeane Marcelle Cavalcante Do</mods:namePart>
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<mods:affiliation id="C157D434128C4E1E802969776AB67037">Universidade Federal do Pará, Programa de Pós-Graduação em Zoologia, Belém, PA, Brazil</mods:affiliation>
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<mods:namePart id="6056767A4FF17533A4E45775E83A4947">Fusari, Lívia Maria</mods:namePart>
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<mods:affiliation id="C3A56E1676484FFD739F47686B9B785A">Departamento de Hidrobiologia, Universidade Federal de São Carlos (UFSCar), São Carlos, SP, Brazil</mods:affiliation>
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<mods:namePart id="EE70B73EA5463C7C085D33176E4A9BF7">Hamada, Neusa</mods:namePart>
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<mods:title id="9FA05B4B3D0ED5C0F260E6C6AF77D910">Zootaxa</mods:title>
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<mods:date id="2DB3B6DDB69C26EB9BC560289315A1E9">2024</mods:date>
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<treatment id="03A387D3FFC1FFF71C87DC67E966AEE2" ID-DOI="http://doi.org/10.5281/zenodo.10611437" ID-Zenodo-Dep="10611437" LSID="urn:lsid:plazi:treatment:03A387D3FFC1FFF71C87DC67E966AEE2" httpUri="http://treatment.plazi.org/id/03A387D3FFC1FFF71C87DC67E966AEE2" lastPageId="16" lastPageNumber="139" pageId="15" pageNumber="138">
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<subSubSection id="C310654EFFC1FFE81C87DC67E880AB7A" box="[151,673,322,349]" pageId="15" pageNumber="138" type="nomenclature">
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<paragraph id="8BB536C5FFC1FFE81C87DC67E880AB7A" blockId="15.[151,673,322,385]" box="[151,673,322,349]" pageId="15" pageNumber="138">
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<heading id="D0FD81A9FFC1FFE81C87DC67E880AB7A" bold="true" box="[151,673,322,349]" fontSize="11" level="1" pageId="15" pageNumber="138" reason="1">
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<emphasis id="B97EEAD7FFC1FFE81C87DC67E880AB7A" bold="true" box="[151,673,322,349]" pageId="15" pageNumber="138">
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<taxonomicName id="4C0A4D46FFC1FFE81C87DC67E81AAB7B" ID-CoL="4N73V" authority="(Navas, 1936)" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[151,571,322,349]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B97EEAD7FFC1FFE81C87DC67EBA8AB7A" bold="true" box="[151,393,322,349]" italics="true" pageId="15" pageNumber="138">Ilyobius brasiliensis</emphasis>
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(Navás, 1936)
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</taxonomicName>
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, sp. rev.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C310654EFFC1FFE81C87DC42EB0FABA6" box="[151,302,359,385]" pageId="15" pageNumber="138" type="description">
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<paragraph id="8BB536C5FFC1FFE81C87DC42EB0FABA6" blockId="15.[151,673,322,385]" box="[151,302,359,385]" pageId="15" pageNumber="138">
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(
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<figureCitation id="13312A40FFC1FFE81C8FDC42EB07ABA6" box="[159,294,359,385]" captionStart="FIGURE 12" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[296,1291,533,1946]" captionTargetId="figure-118@15.[296,1291,533,1946]" captionTargetPageId="15" captionText="FIGURE 12. (A–D) Ilyobius brasiliensis, holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E) Ilyobius nubilus, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="15" pageNumber="138">Fig. 12A–C</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C310654EFFC1FFE81C87DC82E98AABDA" pageId="15" pageNumber="138" type="reference_group">
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<paragraph id="8BB536C5FFC1FFE81C87DC82EB49ABF9" blockId="15.[151,1436,423,509]" pageId="15" pageNumber="138">
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<treatmentCitationGroup id="AB1A11EBFFC1FFE81C87DC82EB49ABF9" pageId="15" pageNumber="138">
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<taxonomicName id="4C0A4D46FFC1FFE81C87DC82E80DAB98" ID-CoL="4N73V" authority="Navas, 1936: 36" authorityName="Navas" authorityPageNumber="36" authorityYear="1936" box="[151,556,423,447]" class="Insecta" family="Sialidae" genus="Protosialis" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="brasiliensis">
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<emphasis id="B97EEAD7FFC1FFE81C87DC82EBA0AB98" box="[151,385,423,447]" italics="true" pageId="15" pageNumber="138">Protosialis brasiliensis</emphasis>
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Navás, 1936: 36
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</taxonomicName>
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||
(original description);
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<treatmentCitation id="0AAB10D4FFC1FFE81F06DC82EE15AB98" author="Contreras-Ramos, A." box="[790,1076,423,447]" page="980" pageId="15" pageNumber="138" year="2006">
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<bibRefCitation id="EF9B4B34FFC1FFE81F06DC82EE15AB98" author="Contreras-Ramos, A." box="[790,1076,423,447]" pageId="15" pageNumber="138" pagination="977" refId="ref7201" refString="Contreras-Ramos, A. (2006) Protosialis ranchograndis, a new species of alderfly from Venezuela, with a redescription of P. brasiliensis Navas (Megaloptera: Sialidae). Proceeding-Entomological Society of Washington, 108, 977." type="journal article" year="2006">Contreras-Ramos 2006: 980</bibRefCitation>
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||
</treatmentCitation>
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(redescription).
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||
<treatmentCitation id="0AAB10D4FFC1FFE818CCDC82EFBDAB98" author="Heckman, C. W." box="[1244,1436,423,447]" page="78" pageId="15" pageNumber="138" year="2017">
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<bibRefCitation id="EF9B4B34FFC1FFE818CCDC82EFBDAB98" author="Heckman, C. W." box="[1244,1436,423,447]" pageId="15" pageNumber="138" pagination="75 - 79" refId="ref7436" refString="Heckman, C. W. (2017) Sialidae. In: Neuroptera (Including Megaloptera). Encyclopedia of South American Aquatic Insects. Springer, Cham, pp. 75 - 79. https: // doi. org / 10.1007 / 978 - 3 - 319 - 35125 - 4 _ 3" type="book chapter" year="2017">Heckman 2017: 78</bibRefCitation>
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</treatmentCitation>
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||
(key of species)
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BB536C5FFC1FFE81C87DCC1E98AABDA" blockId="15.[151,1436,423,509]" box="[151,939,484,509]" pageId="15" pageNumber="138">
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<treatmentCitationGroup id="AB1A11EBFFC1FFE81C87DCC1E819ABDA" box="[151,568,484,509]" pageId="15" pageNumber="138">
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<taxonomicName id="4C0A4D46FFC1FFE81C87DCC1EB45ABDB" ID-CoL="4N73V" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[151,356,484,508]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="brasiliensis">
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||
<emphasis id="B97EEAD7FFC1FFE81C87DCC1EB45ABDB" box="[151,356,484,508]" italics="true" pageId="15" pageNumber="138">Ilyobius brasiliensis</emphasis>
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</taxonomicName>
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||
;
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||
<treatmentCitation id="0AAB10D4FFC1FFE81D61DCC1E819ABDA" author="Liu, X. & Hayashi, F. & Yang, D." box="[369,568,484,509]" page="31" pageId="15" pageNumber="138" year="2015">
|
||
<bibRefCitation id="EF9B4B34FFC1FFE81D61DCC1E819ABDA" author="Liu, X. & Hayashi, F. & Yang, D." box="[369,568,484,509]" pageId="15" pageNumber="138" pagination="18 - 49" refId="ref7542" refString="Liu, X., Hayashi, F. & Yang, D. (2015 a) Phylogeny of the family Sialidae (Insecta: Megaloptera) inferred from morphological data, with implications for generic classification and historical biogeography. Cladistics, 31, 18 - 49. https: // doi. org / 10.1111 / cla. 12071" type="journal article" year="2015">
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||
Liu
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||
<emphasis id="B97EEAD7FFC1FFE81D8BDCC0EBE6ABDB" box="[411,455,484,508]" italics="true" pageId="15" pageNumber="138">et al</emphasis>
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||
. 2015a: 31
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</bibRefCitation>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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||
(phylogeny of
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<taxonomicName id="4C0A4D46FFC1FFE81EC4DCC1E908ABDB" ID-CoL="G9L" authorityName="Leach" authorityYear="1815" box="[724,809,484,508]" class="Insecta" family="Sialidae" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="family">Sialidae</taxonomicName>
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; comb. nov.)
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF75664DFFC1FFE81C87DA97E9ECADC9" ID-DOI="http://doi.org/10.5281/zenodo.10611464" ID-Zenodo-Dep="10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="15" pageNumber="138" startId="15.[151,250,1970,1994]" targetBox="[296,1291,533,1946]" targetPageId="15" targetType="figure">
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<paragraph id="8BB536C5FFC1FFE81C87DA97E9ECADC9" blockId="15.[151,1437,1970,2030]" pageId="15" pageNumber="138">
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||
<emphasis id="B97EEAD7FFC1FFE81C87DA97EB3EADED" bold="true" box="[151,287,1970,1994]" pageId="15" pageNumber="138">FIGURE 12.</emphasis>
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||
(A–D)
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||
<emphasis id="B97EEAD7FFC1FFE81D7DDA97E861ADED" box="[365,576,1970,1994]" italics="true" pageId="15" pageNumber="138">
|
||
<taxonomicName id="4C0A4D46FFC1FFE81D7DDA97E81AADED" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[365,571,1970,1994]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="brasiliensis">Ilyobius brasiliensis</taxonomicName>
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||
,
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||
</emphasis>
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||
holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E)
|
||
<taxonomicName id="4C0A4D46FFC1FFE81CAEDAF3EB40ADC9" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[190,353,2006,2030]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="15" pageNumber="138" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFC1FFE81CAEDAF3EB40ADC9" box="[190,353,2006,2030]" italics="true" pageId="15" pageNumber="138">Ilyobius nubilus</emphasis>
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||
</taxonomicName>
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||
, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm.
|
||
</paragraph>
|
||
</caption>
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||
<subSubSection id="C310654EFFDEFFF71C87DDB3EE80AB42" pageId="16" pageNumber="139" type="materials_examined">
|
||
<paragraph id="8BB536C5FFDEFFF71C87DDB3EB82AAF2" blockId="16.[151,1437,150,1221]" pageId="16" pageNumber="139">
|
||
<emphasis id="B97EEAD7FFDEFFF71C87DDB3EBA7AA97" bold="true" box="[151,390,150,176]" pageId="16" pageNumber="139">Material examined.</emphasis>
|
||
<materialsCitation id="3B623C98FFDEFFF71D82DDB2EB82AAF2" collectingDate="1896-11-09" collectionCode="MZUSP" country="Brazil" location="Luederwaldt" municipality="Ypiranga" pageId="16" pageNumber="139" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
|
||
<typeStatus id="54B18867FFDEFFF71D82DDB2EBDEAA96" box="[402,511,151,177]" pageId="16" pageNumber="139" type="holotype">Holotype</typeStatus>
|
||
: gender unknown (abdomen missing),
|
||
<collectingCountry id="F31D7655FFDEFFF71FDBDDB2EE32AA96" box="[971,1043,151,177]" name="Brazil" pageId="16" pageNumber="139">Brazil</collectingCountry>
|
||
;
|
||
<collectingRegion id="49CEF827FFDEFFF71833DDB2EEBCAA96" box="[1059,1181,151,177]" country="Brazil" name="Sao Paulo" pageId="16" pageNumber="139">São Paulo</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BD1ACBFFFDEFFF718BCDDB2EF33AA96" box="[1196,1298,151,177]" pageId="16" pageNumber="139">Ypiranga</collectingMunicipality>
|
||
<date id="FFB41005FFDEFFF7190DDDB2EFB9AA96" box="[1309,1432,151,177]" pageId="16" pageNumber="139" value="1896-11-09">
|
||
<collectingDate id="EFF0E9EDFFDEFFF7190DDDB2EFB9AA96" box="[1309,1432,151,177]" pageId="16" pageNumber="139" value="1896-11-09">09.xi.1896</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8ED5601EFFDEFFF71C87DD9EEB04AAF2" LSID="urn:lsid:plazi:treatment:03A387D3FFC1FFF71C87DC67E966AEE2:8ED5601EFFDEFFF71C87DD9EEB04AAF2" box="[151,293,187,213]" country="Brazil" municipality="Ypiranga" name="Luederwaldt" pageId="16" pageNumber="139" stateProvince="Sao Paulo">Luederwaldt</location>
|
||
(
|
||
<collectionCode id="ED1BAE00FFDEFFF71D24DD9EEBB9AAF2" box="[308,408,187,213]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="16" pageNumber="139">MZUSP</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB536C5FFDEFFF71CD7DDFBEE80AB42" blockId="16.[151,1437,150,1221]" pageId="16" pageNumber="139">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD7DDFBEBB7AADE" bold="true" box="[199,406,222,249]" pageId="16" pageNumber="139">Taxonomic notes.</emphasis>
|
||
Head (
|
||
<figureCitation id="13312A40FFDEFFF71DF7DDFAE84EAADE" box="[487,623,223,249]" captionStart="FIGURE 12" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[296,1291,533,1946]" captionTargetId="figure-118@15.[296,1291,533,1946]" captionTargetPageId="15" captionText="FIGURE 12. (A–D) Ilyobius brasiliensis, holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E) Ilyobius nubilus, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="16" pageNumber="139">Fig. 12A, B</figureCitation>
|
||
): length =
|
||
<quantity id="4CF29B20FFDEFFF71EF6DDFBE91EAADE" box="[742,831,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="139" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; width (at the widest point) =
|
||
<quantity id="4CF29B20FFDEFFF7189DDDFAEEC7AADE" box="[1165,1254,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="16" pageNumber="139" unit="mm" value="1.9">1.9 mm</quantity>
|
||
; region between the posterior margin of the eyes =
|
||
<quantity id="4CF29B20FFDEFFF71E21DC26E8AFAB3A" box="[561,654,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="139" unit="mm" value="1.6">1.6 mm</quantity>
|
||
. Forewing: length =
|
||
<quantity id="4CF29B20FFDEFFF71F95DC26E9ECAB3A" box="[901,973,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="139" unit="mm" value="9.0">9 mm</quantity>
|
||
; width =
|
||
<quantity id="4CF29B20FFDEFFF71850DC26EEBCAB3A" box="[1088,1181,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="139" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; 4X longer than wide proximally, 3X longer than wide distally; posterior margin almost straight (
|
||
<figureCitation id="13312A40FFDEFFF71801DC02EE5CAB66" box="[1041,1149,295,321]" captionStart="FIGURE 12" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[296,1291,533,1946]" captionTargetId="figure-118@15.[296,1291,533,1946]" captionTargetPageId="15" captionText="FIGURE 12. (A–D) Ilyobius brasiliensis, holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E) Ilyobius nubilus, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="16" pageNumber="139">Fig. 12C</figureCitation>
|
||
). Specimen deteriorated, lacking abdomen, genitalia, antenna flagellum and hindwings, only remaining left forewing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C310654EFFDEFFF71CD7DC4BE8B3ABAE" box="[199,658,366,393]" pageId="16" pageNumber="139" type="distribution">
|
||
<paragraph id="8BB536C5FFDEFFF71CD7DC4BE8B3ABAE" blockId="16.[151,1437,150,1221]" box="[199,658,366,393]" pageId="16" pageNumber="139">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD7DC4BEB40ABAF" bold="true" box="[199,353,366,392]" pageId="16" pageNumber="139">Distribution.</emphasis>
|
||
<collectingCountry id="F31D7655FFDEFFF71D77DC4AEBEEABAE" box="[359,463,367,393]" name="Brazil" pageId="16" pageNumber="139">BRAZIL</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CEF827FFDEFFF71DCBDC4AE8AFABAE" box="[475,654,367,393]" country="Brazil" name="Sao Paulo" pageId="16" pageNumber="139">São Paulo State</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C310654EFFDEFFF71CD7DCB6E966AEE2" pageId="16" pageNumber="139" type="discussion">
|
||
<paragraph id="8BB536C5FFDEFFF71CD7DCB6E86DA9A6" blockId="16.[151,1437,150,1221]" pageId="16" pageNumber="139">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD7DCB6EB1AAB8A" bold="true" box="[199,315,403,429]" pageId="16" pageNumber="139">Remarks.</emphasis>
|
||
In addition to same distribution area, similarities on forewing shape and general coloration were characters used by
|
||
<bibRefCitation id="EF9B4B34FFDEFFF71D7FDC92E815ABF6" author="Liu, X. & Hayashi, F. & Yang, D." box="[367,564,438,465]" pageId="16" pageNumber="139" pagination="18 - 49" refId="ref7542" refString="Liu, X., Hayashi, F. & Yang, D. (2015 a) Phylogeny of the family Sialidae (Insecta: Megaloptera) inferred from morphological data, with implications for generic classification and historical biogeography. Cladistics, 31, 18 - 49. https: // doi. org / 10.1111 / cla. 12071" type="journal article" year="2015">
|
||
Liu
|
||
<emphasis id="B97EEAD7FFDEFFF71D8DDC92EBF7ABF6" box="[413,470,439,465]" italics="true" pageId="16" pageNumber="139">et al.</emphasis>
|
||
(2015a)
|
||
</bibRefCitation>
|
||
to support
|
||
<taxonomicName id="4C0A4D46FFDEFFF71EA5DC92E96FABF6" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[693,846,439,465]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71EA5DC92E96FABF6" box="[693,846,439,465]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
as junior synonym of
|
||
<taxonomicName id="4C0A4D46FFDEFFF7185CDC92EE97ABF6" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[1100,1206,439,465]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF7185CDC92EE97ABF6" box="[1100,1206,439,465]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
. However, the color pattern seems to be well conserved within the
|
||
<taxonomicName id="4C0A4D46FFDEFFF71E85DCFEE967ABD2" box="[661,838,475,501]" pageId="16" pageNumber="139">
|
||
<emphasis id="B97EEAD7FFDEFFF71E85DCFEE8CEABD2" box="[661,751,475,501]" italics="true" pageId="16" pageNumber="139">Ilyobius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
groups (
|
||
<emphasis id="B97EEAD7FFDEFFF71FB8DCFEE9E8ABD3" box="[936,969,475,500]" italics="true" pageId="16" pageNumber="139">e.g</emphasis>
|
||
., members of
|
||
<taxonomicName id="4C0A4D46FFDEFFF71877DCFEEED2ABD3" baseAuthorityName="Banks" baseAuthorityYear="1901" box="[1127,1267,475,500]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B97EEAD7FFDEFFF71877DCFEEED2ABD3" box="[1127,1267,475,500]" italics="true" pageId="16" pageNumber="139">I. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
group share an orange head with postero-ocular black marks, in contrast to the
|
||
<taxonomicName id="4C0A4D46FFDEFFF71F78DCDAE9C3A83E" authorityName="McLachlan" authorityYear="1981" box="[872,994,511,537]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71F78DCDAE9C3A83E" box="[872,994,511,537]" italics="true" pageId="16" pageNumber="139">I. chilensis</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<taxonomicName id="4C0A4D46FFDEFFF7182ADCDAEE85A83E" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[1082,1188,511,537]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF7182ADCDAEE85A83E" box="[1082,1188,511,537]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D46FFDEFFF718CDDCDAEF68A83E" baseAuthorityName="Contreras-Ramos, Fiorentin & Urakami" baseAuthorityYear="2005" box="[1245,1353,511,537]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="hauseri">
|
||
<emphasis id="B97EEAD7FFDEFFF718CDDCDAEF68A83E" box="[1245,1353,511,537]" italics="true" pageId="16" pageNumber="139">I. hauseri</emphasis>
|
||
</taxonomicName>
|
||
, which share a black head on vertex, orange laterally). Furthermore, the similarities on forewing shape of
|
||
<taxonomicName id="4C0A4D46FFDEFFF71912DF06EFBDA81A" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[1282,1436,547,573]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71912DF06EFBDA81A" box="[1282,1436,547,573]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D46FFDEFFF71CD6DF62EB0EA846" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[198,303,583,609]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD6DF62EB0EA846" box="[198,303,583,609]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
was not corroborated in this study. Although these two species share the forewing costal area slightly expanded and with similar wing venation, the general shape of the forewing is different in each species. In
|
||
<taxonomicName id="4C0A4D46FFDEFFF71923DF4EEFBDA8A2" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[1331,1436,619,645]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF71923DF4EEFBDA8A2" box="[1331,1436,619,645]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
the forewing is 3X longer than wide proximally, 3X longer than wide distally, and the posterior margin is slightly rounded (
|
||
<figureCitation id="13312A40FFDEFFF71D12DF96EB4BA8EA" box="[258,362,691,717]" captionStart="FIGURE 12" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[296,1291,533,1946]" captionTargetId="figure-118@15.[296,1291,533,1946]" captionTargetPageId="15" captionText="FIGURE 12. (A–D) Ilyobius brasiliensis, holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E) Ilyobius nubilus, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="16" pageNumber="139">Fig. 12D</figureCitation>
|
||
). On the other hand, in
|
||
<taxonomicName id="4C0A4D46FFDEFFF71E7FDF96E926A8EA" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[623,775,691,717]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71E7FDF96E926A8EA" box="[623,775,691,717]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
the forewing is 4X longer than wide proximally, 3X longer than wide distally, and the posterior margin is almost straight (
|
||
<figureCitation id="13312A40FFDEFFF71F58DFF2E98FA8D6" box="[840,942,727,753]" captionStart="FIGURE 12" captionStartId="15.[151,250,1970,1994]" captionTargetBox="[296,1291,533,1946]" captionTargetId="figure-118@15.[296,1291,533,1946]" captionTargetPageId="15" captionText="FIGURE 12. (A–D) Ilyobius brasiliensis, holotype; (A) head, lateral; (B) head, dorsal; (C) hindwing; D) holotype information; (E) Ilyobius nubilus, hindwing, female. Scale bars, A, B = 0.5 mm; C, E = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611464" httpUri="https://zenodo.org/record/10611464/files/figure.png" pageId="16" pageNumber="139">Fig. 12C</figureCitation>
|
||
).
|
||
<bibRefCitation id="EF9B4B34FFDEFFF71FD1DFF2EEA2A8D6" author="Liu, X. & Hayashi, F. & Yang, D." box="[961,1155,726,753]" pageId="16" pageNumber="139" pagination="55 - 63" refId="ref7604" refString="Liu, X., Hayashi, F. & Yang, D. (2015 b) Taxonomic notes of the Neotropical alderfly genus Ilyobius Enderlein, 1910 (Megaloptera, Sialidae), with description of a new species. Deutsche Entomologische Zeitschrift, 62, 55 - 63. https: // doi. org / 10.3897 / dez. 62.4481" type="journal article" year="2015">
|
||
Liu
|
||
<emphasis id="B97EEAD7FFDEFFF71FFEDFF2EE04A8D6" box="[1006,1061,727,753]" italics="true" pageId="16" pageNumber="139">et al.</emphasis>
|
||
(2015b)
|
||
</bibRefCitation>
|
||
highlighted an important diagnostic characteristic of
|
||
<taxonomicName id="4C0A4D46FFDEFFF71DD8DFDEE811A932" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[456,560,763,789]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF71DD8DFDEE811A932" box="[456,560,763,789]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
: MA vein of hindwing proximally with a veinlet fused to MP (
|
||
<bibRefCitation id="EF9B4B34FFDEFFF718F2DFDEEFB5A932" author="Liu, X. & Hayashi, F. & Yang, D." box="[1250,1428,762,789]" pageId="16" pageNumber="139" pagination="55 - 63" refId="ref7604" refString="Liu, X., Hayashi, F. & Yang, D. (2015 b) Taxonomic notes of the Neotropical alderfly genus Ilyobius Enderlein, 1910 (Megaloptera, Sialidae), with description of a new species. Deutsche Entomologische Zeitschrift, 62, 55 - 63. https: // doi. org / 10.3897 / dez. 62.4481" type="journal article" year="2015">
|
||
Liu
|
||
<emphasis id="B97EEAD7FFDEFFF71900DFDEEF66A932" box="[1296,1351,763,789]" italics="true" pageId="16" pageNumber="139">et al.</emphasis>
|
||
2015b
|
||
</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13312A40FFDEFFF71C8FDE3AEAD6A91E" box="[159,247,799,825]" captionStart="FIGURE 6" captionStartId="9.[151,250,1356,1380]" captionTargetBox="[151,1437,181,1332]" captionTargetId="figure-19@9.[151,1437,181,1332]" captionTargetPageId="9" captionText="FIGURE 6. Ilyobius nubilus, male. (A) forewing; (B) hindwing, the arrow indicates the veinlet between MA and MP. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10611452" httpUri="https://zenodo.org/record/10611452/files/figure.png" pageId="16" pageNumber="139">Fig. 6B</figureCitation>
|
||
); however, it was not possible to verify this characteristic because the hindwings of
|
||
<taxonomicName id="4C0A4D46FFDEFFF7188CDE3AEF15A91E" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[1180,1332,799,825]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF7188CDE3AEF15A91E" box="[1180,1332,799,825]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54B18867FFDEFFF7192ADE3AEFBDA91E" box="[1338,1436,799,825]" pageId="16" pageNumber="139" type="holotype">holotype</typeStatus>
|
||
because they are missing and were not previously described. Therefore, based on available evidence,
|
||
<taxonomicName id="4C0A4D46FFDEFFF71914DE66EFBDA97A" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[1284,1436,835,861]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71914DE66EFBDA97A" box="[1284,1436,835,861]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
is here hypothesized as a valid species.
|
||
</paragraph>
|
||
<paragraph id="8BB536C5FFDEFFF71CD7DEAEEBBDAE36" blockId="16.[151,1437,150,1221]" pageId="16" pageNumber="139">
|
||
<taxonomicName id="4C0A4D46FFDEFFF71CD7DEAEEB8AA982" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[199,427,907,933]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD7DEAEEB8AA982" box="[199,427,907,933]" italics="true" pageId="16" pageNumber="139">Ilyobius brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
seems to be closely related to
|
||
<taxonomicName id="4C0A4D46FFDEFFF71F1EDEAEE958A982" baseAuthorityName="Navas" baseAuthorityYear="1933" box="[782,889,907,933]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B97EEAD7FFDEFFF71F1EDEAEE958A982" box="[782,889,907,933]" italics="true" pageId="16" pageNumber="139">I. nubilus</emphasis>
|
||
</taxonomicName>
|
||
by sharing the general body color, both species have a brown to black head, with two lighter regions extending laterally from the gena region to the base of the head. Thus,
|
||
<taxonomicName id="4C0A4D46FFDEFFF71CCEDEF6EB56A9CA" baseAuthorityName="Navas" baseAuthorityYear="1936" box="[222,375,979,1005]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71CCEDEF6EB56A9CA" box="[222,375,979,1005]" italics="true" pageId="16" pageNumber="139">I. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
appears to belong to
|
||
<taxonomicName id="4C0A4D46FFDEFFF71E7CDEF6E8C7A9CA" authorityName="McLachlan" authorityYear="1981" box="[620,742,979,1005]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="chilensis">
|
||
<emphasis id="B97EEAD7FFDEFFF71E7CDEF6E8C7A9CA" box="[620,742,979,1005]" italics="true" pageId="16" pageNumber="139">I. chilensis</emphasis>
|
||
</taxonomicName>
|
||
group because it has a dark pronotum similar to that of other members of this group.
|
||
</paragraph>
|
||
<paragraph id="8BB536C5FFDEFFF71CD7D93EE966AEE2" blockId="16.[151,1437,150,1221]" pageId="16" pageNumber="139">
|
||
<taxonomicName id="4C0A4D46FFDEFFF71CD7D93EEB00AE12" authorityName="Enderlein" authorityYear="1910" box="[199,289,1051,1077]" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EEAD7FFDEFFF71CD7D93EEB00AE12" box="[199,289,1051,1077]" italics="true" pageId="16" pageNumber="139">Ilyobius</emphasis>
|
||
</taxonomicName>
|
||
is difficult to sample in field studies, which is reflected in few studies on the genus and low representation of specimens in zoological collections (
|
||
<bibRefCitation id="EF9B4B34FFDEFFF71E70D91AE94CAE7E" author="Contreras-Ramos, A." box="[608,877,1087,1113]" pageId="16" pageNumber="139" pagination="808 - 814" refId="ref7242" refString="Contreras-Ramos, A. (2008) Notes on some Neotropical alderflies (Sialidae: Megaloptera). Annals of the Entomological Society of America, 101, 808 - 814. https: // doi. org / 10.1093 / aesa / 101.5.808" type="journal article" year="2008">Contreras-Ramos 2008</bibRefCitation>
|
||
). In addition, the lack of
|
||
<typeStatus id="54B18867FFDEFFF71887D91AEEE6AE7E" box="[1175,1223,1087,1113]" pageId="16" pageNumber="139">type</typeStatus>
|
||
material and poor conditions of
|
||
<typeStatus id="54B18867FFDEFFF71D23D946EB42AE5A" box="[307,355,1123,1149]" pageId="16" pageNumber="139">type</typeStatus>
|
||
specimens are difficulties to be overcome in order to elucidate the taxonomic status of
|
||
<taxonomicName id="4C0A4D46FFDEFFF71952D946EAD1AE86" class="Insecta" family="Sialidae" genus="Ilyobius" kingdom="Animalia" order="Megaloptera" pageId="16" pageNumber="139" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B97EEAD7FFDEFFF71952D946EFBDAE5A" box="[1346,1436,1123,1149]" italics="true" pageId="16" pageNumber="139">Ilyobius</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Collection of specimens at the
|
||
<typeStatus id="54B18867FFDEFFF71E58D9A2E859AE86" box="[584,632,1159,1185]" pageId="16" pageNumber="139">type</typeStatus>
|
||
locality and laboratory rearing make it possible to fill gaps in knowledge about the taxonomy and biological traits of these rare insects.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |