153 lines
20 KiB
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153 lines
20 KiB
XML
<document id="1F1BC310684E8A6AAACE2C83D00F820E" ID-CLB-Dataset="8401" ID-DOI="10.11646/zootaxa.4920.2.1" ID-GBIF-Dataset="ec1a3d33-651e-499c-b775-b42a131ec75a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4475126" ID-ZooBank="C369AA2D-29D5-4DDF-BFE2-EA4FC4AD2703" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611818479954" checkinUser="plazi" docAuthor="Kajihara, Hiroshi" docDate="2021" docId="03A387C8FFDEFFE8FF209F6DAB90C509" docLanguage="en" docName="zootaxa.4920.2.1.pdf" docOrigin="Zootaxa 4920 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oerstediina" docType="treatment" docVersion="7" lastPageNumber="158" masterDocId="FF9AFFB0FFD8FFEFFFB79937A815C752" masterDocTitle="Higher classification of the Monostilifera (Nemertea: Hoplonemertea)" masterLastPageNumber="199" masterPageNumber="151" pageNumber="157" updateTime="1698925697009" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="33F7B69978C018EF5E55B6EED48DD7CF">Higher classification of the Monostilifera (Nemertea: Hoplonemertea)</mods:title>
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<mods:namePart id="50DFE83CB280851393366AF1686DE78C">Kajihara, Hiroshi</mods:namePart>
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<mods:title id="14E9F79915C570E61D14B03F508EAF41">Zootaxa</mods:title>
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<mods:date id="E01B1915D9BFE7831D0C03A3390A5D6E">2021</mods:date>
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<treatment id="03A387C8FFDEFFE8FF209F6DAB90C509" ID-DOI="http://doi.org/10.5281/zenodo.4519275" ID-GBIF-Taxon="177309804" ID-Zenodo-Dep="4519275" LSID="urn:lsid:plazi:treatment:03A387C8FFDEFFE8FF209F6DAB90C509" httpUri="http://treatment.plazi.org/id/03A387C8FFDEFFE8FF209F6DAB90C509" lastPageId="7" lastPageNumber="158" pageId="6" pageNumber="157">
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<subSubSection id="C3106555FFDEFFE9FF209F6DA9E3C1EE" pageId="6" pageNumber="157" type="nomenclature">
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<paragraph id="8BB536DEFFDEFFE9FF209F6DA9E7C126" blockId="6.[151,498,1626,1652]" box="[151,498,1626,1652]" pageId="6" pageNumber="157">
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<heading id="D0FD81B2FFDEFFE9FF209F6DA9E7C126" bold="true" box="[151,498,1626,1652]" fontSize="11" level="1" pageId="6" pageNumber="157" reason="1">
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<emphasis id="B97EEACCFFDEFFE9FF209F6DA9E7C126" bold="true" box="[151,498,1626,1652]" pageId="6" pageNumber="157">
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Infraorder
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<taxonomicName id="4C0A4D5DFFDEFFE9FE929F6DA9E7C126" box="[293,498,1626,1652]" class="Enopla" higherTaxonomySource="GBIF" infraOrder="Oerstediina" kingdom="Animalia" order="Monostilifera" pageId="6" pageNumber="157" phylum="Nemertea" rank="infraOrder">OERSTEDIINA</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB536DEFFDEFFE9FF209F94A9E3C1EE" blockId="6.[151,1436,1699,1833]" box="[151,502,1699,1725]" pageId="6" pageNumber="157">
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<emphasis id="B97EEACCFFDEFFE9FF209F94A9E3C1EE" box="[151,502,1699,1725]" italics="true" pageId="6" pageNumber="157">Morphological circumscription</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3106555FFDEFFE9FF709FF0A96DC08F" pageId="6" pageNumber="157" type="description">
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<paragraph id="8BB536DEFFDEFFE9FF709FF0A9C5C07B" blockId="6.[151,1436,1699,1833]" pageId="6" pageNumber="157">Eumonostiliferans with the cephalic blood vessels protruding into the rhynchocoel as they pass through the cerebral ring, often forming a pair of vascular plugs; the mid-dorsal vessel not penetrating into the rhynchocoel to form a single vascular plug.</paragraph>
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<paragraph id="8BB536DEFFDEFFE9FF209E60A959C023" blockId="6.[151,1436,1879,2013]" box="[151,332,1879,1905]" pageId="6" pageNumber="157">
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<emphasis id="B97EEACCFFDEFFE9FF209E60A959C023" box="[151,332,1879,1905]" italics="true" pageId="6" pageNumber="157">Clade definition</emphasis>
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</paragraph>
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<paragraph id="8BB536DEFFDEFFE9FF709E4CA9F5C0EB" blockId="6.[151,1436,1879,2013]" pageId="6" pageNumber="157">
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Eumonostiliferans that are more closely related to
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<taxonomicName id="4C0A4D5DFFDEFFE9FCB39E4CACA0C0C4" authority="(Abildgaard, 1806)" baseAuthorityName="Abildgaard" baseAuthorityYear="1806" box="[772,1205,1915,1942]" class="Enopla" family="Oerstediidae" genus="Oerstedia" kingdom="Animalia" order="Monostilifera" pageId="6" pageNumber="157" phylum="Nemertea" rank="species" species="dorsalis">
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<emphasis id="B97EEACCFFDEFFE9FCB39E4CABC0C0C7" box="[772,981,1915,1941]" italics="true" pageId="6" pageNumber="157">Oerstedia dorsalis</emphasis>
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(
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<bibRefCitation id="EF9B4B2FFFDEFFE9FC529E4CACB8C0C4" author="Abildgaard, P. C." box="[997,1197,1915,1942]" pageId="6" pageNumber="157" pagination="25" refId="ref18962" refString="Abildgaard, P. C. (1806) Planaria dorsalis. In: Muller, O. F. (Author) & Abildgaard, P. C., Vahl, M., Holten, J. S. & Rathke, J. (who prepared descriptions and figures), Zoologia Danica sev animalium Daniae et Norvegiae rariorvm ac minus notorvm descriptiones et historia. Volvmen qvartvm. Vol. 4. Explicationi iconvm fascicvli qvarti ejvsdem operis inserviens. N. Christensen, Havniae (locative case of Havnia, Copenhagen), pp. 25. [https: // www. biodiversitylibrary. org / item / 104862 # page / 231 / mode / 1 up]" type="book chapter" year="1806">Abildgaard, 1806</bibRefCitation>
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)
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</taxonomicName>
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than to
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<taxonomicName id="4C0A4D5DFFDEFFE9FAA49E4CA9C9C0EB" authority="(Johnston, 1828)" baseAuthorityName="Johnston" baseAuthorityYear="1828" class="Enopla" family="Amphiporidae" genus="Amphiporus" kingdom="Animalia" order="Monostilifera" pageId="6" pageNumber="157" phylum="Nemertea" rank="species" species="lactifloreus">
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<emphasis id="B97EEACCFFDEFFE9FAA49E4CA903C0EB" italics="true" pageId="6" pageNumber="157">Amphiporus lactifloreus</emphasis>
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(
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<bibRefCitation id="EF9B4B2FFFDEFFE9FE939EA8A9C1C0EB" author="Johnston, G." box="[292,468,1951,1977]" pageId="6" pageNumber="157" pagination="486 - 491" refId="ref27377" refString="Johnston, G. (1828) Contributions to the British fauna. Zoological Journal, 3, 486 - 491. [https: // www. biodiversitylibrary. org / item / 19597 # page / 548 / mode / 1 up]" type="journal article" year="1828">Johnston, 1828</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BB536DEFFDEFFE9FF209EF4A96DC08F" blockId="6.[151,1436,1879,2013]" box="[151,376,1987,2013]" pageId="6" pageNumber="157">
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<emphasis id="B97EEACCFFDEFFE9FF209EF4A96DC08F" box="[151,376,1987,2013]" italics="true" pageId="6" pageNumber="157">Constituent families</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3106555FFDFFFE8FF7099A0ABC5C7E3" box="[199,976,151,177]" pageId="7" pageNumber="158" type="reference_group">
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<paragraph id="8BB536DEFFDFFFE8FF7099A0ABC5C7E3" blockId="7.[199,976,151,177]" box="[199,976,151,177]" pageId="7" pageNumber="158">
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<taxonomicName id="4C0A4D5DFFDFFFE8FF7099A0AA33C7E3" authority="Chernyshev, 1993" authorityName="Chernyshev" authorityYear="1993" box="[199,550,151,177]" class="Enopla" family="Oerstediidae" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="family">
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Oerstediidae
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<bibRefCitation id="EF9B4B2FFFDFFFE8FEEB99A0AA33C7E3" author="Chernyshev, A. V." box="[348,550,151,177]" pageId="7" pageNumber="158" pagination="11 - 20" refId="ref20729" refString="Chernyshev, A. V. (1993) A review of the genera of nemerteans allied to Oerstedia (Monostilifera, Tetrastemmatidae) with description of four new species. Zoologischeskii Zhurnal, 72, 11 - 20." type="journal article" year="1993">Chernyshev, 1993</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C0A4D5DFFDFFFE8FDEB99A0ABD9C7E3" authority="Gibson, 1990 a" authorityName="Gibson" authorityYear="1990" box="[604,972,151,177]" class="Enopla" family="Plectonemertidae" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="family">
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Plectonemertidae
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<bibRefCitation id="EF9B4B2FFFDFFFE8FC9299A0ABD9C7E3" author="Gibson, R." box="[805,972,151,177]" pageId="7" pageNumber="158" refId="ref24160" refString="Gibson, R. (1990 a) The macrobenthic nemertean fauna of Hong Kong. In: Morton, B. S. (Ed.), Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Vol. 1. Hong Kong University Press, Hong Kong, pp. 33 - 212." type="book" year="1990">Gibson, 1990a</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3106555FFDFFFE8FF2099EEAB90C509" pageId="7" pageNumber="158" type="discussion">
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<paragraph id="8BB536DEFFDFFFE8FF2099EEA8ECC7A1" blockId="7.[151,1437,217,604]" box="[151,249,217,243]" pageId="7" pageNumber="158">
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<emphasis id="B97EEACCFFDFFFE8FF2099EEA8ECC7A1" box="[151,249,217,243]" italics="true" pageId="7" pageNumber="158">Remarks</emphasis>
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</paragraph>
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<paragraph id="8BB536DEFFDFFFE8FF7099C9AB90C509" blockId="7.[151,1437,217,604]" pageId="7" pageNumber="158">
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As far as I am aware,
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<bibRefCitation id="EF9B4B2FFFDFFFE8FE0099C9AAD3C645" author="Moore, J. & Gibson, R." box="[439,710,253,280]" pageId="7" pageNumber="158" pagination="75 - 86" refId="ref30674" refString="Moore, J. & Gibson, R. (1988) Marine relatives of terrestrial nemerteans: the genus Prosadenoporus Burger, 1890 (Hoplonemertea). Hydrobiologia, 156, 75 - 86. https: // doi. org / 10.1007 / BF 00027980" type="journal article" year="1988">Moore & Gibson (1988)</bibRefCitation>
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were the first to envisage
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<taxonomicName id="4C0A4D5DFFDFFFE8FC5F99CAAD3CC645" authority="Quatrefages, 1846" authorityName="Quatrefages" authorityYear="1846" box="[1000,1321,253,279]" class="Enopla" family="Oerstediidae" genus="Oerstedia" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="genus">
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<emphasis id="B97EEACCFFDFFFE8FC5F99CAAC43C645" box="[1000,1110,253,279]" italics="true" pageId="7" pageNumber="158">Oerstedia</emphasis>
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<bibRefCitation id="EF9B4B2FFFDFFFE8FBEA99CAAD3CC645" author="Quatrefages, A. de" box="[1117,1321,253,279]" pageId="7" pageNumber="158" pagination="173 - 303" refId="ref31808" refString="Quatrefages, A. de (1846) Etude sur les types inferieurs de l'embrachement des anneles. Annales des Sciences Naturelles, Zoologie, Serie 3, 6, 173 - 303. [https: // www. biodiversitylibrary. org / item / 47986 # page / 179 / mode / 1 up]" type="journal article" year="1846">Quatrefages, 1846</bibRefCitation>
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</taxonomicName>
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as closely related to
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<taxonomicName id="4C0A4D5DFFDFFFE8FEB19816A9DDC669" authorityName="Gibson" authorityYear="1990" box="[262,456,289,315]" class="Enopla" family="Plectonemertidae" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="family">Plectonemertidae</taxonomicName>
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by having two vascular plugs. This conjecture was later confirmed with molecular phylogenetic analyses, first by
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<bibRefCitation id="EF9B4B2FFFDFFFE8FE719871AACEC60D" author="Mateos, E. & Giribet, G." box="[454,731,325,352]" pageId="7" pageNumber="158" pagination="235 - 243" refId="ref30003" refString="Mateos, E. & Giribet, G. (2008) Exploring the molecular diversity of terrestrial nemerteans (Hoplonemertea, Monostilifera, Acteonemertidae) in a continental landmass. Zoologica Scripta, 37, 235 - 243. https: // doi. org / 10.1111 / j. 1463 - 6409.2008.00324. x" type="journal article" year="2008">Mateos & Giribet (2008)</bibRefCitation>
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and then more convincingly by
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<bibRefCitation id="EF9B4B2FFFDFFFE8FBF69872AD40C60D" author="Kajihara, H. & Kuris, A." box="[1089,1365,325,351]" pageId="7" pageNumber="158" pagination="1 - 15" refId="ref28316" refString="Kajihara, H. & Kuris, A. (2013) Ovicides paralithodis (Nemertea, Carcinonemertidae), a new species of symbiotic egg predator of the red king crab Paralithodes camtchaticus (Tilesius, 1815) (Decapoda, Anomura). ZooKeys, 258, 1 - 15. http: // doi. org / 10.3897 / zookeys. 258.4260" type="journal article" year="2013">Kajihara & Kuris (2013)</bibRefCitation>
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,
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<bibRefCitation id="EF9B4B2FFFDFFFE8FAD79872A90BC6D1" author="Kvist, S. & Laumer, C. E. & Junoy, J. & Giribet, G." pageId="7" pageNumber="158" pagination="287 - 308" refId="ref29216" refString="Kvist, S., Laumer, C. E., Junoy, J. & Giribet, G. (2014) New insights into the phylogeny, systematics and DNA barcoding of Nemertea. Invertebrate Systematics, 28, 287 - 308. https: // doi. org / 10.1071 / IS 13061" type="journal article" year="2014">
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Kvist
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<emphasis id="B97EEACCFFDFFFE8FF20985DA8DFC6D1" box="[151,202,361,387]" italics="true" pageId="7" pageNumber="158">et al</emphasis>
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. (2014
|
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</bibRefCitation>
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,
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<bibRefCitation id="EF9B4B2FFFDFFFE8FE9C985EA972C6D6" author="Kvist, S. & Chernyshev, A. V. & Giribet, G." box="[299,359,361,388]" pageId="7" pageNumber="158" pagination="105 - 119" refId="ref29273" refString="Kvist, S., Chernyshev, A. V. & Giribet, G. (2015) Phylogeny of Nemertea with special interest in the placement of diversity from Far East Russia and northeast Asia. Hydrobiologia, 760, 105 - 119. https: // doi. org / 10.1007 / s 10750 - 015 - 2310 - 5" type="journal article" year="2015">2015</bibRefCitation>
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), and
|
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<bibRefCitation id="EF9B4B2FFFDFFFE8FE1A985EAB3FC6D1" author="Chernyshev, A. V. & Polyakova, N. E." box="[429,810,361,387]" pageId="7" pageNumber="158" pagination="64 - 73" refId="ref21353" refString="Chernyshev, A. V. & Polyakova, N. E. (2018 a) Nemerteans of the Vema-TRANSIT expedition: First data on diversity with description of two new genera and species. Deep-Sea Research, Part II, 148, 64 - 73. https: // doi. org / 10.1016 / j. dsr 2.2017.06.004" type="journal article" year="2018">
|
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Chernyshev & Polyakova (2018
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<bibRefCitation id="EF9B4B2FFFDFFFE8FCAA985EAB3FC6D1" author="Chernyshev, A. V. & Polyakova, N. E." box="[797,810,361,387]" pageId="7" pageNumber="158" pagination="102148" refId="ref21485" refString="Chernyshev, A. V. & Polyakova, N. E. (2019) Nemerteans from the deep-sea expedition KuramBio II with descriptions of three new hoplonemerteans from the Kuril-Kamchatka Trench. Progress in Oceanography, 178, 102148. https: // doi. org / 10.1016 / j. pocean. 2019.102148" type="journal article" year="2019">a</bibRefCitation>
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</bibRefCitation>
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, b, 2019) [
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<taxonomicName id="4C0A4D5DFFDFFFE8FC1F985EAC03C6D1" authorityName="Quatrefages" authorityYear="1846" box="[936,1046,361,387]" class="Enopla" family="Oerstediidae" genus="Oerstedia" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="genus">
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<emphasis id="B97EEACCFFDFFFE8FC1F985EAC03C6D1" box="[936,1046,361,387]" italics="true" pageId="7" pageNumber="158">Oerstedia</emphasis>
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||
</taxonomicName>
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was not included in the extensive analysis by
|
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<bibRefCitation id="EF9B4B2FFFDFFFE8FEA998BAAA0FC6FA" author="Andrade, S. C. S. & Strand, M & Schwartz, M. & Chen, H. & Kajihara, H. & Sun, S. & Junoy, J. & Thiel, M. & Norenburg, J. L. & Turbeville, J. M. & Giribet, G. & Sundberg, P." box="[286,538,397,424]" pageId="7" pageNumber="158" pagination="141 - 159" refId="ref19155" refString="Andrade, S. C. S., Strand, M, Schwartz, M., Chen, H., Kajihara, H., D ˆ hren, J. v., Sun, S., Junoy, J., Thiel, M., Norenburg, J. L., Turbeville, J. M., Giribet, G. & Sundberg, P. (2012) Disentangling ribbon worm relationships: multi-locus analysis supports traditional classification on the phylum Nemertea. Cladistics, 28, 141 - 159. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00376. x" type="journal article" year="2012">
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Andrade
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<emphasis id="B97EEACCFFDFFFE8FE3098B9A9AFC6F5" box="[391,442,397,423]" italics="true" pageId="7" pageNumber="158">et al</emphasis>
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. (2012)
|
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</bibRefCitation>
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]. The clade comprising
|
||
<taxonomicName id="4C0A4D5DFFDFFFE8FC9C98BAABFFC6F5" authorityName="Chernyshev" authorityYear="2005" box="[811,1002,397,423]" class="Enopla" family="Acteonemertidae" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="family">Acteonemertidae</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C0A4D5DFFDFFFE8FBBB98BAAC6FC6F5" authorityName="Quatrefages" authorityYear="1846" box="[1036,1146,397,423]" class="Enopla" family="Oerstediidae" genus="Oerstedia" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="158" phylum="Nemertea" rank="genus">
|
||
<emphasis id="B97EEACCFFDFFFE8FBBB98BAAC6FC6F5" box="[1036,1146,397,423]" italics="true" pageId="7" pageNumber="158">Oerstedia</emphasis>
|
||
</taxonomicName>
|
||
and their kin was named
|
||
<taxonomicName id="4C0A4D5DFFDFFFE8FF209886A90DC699" box="[151,280,433,459]" class="Enopla" higherTaxonomySource="GBIF" infraOrder="Oerstediina" kingdom="Animalia" order="Monostilifera" pageId="7" pageNumber="157" phylum="Nemertea" rank="infraOrder">Oerstediina</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B4B2FFFDFFFE8FE919886AA9FC699" author="Chernyshev, A. V. & Polyakova, N. E." box="[294,650,433,459]" pageId="7" pageNumber="158" pagination="102148" refId="ref21485" refString="Chernyshev, A. V. & Polyakova, N. E. (2019) Nemerteans from the deep-sea expedition KuramBio II with descriptions of three new hoplonemerteans from the Kuril-Kamchatka Trench. Progress in Oceanography, 178, 102148. https: // doi. org / 10.1016 / j. pocean. 2019.102148" type="journal article" year="2019">Chernyshev & Polyakova 2019</bibRefCitation>
|
||
), with neither Linnaean rank nor morphological circumscription. Here I place it at the infraorder rank. The suffix -ina must be used for the name of a subtribe when the taxon is placed on any of family group ranks in accordance with Article 29.2 of the Code (ICZN 1999); however, the Code (ICZN 1999) does not forbid the usage of -ina for ranks above the family group. While this may lead to confusion, I leave the ending as it was proposed by
|
||
<bibRefCitation id="EF9B4B2FFFDFFFE8FDBB9B76AB95C509" author="Chernyshev, A. V. & Polyakova, N. E." box="[524,896,577,603]" pageId="7" pageNumber="158" pagination="102148" refId="ref21485" refString="Chernyshev, A. V. & Polyakova, N. E. (2019) Nemerteans from the deep-sea expedition KuramBio II with descriptions of three new hoplonemerteans from the Kuril-Kamchatka Trench. Progress in Oceanography, 178, 102148. https: // doi. org / 10.1016 / j. pocean. 2019.102148" type="journal article" year="2019">Chernyshev & Polyakova (2019)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |