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<mods:title id="15E74DF440B9F07C02674F1375C91C8C">Revision of theGonioctena flavoplagiata species-group (Coleoptera: Chrysomelidae: Chrysomelinae), with descriptions of two new species from China and Laos</mods:title>
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<mods:namePart id="F6776C97001E6033EEF3F2A58510243F">Cho, Hee-Wook</mods:namePart>
<mods:affiliation id="9F4735D9F0CAD1FB9D884F04D5462456">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland</mods:affiliation>
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<mods:namePart id="D81B5630A488F988441451AEF461A64D">Borowiec, Lech</mods:namePart>
<mods:affiliation id="957700FCD7B892A30D05048523096BE8">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland</mods:affiliation>
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<treatment id="039B87BBFFE02467FE80BCC70E1BFE18" ID-DOI="http://doi.org/10.5281/zenodo.5448791" ID-GBIF-Taxon="194660307" ID-Zenodo-Dep="5448791" LSID="urn:lsid:plazi:treatment:039B87BBFFE02467FE80BCC70E1BFE18" httpUri="http://treatment.plazi.org/id/039B87BBFFE02467FE80BCC70E1BFE18" lastPageId="2" lastPageNumber="757" pageId="1" pageNumber="756">
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<heading id="D0C581C1FFE02464FE80BCC709F0FB35" bold="true" box="[293,909,1190,1215]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="756" reason="3">
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<taxonomicName id="4C324D2EFFE02464FE80BCC70896FB35" baseAuthorityName="Jacoby" baseAuthorityYear="1890" box="[293,747,1190,1215]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="756" phylum="Arthropoda" rank="species" species="flavoplagiata" subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE02464FE80BCC70BD9FB34" bold="true" box="[293,420,1191,1215]" italics="true" pageId="1" pageNumber="756">Gonioctena</emphasis>
(
<emphasis id="B946EABFFFE02464FE11BCC60833FB35" bold="true" box="[436,590,1190,1214]" italics="true" pageId="1" pageNumber="756">Asiphytodecta</emphasis>
)
<emphasis id="B946EABFFFE02464FDF8BCC60896FB35" bold="true" box="[605,747,1190,1214]" italics="true" pageId="1" pageNumber="756">flavoplagiata</emphasis>
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species-group
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<emphasis id="B946EABFFFE02464FFD4BCB60A9BFB66" bold="true" box="[113,230,1238,1261]" pageId="1" pageNumber="756">Diagnosis.</emphasis>
Body length
<quantity id="4CCA9B48FFE02464FED4BCB60B89FB66" box="[369,500,1238,1261]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.6" metricValueMin="6.2" pageId="1" pageNumber="756" unit="mm" value="6.9" valueMax="7.6" valueMin="6.2">6.27.6 mm</quantity>
. Dorsum brown to blackish brown with two pairs of yellowish brown spots on elytra. Anterior margin of labrum with tooth-like projection.Antennae reaching pronotal base, with last four antennomeres strongly widened and flattened. Elytral surface mostly covered with irregular, coarse and large punctures except humeri, elytral suture and lateral margins. Aedeagus variable in shape, very thin or thick with narrow or broad apical process, setose apicolaterally. Spermatheca C-shaped and very thin.
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<emphasis id="B946EABFFFE02464FF57BDD609BDFA46" bold="true" box="[242,960,1461,1486]" pageId="1" pageNumber="756">
Key to the species of the
<taxonomicName id="4C324D2EFFE02464FDAFBDD60963FA46" baseAuthorityName="Jacoby" baseAuthorityYear="1890" box="[522,798,1461,1486]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="756" phylum="Arthropoda" rank="species" species="flavoplagiata">
<emphasis id="B946EABFFFE02464FDAFBDD60963FA46" bold="true" box="[522,798,1461,1486]" italics="true" pageId="1" pageNumber="756">Gonioctena flavoplagiata</emphasis>
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species-group
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<keyLead id="30C39598FFE02464FFD4BD850E3DFA77" box="[113,1088,1509,1532]" pageId="1" pageNumber="756">1 Venter largely black to dark brown. ............................................................................... 2</keyLead>
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<keyLead id="30C39598FFE02464FFD4BE650E3DF997" box="[113,1088,1541,1565]" pageId="1" pageNumber="756">
Venter largely reddish-brown. ..................
<taxonomicName id="4C324D2EFFE02464FDC4BE650E3DF997" authority="Cho &amp; Borowiec, 2016" authorityName="Cho &amp; Borowiec" authorityYear="2016" box="[609,1088,1541,1565]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="756" phylum="Arthropoda" rank="species" species="mausonensis" subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE02464FDC4BE65093DF996" bold="true" box="[609,832,1541,1565]" pageId="1" pageNumber="756">
<emphasis id="B946EABFFFE02464FDC4BE650809F996" bold="true" box="[609,628,1541,1565]" italics="true" pageId="1" pageNumber="756">G</emphasis>
. (
<emphasis id="B946EABFFFE02464FD2FBE6508DFF996" bold="true" box="[650,674,1541,1565]" italics="true" pageId="1" pageNumber="756">A.</emphasis>
)
<emphasis id="B946EABFFFE02464FD17BE65093DF996" bold="true" box="[690,832,1541,1565]" italics="true" pageId="1" pageNumber="756">mausonensis</emphasis>
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<bibRefCitation id="EFA34B5CFFE02464FCE2BE650E3DF997" author="Cho &amp; Borowiec" box="[839,1088,1541,1564]" firstAuthor="Cho" pageId="1" pageNumber="756" pagination="168 - 184" refId="ref7222" refString="CHO H. W. &amp; BOROWIEC L. 2016: On the genus Gonioctena Chevrolat (Coleoptera: Chrysomelidae: Chrysomelinae), with descriptions of seven new species from the Oriental region and Palaearctic China. Zootaxa 4067: 168 - 184." type="journal article" year="2016">Cho &amp; Borowiec, 2016</bibRefCitation>
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2 Apical antennomere widely rounded at apex (
<figureCitation id="13092A28FFE02464FDD8BE450885F9B7" box="[637,760,1573,1596]" captionStart="Figs 1321. 1318" captionStartId="9.[113,150,1493,1512]" captionTargetBox="[147,1065,226,1404]" captionTargetId="figure-14@9.[111,1092,216,1418]" captionTargetPageId="9" captionText="Figs 1321. 1318 Gonioctena (A.) eburoides (Achard, 1924): 13 antenna; 14 aedeagus (dorsal and lateral views); 1517 variation of apex of aedeagus (1516 Sa Pa; 17 Tam Dao); 18 spermatheca. 1921 G. (A.) flavoplagiata (Jacoby, 1890): 19 antenna; 20 aedeagus, paralectotype (dorsal view, detail of apex in dorsal view and lateral view); 21 spermatheca. Scale bars = 1.0 mm (except Figs 18, 21 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310364" httpUri="https://zenodo.org/record/5310364/files/figure.png" pageId="1" pageNumber="756">Figs 13, 19</figureCitation>
,
<figureCitation id="13092A28FFE02464FCA6BE45095CF9B7" box="[771,801,1573,1596]" captionStart="Figs 2228. 2224" captionStartId="10.[151,188,1465,1484]" captionTargetBox="[174,1105,223,1419]" captionTargetId="figure-10@10.[148,1129,217,1454]" captionTargetPageId="10" captionText="Figs 2228. 2224 Gonioctena (A.) jindrai sp. nov.: 22 antenna; 23 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view); 24 spermatheca. 2526 G. (A.) mausonensis Cho &amp; Borowiec, 2016 (after CHO &amp; BOROWIEC 2016): 25 antenna; 26 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view). 2728 G. (A.) truncaticornis sp. nov.: 27 antenna; 28 aedeagus, holotype (dorsal and lateral views). Scale bars = 1.0 mm (except Fig 24 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310366" httpUri="https://zenodo.org/record/5310366/files/figure.png" pageId="1" pageNumber="756">25</figureCitation>
). ..................................... 3
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<keyLead id="30C39598FFE02464FFD4BE250E3DF9D7" box="[113,1088,1605,1629]" pageId="1" pageNumber="756">
Apical antennomere rather truncate at apex (
<figureCitation id="13092A28FFE02464FDD7BE2508BCF9D7" box="[626,705,1605,1628]" captionStart="Figs 2228. 2224" captionStartId="10.[151,188,1465,1484]" captionTargetBox="[174,1105,223,1419]" captionTargetId="figure-10@10.[148,1129,217,1454]" captionTargetPageId="10" captionText="Figs 2228. 2224 Gonioctena (A.) jindrai sp. nov.: 22 antenna; 23 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view); 24 spermatheca. 2526 G. (A.) mausonensis Cho &amp; Borowiec, 2016 (after CHO &amp; BOROWIEC 2016): 25 antenna; 26 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view). 2728 G. (A.) truncaticornis sp. nov.: 27 antenna; 28 aedeagus, holotype (dorsal and lateral views). Scale bars = 1.0 mm (except Fig 24 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310366" httpUri="https://zenodo.org/record/5310366/files/figure.png" pageId="1" pageNumber="756">Fig. 27</figureCitation>
). ...
<emphasis id="B946EABFFFE02464FD52BE250E3DF9D7" bold="true" box="[759,1088,1605,1629]" pageId="1" pageNumber="756">
<taxonomicName id="4C324D2EFFE02464FD52BE250998F9D6" authority="Cho &amp; Borowiec, 2016" authorityName="Cho &amp; Borowiec" authorityYear="2016" box="[759,997,1605,1629]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="756" phylum="Arthropoda" rank="species" species="truncaticornis" status="sp. nov." subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE02464FD52BE250977F9D6" bold="true" box="[759,778,1605,1629]" italics="true" pageId="1" pageNumber="756">G</emphasis>
. (
<emphasis id="B946EABFFFE02464FC84BE250944F9D6" bold="true" box="[801,825,1605,1629]" italics="true" pageId="1" pageNumber="756">A.</emphasis>
)
<emphasis id="B946EABFFFE02464FCECBE250998F9D6" bold="true" box="[841,997,1605,1629]" italics="true" pageId="1" pageNumber="756">truncaticornis</emphasis>
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3 Aedeagus strongly narrowed before short apical process (
<figureCitation id="13092A28FFE32467FC84B8BC0908FF78" box="[801,885,220,243]" captionStart="Figs 1321. 1318" captionStartId="9.[113,150,1493,1512]" captionTargetBox="[147,1065,226,1404]" captionTargetId="figure-14@9.[111,1092,216,1418]" captionTargetPageId="9" captionText="Figs 1321. 1318 Gonioctena (A.) eburoides (Achard, 1924): 13 antenna; 14 aedeagus (dorsal and lateral views); 1517 variation of apex of aedeagus (1516 Sa Pa; 17 Tam Dao); 18 spermatheca. 1921 G. (A.) flavoplagiata (Jacoby, 1890): 19 antenna; 20 aedeagus, paralectotype (dorsal view, detail of apex in dorsal view and lateral view); 21 spermatheca. Scale bars = 1.0 mm (except Figs 18, 21 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310364" httpUri="https://zenodo.org/record/5310364/files/figure.png" pageId="2" pageNumber="757">Figs 20</figureCitation>
,
<figureCitation id="13092A28FFE32467FC25B8BC09E3FF78" box="[896,926,220,243]" captionStart="Figs 2228. 2224" captionStartId="10.[151,188,1465,1484]" captionTargetBox="[174,1105,223,1419]" captionTargetId="figure-10@10.[148,1129,217,1454]" captionTargetPageId="10" captionText="Figs 2228. 2224 Gonioctena (A.) jindrai sp. nov.: 22 antenna; 23 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view); 24 spermatheca. 2526 G. (A.) mausonensis Cho &amp; Borowiec, 2016 (after CHO &amp; BOROWIEC 2016): 25 antenna; 26 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view). 2728 G. (A.) truncaticornis sp. nov.: 27 antenna; 28 aedeagus, holotype (dorsal and lateral views). Scale bars = 1.0 mm (except Fig 24 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310366" httpUri="https://zenodo.org/record/5310366/files/figure.png" pageId="2" pageNumber="757">23</figureCitation>
). ........................ 4
</keyLead>
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<paragraph id="8B8D36ADFFE32467FF32B89C0E1BFEB8" blockId="2.[151,1126,220,403]" pageId="2" pageNumber="757">
<keyLead id="30C39598FFE32467FF32B89C0E1BFEB8" pageId="2" pageNumber="757">
Aedeagus moderately narrowed from apical 1/3 to apex (
<figureCitation id="13092A28FFE32467FCB9B89C09E4FE98" box="[796,921,252,275]" captionStart="Figs 1321. 1318" captionStartId="9.[113,150,1493,1512]" captionTargetBox="[147,1065,226,1404]" captionTargetId="figure-14@9.[111,1092,216,1418]" captionTargetPageId="9" captionText="Figs 1321. 1318 Gonioctena (A.) eburoides (Achard, 1924): 13 antenna; 14 aedeagus (dorsal and lateral views); 1517 variation of apex of aedeagus (1516 Sa Pa; 17 Tam Dao); 18 spermatheca. 1921 G. (A.) flavoplagiata (Jacoby, 1890): 19 antenna; 20 aedeagus, paralectotype (dorsal view, detail of apex in dorsal view and lateral view); 21 spermatheca. Scale bars = 1.0 mm (except Figs 18, 21 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310364" httpUri="https://zenodo.org/record/5310364/files/figure.png" pageId="2" pageNumber="757">Figs 1417</figureCitation>
). ........................... .....................................................................................
<taxonomicName id="4C324D2EFFE32467FCA5B97B0E1BFEB8" authority="(Achard, 1924)" baseAuthorityName="Achard" baseAuthorityYear="1924" box="[768,1126,283,307]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="757" phylum="Arthropoda" rank="species" species="eburoides" subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE32467FCA5B97B09C0FEB8" bold="true" box="[768,957,283,307]" pageId="2" pageNumber="757">
<emphasis id="B946EABFFFE32467FCA5B97B096EFEB8" bold="true" box="[768,787,283,307]" italics="true" pageId="2" pageNumber="757">G</emphasis>
. (
<emphasis id="B946EABFFFE32467FC8FB97B093FFEB8" bold="true" box="[810,834,283,307]" italics="true" pageId="2" pageNumber="757">A.</emphasis>
)
<emphasis id="B946EABFFFE32467FCF7B97B09C0FEB8" bold="true" box="[850,957,283,307]" italics="true" pageId="2" pageNumber="757">eburoides</emphasis>
</emphasis>
(
<bibRefCitation id="EFA34B5CFFE32467FC6FB97C0E22FEB8" author="Achard" box="[970,1119,284,307]" firstAuthor="Achard" pageId="2" pageNumber="757" pagination="31 - 37" refId="ref6901" refString="ACHARD J. 1924: Les Phytodecta et leurs variations. Especes et varietes nouvelles. Rectifications synonymiques. Casopis Ceskoslovenske Spolecnosti Entomologicke 21: 31 - 37." type="journal article" year="1924">Achard, 1924</bibRefCitation>
)
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</paragraph>
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<keyStep id="30C62E08FFE32467FF32B95C0E1BFE18" pageId="2" pageNumber="757">
<paragraph id="8B8D36ADFFE32467FF32B95C0E1BFED8" blockId="2.[151,1126,220,403]" box="[151,1126,315,339]" pageId="2" pageNumber="757">
<keyLead id="30C39598FFE32467FF32B95C0E1BFED8" box="[151,1126,315,339]" pageId="2" pageNumber="757">
4 Apical process of aedeagus thin and elongate-oval (
<figureCitation id="13092A28FFE32467FD41B95C0949FED8" box="[740,820,316,339]" captionStart="Figs 2228. 2224" captionStartId="10.[151,188,1465,1484]" captionTargetBox="[174,1105,223,1419]" captionTargetId="figure-10@10.[148,1129,217,1454]" captionTargetPageId="10" captionText="Figs 2228. 2224 Gonioctena (A.) jindrai sp. nov.: 22 antenna; 23 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view); 24 spermatheca. 2526 G. (A.) mausonensis Cho &amp; Borowiec, 2016 (after CHO &amp; BOROWIEC 2016): 25 antenna; 26 aedeagus, holotype (dorsal view, detail of apex in dorsal view and lateral view). 2728 G. (A.) truncaticornis sp. nov.: 27 antenna; 28 aedeagus, holotype (dorsal and lateral views). Scale bars = 1.0 mm (except Fig 24 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310366" httpUri="https://zenodo.org/record/5310366/files/figure.png" pageId="2" pageNumber="757">Fig. 23</figureCitation>
). ....
<emphasis id="B946EABFFFE32467FCCBB95B0E1BFED8" bold="true" box="[878,1126,315,339]" pageId="2" pageNumber="757">
<taxonomicName id="4C324D2EFFE32467FCCBB95B0E77FED8" authority="Cho &amp; Borowiec, 2016" authorityName="Cho &amp; Borowiec" authorityYear="2016" box="[878,1034,315,339]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="757" phylum="Arthropoda" rank="species" species="jindrai" status="sp. nov." subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE32467FCCBB95B09FCFED8" bold="true" box="[878,897,315,339]" italics="true" pageId="2" pageNumber="757">G</emphasis>
. (
<emphasis id="B946EABFFFE32467FC3DB95B09CDFED8" bold="true" box="[920,944,315,339]" italics="true" pageId="2" pageNumber="757">A.</emphasis>
)
<emphasis id="B946EABFFFE32467FC65B95B0E77FED8" bold="true" box="[960,1034,315,339]" italics="true" pageId="2" pageNumber="757">jindrai</emphasis>
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<taxonomicNameLabel id="A27557C4FFE32467FBB4B95C0E1BFED8" box="[1041,1126,316,339]" pageId="2" pageNumber="757" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8B8D36ADFFE32467FF32B93C0E1BFE18" blockId="2.[151,1126,220,403]" pageId="2" pageNumber="757">
<keyLead id="30C39598FFE32467FF32B93C0E1BFE18" pageId="2" pageNumber="757">
Apical process of aedeagus very thin and parallel-sided (
<figureCitation id="13092A28FFE32467FCBFB93C0917FEF8" box="[794,874,348,371]" captionStart="Figs 1321. 1318" captionStartId="9.[113,150,1493,1512]" captionTargetBox="[147,1065,226,1404]" captionTargetId="figure-14@9.[111,1092,216,1418]" captionTargetPageId="9" captionText="Figs 1321. 1318 Gonioctena (A.) eburoides (Achard, 1924): 13 antenna; 14 aedeagus (dorsal and lateral views); 1517 variation of apex of aedeagus (1516 Sa Pa; 17 Tam Dao); 18 spermatheca. 1921 G. (A.) flavoplagiata (Jacoby, 1890): 19 antenna; 20 aedeagus, paralectotype (dorsal view, detail of apex in dorsal view and lateral view); 21 spermatheca. Scale bars = 1.0 mm (except Figs 18, 21 = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5310364" httpUri="https://zenodo.org/record/5310364/files/figure.png" pageId="2" pageNumber="757">Fig. 20</figureCitation>
). .................................. ................................................................................
<taxonomicName id="4C324D2EFFE32467FD46B91B0E1BFE18" authority="(Jacoby, 1890)" baseAuthorityName="Jacoby" baseAuthorityYear="1890" box="[739,1126,379,403]" class="Insecta" family="Chrysomelidae" genus="Gonioctena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="757" phylum="Arthropoda" rank="species" species="flavoplagiata" subGenus="Asiphytodecta">
<emphasis id="B946EABFFFE32467FD46B91B09BFFE18" bold="true" box="[739,962,379,403]" pageId="2" pageNumber="757">
<emphasis id="B946EABFFFE32467FD46B91B088BFE18" bold="true" box="[739,758,379,403]" italics="true" pageId="2" pageNumber="757">G</emphasis>
. (
<emphasis id="B946EABFFFE32467FCA9B91B0959FE18" bold="true" box="[780,804,379,403]" italics="true" pageId="2" pageNumber="757">A.</emphasis>
)
<emphasis id="B946EABFFFE32467FC91B91B09BFFE18" bold="true" box="[820,962,379,403]" italics="true" pageId="2" pageNumber="757">flavoplagiata</emphasis>
</emphasis>
(
<bibRefCitation id="EFA34B5CFFE32467FC75B91C0E22FE18" author="Jacoby" box="[976,1119,380,403]" firstAuthor="Jacoby" pageId="2" pageNumber="757" pagination="114 - 118" refId="ref7327" refString="JACOBY M. 1890: Descriptions of new species of phytophagous Coleoptera received by Mr. J. H. Leech, from Chang-Yang, China. Entomologist 23: 114 - 118." type="journal article" year="1890">Jacoby, 1890</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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</subSubSection>
</treatment>
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