385 lines
47 KiB
XML
385 lines
47 KiB
XML
<document id="3994FCD853ECF900C376570999E227BB" ID="10.11646/zootaxa.4238.4.5SLASH10410" ID-CLB-Dataset="36329" ID-DOI="10.11646/zootaxa.4238.4.5" ID-GBIF-Dataset="e17e2c89-bf5a-4747-8af1-3ab31d497bd1" ID-ISSN="1175-5326" ID-Zenodo-Dep="375507" ID-ZooBank="8070D6DA-F5DB-4612-A768-2A61D56A2B70" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488876041372" checkinUser="plazi" docAuthor="Quah, Evan S. H., Grismer, L. Lee, Wood, Perry L., Thura, Myint Kyaw, Zin, Thaw, Kyaw, Htet, Lwin, Ngwe, Grismer, Marta S. & Murdoch, Matthew L." docDate="2017" docId="039AE337E475415B16A5FEFDFDBD35D0" docLanguage="en" docName="zootaxa.4238.4.5SLASH10410.pdf" docOrigin="Zootaxa 4238 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyiophis salweenensis Quah, Grismer, Wood, Thura, Zin, Kyaw, Lwin, Grismer & Murdoch, 2017, sp. nov." docType="treatment" docVersion="10" lastPageNumber="575" masterDocId="FFA39B4FE476415F1632FF99FFFC3348" masterDocTitle="A new species of Mud Snake (Serpentes, Homalopsidae, Gyiophis Murphy & Voris, 2014) from Myanmar with a first molecular phylogenetic assessment of the genus" masterLastPageNumber="582" masterPageNumber="571" originalUpdateDomain="Boston" originalUpdateTime="1544887201233" originalUpdateUser="plazi" pageNumber="574" updateTime="1698437695048" updateUser="ExternalLinkService">
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<mods:title id="D4889E2181878119F52C69D3166D9559">A new species of Mud Snake (Serpentes, Homalopsidae, Gyiophis Murphy & Voris, 2014) from Myanmar with a first molecular phylogenetic assessment of the genus</mods:title>
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<mods:namePart id="8C7FB8A868A3884CAEF21715F1130031">Quah, Evan S. H.</mods:namePart>
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<mods:namePart id="E0417EBB3A884902E8CDA895294C6263">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="3843DC3864990744367CE1247DC593D6">Wood, Perry L.</mods:namePart>
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<mods:roleTerm id="1D66BE68ECBFE412974DF83B22273046">Author</mods:roleTerm>
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<mods:namePart id="34A859A369B85253AAF4FEBB6C141BFD">Thura, Myint Kyaw</mods:namePart>
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<mods:roleTerm id="E4BDE35EE63C582EE61228B76361964C">Author</mods:roleTerm>
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<mods:namePart id="76F1E8F29051E7814B3049EA7CD82852">Zin, Thaw</mods:namePart>
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<mods:roleTerm id="33715E6A0A65B2A8E3C11530C8C88A82">Author</mods:roleTerm>
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<mods:namePart id="2CB1DF59B9580A095204A24048D847CE">Kyaw, Htet</mods:namePart>
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<mods:namePart id="DB3C9C0EE04BCFEBC4F087B3B7296B64">Lwin, Ngwe</mods:namePart>
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<mods:namePart id="EC5B5C17C434E3C6D9F1210EF450AB7A">Grismer, Marta S.</mods:namePart>
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<mods:namePart id="7C00BB1639AC3BB2AE52BF56A2615266">Murdoch, Matthew L.</mods:namePart>
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<mods:typeOfResource id="86EE39B31ED01F531FEA696E94F6F5EC">text</mods:typeOfResource>
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<mods:title id="E6F4610DA5DC4519CE7F5E39C223ABE3">Zootaxa</mods:title>
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<mods:date id="095DE2863BB333EB726AEDB10634E135">2017</mods:date>
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<mods:number id="9A8A5D2E228005900ED9A175D532198F">4238</mods:number>
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<mods:detail id="37D1F63AD36125C49B107E13EA795828" type="issue">
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<mods:number id="C5184973F5F4A2384012CB7359033360">4</mods:number>
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<mods:start id="A553ADA2CAE7B0B63583CF41B91FEC66">571</mods:start>
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<mods:classification id="1B44E3A11CFFB897A77EECBF42869E48">journal article</mods:classification>
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<mods:identifier id="88CE7452B1F93308F53947BBBB978337" type="DOI">10.11646/zootaxa.4238.4.5</mods:identifier>
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<mods:identifier id="84C9DDB62396E7970414517DA4D3AFA7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D3438B3CC3FB95C3F894E3060769B3FE" type="Zenodo-Dep">375507</mods:identifier>
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<treatment id="039AE337E475415B16A5FEFDFDBD35D0" ID-DOI="http://doi.org/10.5281/zenodo.5684693" ID-GBIF-Taxon="127719449" ID-Zenodo-Dep="5684693" LSID="urn:lsid:plazi:treatment:039AE337E475415B16A5FEFDFDBD35D0" httpUri="http://treatment.plazi.org/id/039AE337E475415B16A5FEFDFDBD35D0" lastPageId="4" lastPageNumber="575" pageId="3" pageNumber="574">
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<subSubSection id="C32901AAE475415C16A5FEFDFEFB328C" pageId="3" pageNumber="574" type="nomenclature">
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<paragraph id="8B8C5221E475415C16A5FEFDFDF73237" blockId="3.[151,523,356,453]" box="[151,523,356,383]" pageId="3" pageNumber="574">
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<heading id="D0C4E54DE475415C16A5FEFDFDF73237" bold="true" box="[151,523,356,383]" fontSize="11" level="1" pageId="3" pageNumber="574" reason="1">
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<emphasis id="B9478E33E475415C16A5FEFDFDF73237" bold="true" box="[151,523,356,383]" pageId="3" pageNumber="574">
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<taxonomicName id="4C3329A2E475415C16A5FEFDFE5A3236" box="[151,422,356,382]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
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<emphasis id="B9478E33E475415C16A5FEFDFE5A3236" bold="true" box="[151,422,356,382]" italics="true" pageId="3" pageNumber="574">Gyiophis salweenensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2743348E475415C179FFEFCFDF73237" box="[429,523,357,383]" pageId="3" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8C5221E475415C16A5FE1EFEFB328C" blockId="3.[151,523,356,453]" pageId="3" pageNumber="574">
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Salween River Basin Mud Snake
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<figureCitation id="13084EA4E475415C16A5FE35FEFF328C" box="[151,259,428,453]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1957,1979]" captionStartId-1="6.[151,250,1832,1854]" captionStartId-2="7.[151,250,1793,1815]" captionTargetBox-0="[369,1207,193,1906]" captionTargetBox-1="[152,1435,193,1811]" captionTargetBox-2="[152,1435,193,1772]" captionTargetId-0="figure@5.[369,1209,193,1906]" captionTargetId-1="figure@6.[152,1435,193,1811]" captionTargetId-2="figure@7.[152,1435,193,1772]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Distribution of Gyiophis salweenensis sp. nov. as well as G. maculosa and G. vorisi based on Murphy (2007 b) in Myanmar." captionText-1="FIGURE 3. A: Holotype of Gyiophis salweenensis sp. nov. in life. B: Venter of the holotype of Gyiophis salweenensis sp. nov. C: Pattern on the dorsum of the holotype of Gyiophis salweenensis sp. nov. Photographs by Evan Quah." captionText-2="FIGURE 4. Position and number of supralabials and other head scales on the holotype of Gyiophis salweenensis sp. nov., A: The right side of the head. B: The left side of the head. AT = anterior temporal; F = frontal; IF = infralabial; IN = internasal; L = loreal; LMT = lower middle temporal; LPT = lower posterior temporal; M = mental; MPT = middle posterior temporal; N = nasal; P = parietal; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UMT = upper middle temporal; UPT = upper posterior temporal. Photographs by Evan Quah." httpUri-0="https://zenodo.org/record/375509/files/figure.png" httpUri-1="https://zenodo.org/record/375510/files/figure.png" httpUri-2="https://zenodo.org/record/375511/files/figure.png" pageId="3" pageNumber="574">Figs. 2–4</figureCitation>
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.
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C32901AAE475415C16A5FE6DFB12311D" pageId="3" pageNumber="574" type="materials_examined">
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<paragraph id="8B8C5221E475415C16A5FE6DFB12311D" blockId="3.[151,1437,500,813]" pageId="3" pageNumber="574">
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<materialsCitation id="3B5B587CE475415C16A5FE6DFB12311D" ID-GBIF-Occurrence="1438449034" collectingDate="2016-10-08" collectionCode="LSUHC" collectorName="Myint Kyaw Thura & Thaw Zin & Evan S. H. Quah & L. Lee Grismer & Perry L. Wood, Jr. & Marta S. Grismer & Matthew L. Murdoch & Htet Kyaw" country="Myanmar" elevation="44" latitude="16.373783" location="close to Sanpel Cave, Mawlamyine" longLatPrecision="1" longitude="97.77313" pageId="3" pageNumber="574" specimenCode="LSUHC 12960" specimenCount="1" stateProvince="Mon State" typeStatus="holotype">
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<emphasis id="B9478E33E475415C16A5FE6DFEF73144" bold="true" box="[151,267,500,524]" pageId="3" pageNumber="574">
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<typeStatus id="5488EC83E475415C16A5FE6DFEFA3144" box="[151,262,500,524]" pageId="3" pageNumber="574" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult
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<specimenCount id="9D3599A8E475415C176CFE6DFE563145" box="[350,426,500,525]" pageId="3" pageNumber="574" type="female">female</specimenCount>
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(
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<specimenCode id="DB95FA5AE475415C1789FE6DFD973145" box="[443,619,500,525]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="3" pageNumber="574">LSUHC 12960</specimenCode>
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) collected on
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<collectingDate id="EFC98D09E475415C1522FE6DFC383145" box="[784,964,500,525]" pageId="3" pageNumber="574" value="2016-10-08">
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<date id="FF8D74E1E475415C1522FE6DFC383145" box="[784,964,500,525]" pageId="3" pageNumber="574" value="2016-10-08">8 October 2016</date>
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</collectingDate>
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by
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<collectorName id="26C637F7E475415C15C1FE6DFB283145" box="[1011,1236,500,525]" pageId="3" pageNumber="574">Myint Kyaw Thura</collectorName>
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,
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<collectorName id="26C637F7E475415C12D0FE6DFAA93145" box="[1250,1365,500,525]" pageId="3" pageNumber="574">Thaw Zin</collectorName>
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,
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<collectorName id="26C637F7E475415C1351FE6CFEF33178" pageId="3" pageNumber="574">Evan S.H. Quah</collectorName>
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,
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||
<collectorName id="26C637F7E475415C1728FD81FE343178" box="[282,456,535,560]" pageId="3" pageNumber="574">L. Lee Grismer</collectorName>
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||
,
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||
<collectorName id="26C637F7E475415C17E1FD81FD5E3167" box="[467,674,535,560]" pageId="3" pageNumber="574">Perry L. Wood, Jr.</collectorName>
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||
,
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||
<collectorName id="26C637F7E475415C149CFD81FC8F3178" box="[686,883,535,560]" pageId="3" pageNumber="574">Marta S. Grismer</collectorName>
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||
,
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||
<collectorName id="26C637F7E475415C154CFD8EFB913178" box="[894,1133,535,560]" pageId="3" pageNumber="574">Matthew L. Murdoch</collectorName>
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||
and
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||
<collectorName id="26C637F7E475415C1296FD81FAE23178" box="[1188,1310,536,560]" pageId="3" pageNumber="574">Htet Kyaw</collectorName>
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||
from
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<location id="8EEC04FAE475415C1350FD8EFE19311D" LSID="urn:lsid:plazi:treatment:039AE337E475415B16A5FEFDFDBD35D0:8EEC04FAE475415C1350FD8EFE19311D" country="Myanmar" latitude="16.373783" longLatPrecision="1" longitude="97.77313" name="close to Sanpel Cave, Mawlamyine" pageId="3" pageNumber="574" stateProvince="Mon State">close to Sanpel Cave, Mawlamyine</location>
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,
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<collectingRegion id="49F79CC3E475415C17C2FDA4FD9B311C" box="[496,615,572,596]" pageId="3" pageNumber="574">Mon State</collectingRegion>
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,
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<collectingCountry id="F32412B1E475415C1443FDA4FD22311D" box="[625,734,573,597]" name="Myanmar" pageId="3" pageNumber="574">Myanmar</collectingCountry>
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(
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<geoCoordinate id="EE0734E6E475415C14DFFDA5FC86311D" box="[749,890,572,597]" direction="north" orientation="latitude" pageId="3" pageNumber="574" precision="1" value="16.373783">N16°22.427</geoCoordinate>
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,
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<geoCoordinate id="EE0734E6E475415C15B7FDA5FBF2311D" box="[901,1038,572,597]" direction="east" orientation="longitude" pageId="3" pageNumber="574" precision="1" value="97.77313">E97°46.388</geoCoordinate>
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;
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<elevation id="001EB512E475415C122BFDA5FBAE311C" box="[1049,1106,572,596]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="574" unit="m" value="44.0">
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<quantity id="4CCBFFC4E475415C122BFDA5FBAE311C" box="[1049,1106,572,596]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="574" unit="m" value="44.0">44 m</quantity>
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</elevation>
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in elevation).
|
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</materialsCitation>
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C32901AAE475415C16F5FDC6FEFF3064" pageId="3" pageNumber="574" type="diagnosis">
|
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<paragraph id="8B8C5221E475415C16F5FDC6FEFF3064" blockId="3.[151,1437,500,813]" pageId="3" pageNumber="574">
|
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<emphasis id="B9478E33E475415C16F5FDC6FEBD3130" bold="true" box="[199,321,607,632]" pageId="3" pageNumber="574">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C3329A2E475415C177AFDF9FDBB3130" box="[328,583,608,632]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
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||
<emphasis id="B9478E33E475415C177AFDF9FDBB3130" box="[328,583,608,632]" italics="true" pageId="3" pageNumber="574">Gyiophis salweenensis</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A2743348E475415C147CFDF9FD5B3130" box="[590,679,608,632]" pageId="3" pageNumber="574" rank="species">
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||
<emphasis id="B9478E33E475415C147CFDF9FD5B3130" bold="true" box="[590,679,608,632]" pageId="3" pageNumber="574">sp. nov.</emphasis>
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</taxonomicNameLabel>
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||
is separated from all congeners by having a unique combination of the following characters: a narrow rostral scale; the first three dorsal scale rows square; 129 (female) ventral scales; 30/29 (female) paired subcaudals; a divided cloacal plate; eight or nine supralabials; 10 infralabials; a maximum total length of
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||
<quantity id="4CCBFFC4E475415C170EFD55FE6131AC" box="[316,413,716,741]" metricMagnitude="-1" metricUnit="m" metricValue="4.16" pageId="3" pageNumber="574" unit="mm" value="416.0">416 mm</quantity>
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; relative tail length ratio of 0.13; a ventral patterning lacking a central spot on each ventral scale; the presence of a faint stripe on the lower, dorsal scale rows; and four rows of dark spots on the dorsum (
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<tableCitation id="C6B1679AE475415C16ACFC8DFF0E3064" box="[158,242,788,813]" captionStart="TABLE 1" captionStartId="3.[151,239,860,883]" captionTargetBox="[159,1386,942,1532]" captionTargetPageId="3" captionText="TABLE 1. Comparison of the scalation and colour pattern in G. maculosa, G. vorisi and G. salweenensis sp. nov. * data obtained from Murphy 2007 b, – no data available." httpUri="http://table.plazi.org/id/DF4C02A9E475415C16A5FCC5FD4C30DB" pageId="3" pageNumber="574" tableUuid="DF4C02A9E475415C16A5FCC5FD4C30DB">Table 1</tableCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4C02A9E475415C16A5FCC5FD4C30DB" ID-Table-UUID="DF4C02A9E475415C16A5FCC5FD4C30DB" httpUri="http://table.plazi.org/id/DF4C02A9E475415C16A5FCC5FD4C30DB" pageId="3" pageNumber="574" targetBox="[159,1386,942,1532]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="8B8C5221E475415C16A5FCC5FD4C30DB" blockId="3.[151,1436,860,915]" pageId="3" pageNumber="574">
|
||
<emphasis id="B9478E33E475415C16A5FCC5FEF7303B" bold="true" box="[151,267,860,883]" pageId="3" pageNumber="574">TABLE 1.</emphasis>
|
||
Comparison of the scalation and colour pattern in
|
||
<emphasis id="B9478E33E475415C1502FCC5FBE2303B" box="[816,1054,860,883]" italics="true" pageId="3" pageNumber="574">
|
||
<taxonomicName id="4C3329A2E475415C1502FCC5FC4B303B" box="[816,951,860,883]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="maculosa">G. maculosa</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3329A2E475415C15F1FCC5FBE2303B" box="[963,1054,860,883]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="vorisi">G. vorisi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3329A2E475415C1266FCC5FB01303B" box="[1108,1277,860,883]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E475415C1266FCC5FB01303B" box="[1108,1277,860,883]" italics="true" pageId="3" pageNumber="574">G. salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2743348E475415C1336FCC5FAA7303B" box="[1284,1371,860,883]" pageId="3" pageNumber="574" rank="species">
|
||
<emphasis id="B9478E33E475415C1336FCC5FAA7303B" bold="true" box="[1284,1371,860,883]" pageId="3" pageNumber="574">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
*data obtained from Murphy 2007b, – no data available.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C5221E475415C16ADFC37FB8736B4" pageId="3" pageNumber="574">
|
||
<table id="F933A081E475BEA016ADFC37FA9636B4" box="[159,1386,942,1532]" gridcols="4" gridrows="5" pageId="3" pageNumber="574">
|
||
<tr id="35035063E475BEA016ADFC37FA963744" box="[159,1386,942,1036]" gridrow="0" pageId="3" pageNumber="574">
|
||
<th id="76D2391FE475BEA016ADFC37FDCE3744" box="[159,562,942,1036]" gridcol="0" gridrow="0" pageId="3" pageNumber="574">Characteristics Scale rows on neck (ASR)</th>
|
||
<th id="76D2391FE475BEA01466FC37FCD53744" box="[596,809,942,1036]" gridcol="1" gridrow="0" pageId="3" pageNumber="574">
|
||
<taxonomicName id="4C3329A2E475415C1466FC37FCEB308C" box="[596,791,942,964]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B9478E33E475415C1466FC37FCEB308C" box="[596,791,942,964]" italics="true" pageId="3" pageNumber="574">Gyiophis maculosa</emphasis>
|
||
</taxonomicName>
|
||
* (N=1) 25
|
||
</th>
|
||
<th id="76D2391FE475BEA01560FC37FBFC3744" box="[850,1024,942,1036]" gridcol="2" gridrow="0" pageId="3" pageNumber="574">
|
||
<taxonomicName id="4C3329A2E475415C1561FC37FC12308D" box="[851,1006,942,965]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="vorisi">
|
||
<emphasis id="B9478E33E475415C1561FC37FC12308D" box="[851,1006,942,965]" italics="true" pageId="3" pageNumber="574">Gyiophis vorisi</emphasis>
|
||
</taxonomicName>
|
||
* (N=11) 26–28
|
||
</th>
|
||
<th id="76D2391FE475BEA0121CFC37FA963744" box="[1070,1386,942,1036]" gridcol="3" gridrow="0" pageId="3" pageNumber="574">
|
||
<taxonomicName id="4C3329A2E475415C121CFC37FAE8308C" box="[1070,1300,942,964]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="574" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E475415C121CFC37FAE8308C" box="[1070,1300,942,964]" italics="true" pageId="3" pageNumber="574">Gyiophis salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2743348E475415C1328FC36FA96308C" box="[1306,1386,943,964]" pageId="3" pageNumber="574" rank="species">
|
||
<emphasis id="B9478E33E475415C1328FC36FA96308C" bold="true" box="[1306,1386,943,964]" pageId="3" pageNumber="574">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(N=1) 26
|
||
</th>
|
||
</tr>
|
||
<tr id="35035063E475BEA016ADFBB9FA9637C0" box="[159,1386,1056,1160]" gridrow="1" pageId="3" pageNumber="574">
|
||
<th id="76D2391FE475BEA016ADFBB9FDCE37C0" box="[159,562,1056,1160]" gridcol="0" gridrow="1" pageId="3" pageNumber="574">Scale rows at mid-body (MSR) Scale rows near vent (PSR) Scale shape in first 3 rows</th>
|
||
<td id="76D2391FE475BEA01466FBB9FCD537C0" box="[596,809,1056,1160]" gridcol="1" gridrow="1" pageId="3" pageNumber="574">25 22 Ovate</td>
|
||
<td id="76D2391FE475BEA01560FBB9FBFC37C0" box="[850,1024,1056,1160]" gridcol="2" gridrow="1" pageId="3" pageNumber="574">25 21–23 Square</td>
|
||
<td id="76D2391FE475BEA0121CFBB9FA9637C0" box="[1070,1386,1056,1160]" gridcol="3" gridrow="1" pageId="3" pageNumber="574">25 20 Square</td>
|
||
</tr>
|
||
<tr id="35035063E475BEA016ADFB05FA96364C" box="[159,1386,1180,1284]" gridrow="2" pageId="3" pageNumber="574">
|
||
<th id="76D2391FE475BEA016ADFB05FDCE364C" box="[159,562,1180,1284]" gridcol="0" gridrow="2" pageId="3" pageNumber="574">Supralabials Ventral scales in males Ventral scales in females</th>
|
||
<td id="76D2391FE475BEA01466FB05FCD5364C" box="[596,809,1180,1284]" gridcol="1" gridrow="2" pageId="3" pageNumber="574">8 - 122</td>
|
||
<td id="76D2391FE475BEA01560FB05FBFC364C" box="[850,1024,1180,1284]" gridcol="2" gridrow="2" pageId="3" pageNumber="574">8 147– 152 142–148</td>
|
||
<td id="76D2391FE475BEA0121CFB05FA96364C" box="[1070,1386,1180,1284]" gridcol="3" gridrow="2" pageId="3" pageNumber="574">9/8 - 129</td>
|
||
</tr>
|
||
<tr id="35035063E475BEA016ADFA81FA9636C8" box="[159,1386,1304,1408]" gridrow="3" pageId="3" pageNumber="574">
|
||
<th id="76D2391FE475BEA016ADFA81FDCE36C8" box="[159,562,1304,1408]" gridcol="0" gridrow="3" pageId="3" pageNumber="574">Subcaudal scales in males Subcaudal scales in females Central spot on each ventral</th>
|
||
<td id="76D2391FE475BEA01466FA81FCD536C8" box="[596,809,1304,1408]" gridcol="1" gridrow="3" pageId="3" pageNumber="574">- 32/31 Present</td>
|
||
<td id="76D2391FE475BEA01560FA81FBFC36C8" box="[850,1024,1304,1408]" gridcol="2" gridrow="3" pageId="3" pageNumber="574">51–58 41–53 None</td>
|
||
<td id="76D2391FE475BEA0121CFA81FA9636C8" box="[1070,1386,1304,1408]" gridcol="3" gridrow="3" pageId="3" pageNumber="574">- 30/29 None</td>
|
||
</tr>
|
||
<tr id="35035063E475BEA016ADFA0DFA9636B4" box="[159,1386,1428,1532]" gridrow="4" pageId="3" pageNumber="574">
|
||
<th id="76D2391FE475BEA016ADFA0DFDCE36B4" box="[159,562,1428,1532]" gridcol="0" gridrow="4" pageId="3" pageNumber="574">Stripe on dorsal rows 2–4 Number of rows of spots on the dorsum Width of rostral scale</th>
|
||
<td id="76D2391FE475BEA01466FA0DFCD536B4" box="[596,809,1428,1532]" gridcol="1" gridrow="4" pageId="3" pageNumber="574">Absent 5–6 Narrow</td>
|
||
<td id="76D2391FE475BEA01560FA0DFBFC36B4" box="[850,1024,1428,1532]" gridcol="2" gridrow="4" pageId="3" pageNumber="574">Present 3 Broad</td>
|
||
<td id="76D2391FE475BEA0121CFA0DFA9636B4" box="[1070,1386,1428,1532]" gridcol="3" gridrow="4" pageId="3" pageNumber="574">Present (Faint) 4 Narrow</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="C32901AAE475415B16F5F9D8FAD13070" lastPageId="4" lastPageNumber="575" pageId="3" pageNumber="574" type="description">
|
||
<paragraph id="8B8C5221E475415B16F5F9D8FA86339D" blockId="3.[151,1436,1601,2022]" lastBlockId="4.[151,1437,151,1688]" lastPageId="4" lastPageNumber="575" pageId="3" pageNumber="574">
|
||
<emphasis id="B9478E33E475415C16F5F9D8FD8D3512" bold="true" box="[199,625,1601,1626]" pageId="3" pageNumber="574">
|
||
Description of
|
||
<typeStatus id="5488EC83E475415C1749F9D8FE1E3511" box="[379,482,1601,1625]" pageId="3" pageNumber="574" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13084EA4E475415C17C1F9D8FD983512" box="[499,612,1601,1626]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1832,1854]" captionStartId-1="7.[151,250,1793,1815]" captionTargetBox-0="[152,1435,193,1811]" captionTargetBox-1="[152,1435,193,1772]" captionTargetId-0="figure@6.[152,1435,193,1811]" captionTargetId-1="figure@7.[152,1435,193,1772]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 3. A: Holotype of Gyiophis salweenensis sp. nov. in life. B: Venter of the holotype of Gyiophis salweenensis sp. nov. C: Pattern on the dorsum of the holotype of Gyiophis salweenensis sp. nov. Photographs by Evan Quah." captionText-1="FIGURE 4. Position and number of supralabials and other head scales on the holotype of Gyiophis salweenensis sp. nov., A: The right side of the head. B: The left side of the head. AT = anterior temporal; F = frontal; IF = infralabial; IN = internasal; L = loreal; LMT = lower middle temporal; LPT = lower posterior temporal; M = mental; MPT = middle posterior temporal; N = nasal; P = parietal; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UMT = upper middle temporal; UPT = upper posterior temporal. Photographs by Evan Quah." httpUri-0="https://zenodo.org/record/375510/files/figure.png" httpUri-1="https://zenodo.org/record/375511/files/figure.png" pageId="3" pageNumber="574">Figs. 3–4</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Head depressed, distinct from neck; snout short and rounded, rostrum tapers downwards; rostral scale pentagonal, nearly as broad as tall, visible from above; eye moderate in size, pupils rounded; nasals semi-divided with nasal groove contacting first supralabial, nare in center (on left side of snout is a small, aberrant, triangular scale positioned between rostral, nasal and first supralabial); internasal small, quadrangular, not in contact with loreal; loreal 1/1, quadrangular, contacting first three supralabials on right side and supralabials 1, 2 and 4 on left side; preocular 1/1; supraocular 1/1; postoculars 2/2; prefrontals 2, in broad contact with each other, frontal, internasal, loreals, preoculars, supraoculars, and posterior tip of nasals; frontal pentagonal, 1.4 times longer than supraocular; parietals elongate; temporal scale formula 1 + 2 + 3; supralabials 9/ 8, largest supralabial 7/6, smallest 3/8, supralabial entering orbit 5/4 (on left side of head there is fragmentation of supralabial scales with third supralabial being small and sandwiched beneath second and fourth supralabials (
|
||
<figureCitation id="13084EA4E475415C135DF81CFF5B348A" captionStart="FIGURE 4" captionStartId="7.[151,250,1793,1815]" captionTargetBox="[152,1435,193,1772]" captionTargetId="figure@7.[152,1435,193,1772]" captionTargetPageId="7" captionText="FIGURE 4. Position and number of supralabials and other head scales on the holotype of Gyiophis salweenensis sp. nov., A: The right side of the head. B: The left side of the head. AT = anterior temporal; F = frontal; IF = infralabial; IN = internasal; L = loreal; LMT = lower middle temporal; LPT = lower posterior temporal; M = mental; MPT = middle posterior temporal; N = nasal; P = parietal; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UMT = upper middle temporal; UPT = upper posterior temporal. Photographs by Evan Quah." httpUri="https://zenodo.org/record/375511/files/figure.png" pageId="3" pageNumber="574">Fig. 4</figureCitation>
|
||
b); infralabials 10/10, seventh elongate, first four in contact with anterior chin shield; anterior pair of chin shields largest and rounded, second pair small; 26 ASR, 25 MSR, 20 PSR; scales of first three rows of dorsal scales square; 129 ventrals; cloacal plate divided; subcaudal scales 30/29. Body short, somewhat stocky; tail short; SVL
|
||
<quantity id="4CCBFFC4E472415B130AFF0EFA6433E7" box="[1336,1432,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="3.6399999999999997" pageId="4" pageNumber="575" unit="mm" value="364.0">364 mm</quantity>
|
||
; TaL
|
||
<quantity id="4CCBFFC4E472415B16F9FF25FEEA339D" box="[203,278,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="575" unit="mm" value="52.0">52mm</quantity>
|
||
; TL
|
||
<quantity id="4CCBFFC4E472415B1778FF25FE5F339D" box="[330,419,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="4.16" pageId="4" pageNumber="575" unit="mm" value="416.0">416mm</quantity>
|
||
. There is a puncture wound on the right side of the dorsum at the position of ventral 63.
|
||
</paragraph>
|
||
<paragraph id="8B8C5221E472415B16F5FF46FAD13070" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="575">
|
||
<emphasis id="B9478E33E472415B16F5FF46FDC933B0" bold="true" box="[199,565,223,248]" pageId="4" pageNumber="575">
|
||
Colouration in life (
|
||
<figureCitation id="13084EA4E472415B1785FF46FDDB33B0" box="[439,551,223,248]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1957,1979]" captionStartId-1="6.[151,250,1832,1854]" captionStartId-2="7.[151,250,1793,1815]" captionTargetBox-0="[369,1207,193,1906]" captionTargetBox-1="[152,1435,193,1811]" captionTargetBox-2="[152,1435,193,1772]" captionTargetId-0="figure@5.[369,1209,193,1906]" captionTargetId-1="figure@6.[152,1435,193,1811]" captionTargetId-2="figure@7.[152,1435,193,1772]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Distribution of Gyiophis salweenensis sp. nov. as well as G. maculosa and G. vorisi based on Murphy (2007 b) in Myanmar." captionText-1="FIGURE 3. A: Holotype of Gyiophis salweenensis sp. nov. in life. B: Venter of the holotype of Gyiophis salweenensis sp. nov. C: Pattern on the dorsum of the holotype of Gyiophis salweenensis sp. nov. Photographs by Evan Quah." captionText-2="FIGURE 4. Position and number of supralabials and other head scales on the holotype of Gyiophis salweenensis sp. nov., A: The right side of the head. B: The left side of the head. AT = anterior temporal; F = frontal; IF = infralabial; IN = internasal; L = loreal; LMT = lower middle temporal; LPT = lower posterior temporal; M = mental; MPT = middle posterior temporal; N = nasal; P = parietal; PF = prefrontal; PO = postocular; PrO = preocular; R = rostral; SL = supralabial; SO = supraocular; UMT = upper middle temporal; UPT = upper posterior temporal. Photographs by Evan Quah." httpUri-0="https://zenodo.org/record/375509/files/figure.png" httpUri-1="https://zenodo.org/record/375510/files/figure.png" httpUri-2="https://zenodo.org/record/375511/files/figure.png" pageId="4" pageNumber="575">Figs. 2–4</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The colour of each ventral and subcaudal scale is uniform cream with dark edging on the anterior right and left corners of each scale. These markings merge with the dark spot on the anterior half of each scale of the first dorsal scale row and some on the second row of dorsal scales to form a zig-zag stripe along the dorsoventral edge that runs the length of the body and tail. There are no central spots on the ventral scales except for ventrals 21, 54 and 55 that are extensions of the dark edgings of the corners of the scales. The chin and throat are cream and the infralabials, chin shields, and scales on the throat are edged in dark-grey. The posterior portion of dorsal scale rows one, two, three, and four form a faint cream stripe that is most prominent on the anterior portion on the body near the jaw and neck. The ground colour of the top of the head and the dorsum is greyish-brown. On dorsal scale rows 5–7 there are a series of large black spots 3–7 scales wide along the flanks and the anterior portion of the body near the neck. Some of these spots merge to form an irregularly shaped stripe. On the back are two rows of smaller black spots approximately 2–4 scales in width and usually on dorsal scale rows 11–14 and rows 15–17. Some single, darker scales are interspersed along the body on dorsal scale rows nine or 10. The tail is greyish brown with black spots. The head is speckled with small dark-grey spots with some larger spots; one on the frontal; another beneath the eye on the lower postocular and supralabials 4/5 and six. There is a broad dark-coloured streak across the anterior temporal, lower middle temporal, and the seventh and eighth supralabials; one at the corner of the jaw; and another two-scale wide streak on the crown from the posterior tip of the parietals extending posteriorly the length of five dorsal scales. The supralabials are cream and edged in dark-grey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32901AAE472415B16F5FCDAFCB33645" pageId="4" pageNumber="575" type="distribution">
|
||
<paragraph id="8B8C5221E472415B16F5FCDAFE3D30EC" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="575">
|
||
<emphasis id="B9478E33E472415B16F5FCDAFE423014" bold="true" box="[199,446,835,860]" pageId="4" pageNumber="575">
|
||
Distribution (
|
||
<figureCitation id="13084EA4E472415B175BFCDAFE533014" box="[361,431,835,860]" captionStart="FIGURE 2" captionStartId="5.[151,250,1957,1979]" captionTargetBox="[369,1207,193,1906]" captionTargetId="figure@5.[369,1209,193,1906]" captionTargetPageId="5" captionText="FIGURE 2. Distribution of Gyiophis salweenensis sp. nov. as well as G. maculosa and G. vorisi based on Murphy (2007 b) in Myanmar." httpUri="https://zenodo.org/record/375509/files/figure.png" pageId="4" pageNumber="575">Fig. 2</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<taxonomicName id="4C3329A2E472415B17F7FCDDFD383014" box="[453,708,836,860]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E472415B17F7FCDDFD383014" box="[453,708,836,860]" italics="true" pageId="4" pageNumber="575">Gyiophis salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2743348E472415B14FEFCDDFCDA3014" box="[716,806,836,860]" pageId="4" pageNumber="575" rank="species">
|
||
<emphasis id="B9478E33E472415B14FEFCDDFCDA3014" bold="true" box="[716,806,836,860]" pageId="4" pageNumber="575">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is only known from the
|
||
<typeStatus id="5488EC83E472415B120FFCDCFB913015" box="[1085,1133,837,861]" pageId="4" pageNumber="575">type</typeStatus>
|
||
locality near Sanpel Cave,
|
||
<collectingRegion id="49F79CC3E472415B16A5FCF1FF303037" box="[151,204,872,895]" country="Myanmar" name="Mon" pageId="4" pageNumber="575">Mon</collectingRegion>
|
||
State,
|
||
<collectingCountry id="F32412B1E472415B1728FCF1FE7530C8" box="[282,393,872,896]" name="Myanmar" pageId="4" pageNumber="575">Myanmar</collectingCountry>
|
||
. It is expected to be wider ranging throughout the Salween River Basin and found wherever appropriate habitat occurs.
|
||
</paragraph>
|
||
<paragraph id="8B8C5221E472415B16F5FC29FCB33645" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="575">
|
||
<emphasis id="B9478E33E472415B16F5FC29FE163080" bold="true" box="[199,490,943,968]" pageId="4" pageNumber="575">
|
||
Natural history (
|
||
<figureCitation id="13084EA4E472415B17A1FC36FE203080" box="[403,476,943,968]" captionStart="FIGURE 5" captionStartId="8.[151,250,911,933]" captionTargetBox="[151,1436,285,890]" captionTargetId="figure@8.[151,1436,193,890]" captionTargetPageId="8" captionText="FIGURE 5. Flooded grassland habitat near where Gyiophis salweenensis sp. nov. was found. Photograph by Evan Quah." httpUri="https://zenodo.org/record/375512/files/figure.png" pageId="4" pageNumber="575">Fig. 5</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="5488EC83E472415B141AFC36FD763080" box="[552,650,943,968]" pageId="4" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
was found at approximately 1930 hours crossing a narrow dirt road between flooded fields that we presume to be its natural habitat. The weather was overcast and it rained later that night. Many other species of homalopsids are semiaquatic and commonly found in streams, rivers, ponds, and flooded rice fields (
|
||
<bibRefCitation id="EFA22FD0E472415B174AFB85FDDA377C" author="Murphy" box="[376,550,1052,1077]" pageId="4" pageNumber="575" refString="Murphy, J. C. (2007 a) Homalopsid snakes, evolution in the mud. Krieger Publishing, Malabar, 260 pp." type="book" year="2007" yearSuffix="a">Murphy 2007a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA22FD0E472415B1401FB85FCDC377C" author="Stuebing" box="[563,800,1052,1077]" pageId="4" pageNumber="575" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo (second edition). Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="B9478E33E472415B14ADFB84FD2E377C" box="[671,722,1052,1076]" italics="true" pageId="4" pageNumber="575">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C3329A2E472415B1506FB85FBC9377C" box="[820,1077,1052,1076]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E472415B1506FB85FBC9377C" box="[820,1077,1052,1076]" italics="true" pageId="4" pageNumber="575">Gyiophis salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2743348E472415B120CFB85FB66377C" box="[1086,1178,1052,1076]" pageId="4" pageNumber="575" rank="species">
|
||
<emphasis id="B9478E33E472415B120CFB85FB66377C" bold="true" box="[1086,1178,1052,1076]" pageId="4" pageNumber="575">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is expected to share a similar life history and the valvular nostrils located dorsally on the snout indicate this species probably spends a large part of its life in the water. Homalopsid snakes such as
|
||
<emphasis id="B9478E33E472415B1564FBFDFBD73734" box="[854,1067,1124,1148]" italics="true" pageId="4" pageNumber="575">
|
||
<taxonomicName id="4C3329A2E472415B1564FBFDFBDB3734" box="[854,1063,1124,1148]" class="Reptilia" family="Homalopsidae" genus="Enhydris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="enhydris">Enhydris enhydris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C3329A2E472415B1206FBFDFF2437E8" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Reptilia" family="Homalopsidae" genus="Homalopsis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="buccata">
|
||
<emphasis id="B9478E33E472415B1206FBFDFAE03734" box="[1076,1308,1124,1148]" italics="true" pageId="4" pageNumber="575">Homalopsis buccata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA22FD0E472415B131FFBFDFF2C37E8" author="Linnaeus" pageId="4" pageNumber="575" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata, Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3329A2E472415B1724FB11FDEC37E8" box="[278,528,1160,1184]" class="Reptilia" family="Homalopsidae" genus="Hypsiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="plumbea">
|
||
<emphasis id="B9478E33E472415B1724FB11FDEC37E8" box="[278,528,1160,1184]" italics="true" pageId="4" pageNumber="575">Hypsiscopus plumbea</emphasis>
|
||
</taxonomicName>
|
||
have been observed crawling on land and crossing roads during wet weather (
|
||
<bibRefCitation id="EFA22FD0E472415B16ACFB35FE7A378D" author="Voris" box="[158,390,1196,1221]" pageId="4" pageNumber="575" refString="Voris, H. K. & Karns, D. R. (1996) Habitat utilization, movements, and activity patterns of Enhydris plumbea (Serpentes: Homalopsinae) in a rice paddy wetland in Borneo, Herpetological Natural History, 4, 111 - 126." type="journal article" year="1996">Voris & Karns 1996</bibRefCitation>
|
||
; Lim & D’Rozario 2009; EQSH personal obs.) and the
|
||
<typeStatus id="5488EC83E472415B15CDFB35FB9D378D" box="[1023,1121,1196,1221]" pageId="4" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C3329A2E472415B12BAFB35FAC5378C" box="[1160,1337,1196,1220]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E472415B12BAFB35FAC5378C" box="[1160,1337,1196,1220]" italics="true" pageId="4" pageNumber="575">G. salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9478E33E472415B1373FB35FA60378C" bold="true" box="[1345,1436,1196,1220]" pageId="4" pageNumber="575">
|
||
<taxonomicNameLabel id="A2743348E472415B1373FB35FA60378C" box="[1345,1436,1196,1220]" pageId="4" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
could have been dispersing to a new area as well. The
|
||
<typeStatus id="5488EC83E472415B14C4FB56FCA437A0" box="[758,856,1231,1256]" pageId="4" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
also had a puncture wound on its back which might have come from an encounter with a predator such as a heron.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32901AAE472415B16F5FA81FDBD35D0" pageId="4" pageNumber="575" type="etymology">
|
||
<paragraph id="8B8C5221E472415B16F5FA81FE6136D4" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="575">
|
||
<emphasis id="B9478E33E472415B16F5FA81FEB23678" bold="true" box="[199,334,1304,1328]" pageId="4" pageNumber="575">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C3329A2E472415B140DFA81FD2D3678" box="[575,721,1304,1328]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="salweenensis">
|
||
<emphasis id="B9478E33E472415B140DFA81FD2D3678" box="[575,721,1304,1328]" italics="true" pageId="4" pageNumber="575">salweenensis</emphasis>
|
||
</taxonomicName>
|
||
is in reference to area where the
|
||
<typeStatus id="5488EC83E472415B1261FA8EFB493678" box="[1107,1205,1303,1328]" pageId="4" pageNumber="575" type="holotype">holotype</typeStatus>
|
||
was found which is close to the vicinity of the Salween River near the city of Mawlamyine. The suffix
|
||
<emphasis id="B9478E33E472415B125CFAA4FB5A361C" box="[1134,1190,1341,1364]" italics="true" pageId="4" pageNumber="575">ensis</emphasis>
|
||
is a Latin derivation meaning “from” or “inhabiting.” It renders the specific epithet an adjective that must be in grammatical accord with the gender of
|
||
<taxonomicName id="4C3329A2E472415B1703FA1DFE6A36D4" box="[305,406,1412,1436]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9478E33E472415B1703FA1DFE6A36D4" box="[305,406,1412,1436]" italics="true" pageId="4" pageNumber="575">Gyiophis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8C5221E472415B16F5FA3EFDBD35D0" blockId="4.[151,1437,151,1688]" pageId="4" pageNumber="575">
|
||
<emphasis id="B9478E33E472415B16F5FA3EFE9D3688" bold="true" box="[199,353,1447,1472]" pageId="4" pageNumber="575">Comparison.</emphasis>
|
||
<taxonomicName id="4C3329A2E472415B1758FA31FD963688" box="[362,618,1448,1472]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="salweenensis" status="sp. nov.">
|
||
<emphasis id="B9478E33E472415B1758FA31FD963688" box="[362,618,1448,1472]" italics="true" pageId="4" pageNumber="575">Gyiophis salweenensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2743348E472415B1440FA31FD313688" box="[626,717,1448,1472]" pageId="4" pageNumber="575" rank="species">
|
||
<emphasis id="B9478E33E472415B1440FA31FD313688" bold="true" box="[626,717,1448,1472]" pageId="4" pageNumber="575">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguishable from
|
||
<taxonomicName id="4C3329A2E472415B15D0FA31FB963688" box="[994,1130,1448,1472]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="maculosa">
|
||
<emphasis id="B9478E33E472415B15D0FA31FB963688" box="[994,1130,1448,1472]" italics="true" pageId="4" pageNumber="575">G. maculosa</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the dorsal scales of first three rows (square vs. ovate), the ventral scale pattern (absence of a central spot on each ventral scale vs. its presence), and a stripe running through the scales of the lower dorsal scale row (faint one vs. absent). It is further distinguished from
|
||
<taxonomicName id="4C3329A2E472415B17F0F98DFDDC3564" box="[450,544,1556,1580]" class="Reptilia" family="Homalopsidae" genus="Gyiophis" higherTaxonomySource="CoL" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="575" phylum="Chordata" rank="species" species="vorisi">
|
||
<emphasis id="B9478E33E472415B17F0F98DFDDC3564" box="[450,544,1556,1580]" italics="true" pageId="4" pageNumber="575">G. vorisi</emphasis>
|
||
</taxonomicName>
|
||
by its lower number of ventrals (129 vs. 142–152), lower number of subcaudals (30/29 vs. 41–58), shape of the rostral scale (narrow vs. broad), and the number of rows of spots on the dorsum (four vs. three) (
|
||
<tableCitation id="C6B1679AE472415B1761F9C5FE54353C" box="[339,424,1628,1653]" captionStart="TABLE 1" captionStartId="3.[151,239,860,883]" captionTargetBox="[159,1386,942,1532]" captionTargetPageId="3" captionText="TABLE 1. Comparison of the scalation and colour pattern in G. maculosa, G. vorisi and G. salweenensis sp. nov. * data obtained from Murphy 2007 b, – no data available." httpUri="http://table.plazi.org/id/DF4C02A9E475415C16A5FCC5FD4C30DB" pageId="4" pageNumber="575" tableUuid="DF4C02A9E475415C16A5FCC5FD4C30DB">Table 1</tableCitation>
|
||
). It differs from the other species of homalopsids found in
|
||
<collectingCountry id="F32412B1E472415B1264F9C4FB3F353D" box="[1110,1219,1629,1653]" name="Myanmar" pageId="4" pageNumber="575">Myanmar</collectingCountry>
|
||
by unique suite of characters presented in the key below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |