435 lines
50 KiB
XML
435 lines
50 KiB
XML
<document id="52D13C5D38A627AB457CF0BB05C072AE" ID-DOI="10.5281/zenodo.193804" ID-GBIF-Dataset="844d15b3-9a8e-4b1e-b667-8c0cdf9ee6f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="193804" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460173208284" checkinUser="plazi" docAuthor="Huang, Di-Ying & Nel, Andre" docDate="2010" docId="0397D95FFFF3867750F3FE0A998C0FA9" docLanguage="en" docName="zt02387p062.pdf" docOrigin="Zootaxa 2387" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Protoliupanshania wangi Huang & Nel, 2010, gen. nov." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="FFAEA127FFF286735064FF97986E0B0B" masterDocTitle="Protoliupanshania wangi, a new genus and species from the Chinese Early Cretaceous (Odonata: Aeshnoptera: Liupanshaniidae)" masterLastPageNumber="62" masterPageNumber="57" pageNumber="58" updateTime="1698236004121" updateUser="plazi">
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<mods:mods id="9E0F6CB56060B21FCB5931946B475479" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A65E97F034E7DC40834F5FA0225C052A">
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<mods:title id="AA983492B46A7AB6C43FC0FDC1E8E153">Protoliupanshania wangi, a new genus and species from the Chinese Early Cretaceous (Odonata: Aeshnoptera: Liupanshaniidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2108EDB9B3812B7578E24E04B5B1A3EE" type="personal">
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<mods:role id="064BE5125372D91587E1BD990CD626B8">
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<mods:roleTerm id="E55F7669847E2237111D7F2B043DAA0B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6C49561AC810FB64EF2E7ED995B182F2">Huang, Di-Ying</mods:namePart>
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</mods:name>
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<mods:name id="B1E0FBF93A0254741154EC761B48647E" type="personal">
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<mods:role id="8A9D2C89E3590484CF4EE24F86405799">
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<mods:roleTerm id="6B861A827C2CC1E74DD2B4F3BF89DAE0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="63940B110D9775B567A0E963878E9239">Nel, Andre</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A5279F4F57ECCCD054744B6B9203643B">text</mods:typeOfResource>
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<mods:relatedItem id="F267E5CE7C7F8296743C6203A9C4E194" type="host">
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<mods:titleInfo id="00AA1F422DCD83853A608BFC60105901">
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<mods:title id="903AB1A893CDB091320F635C29AB088F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="87121D6C25B0BDF379C5A22CD3A7732A">
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<mods:date id="D671E871F56062A8E18EA794E8EEFFBF">2010</mods:date>
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<mods:detail id="54585AF94C80CFF70CB7E88EF8CDAD71" type="volume">
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<mods:number id="FB529AD205C332A394F40B8CAE07D69C">2387</mods:number>
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</mods:detail>
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<mods:extent id="341C2952C0D4D1A7321F2ADD4D46B2B9" unit="page">
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<mods:start id="E60508EFEA2E38A95723BDA280E48554">57</mods:start>
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<mods:end id="0C2BD78F938EBE7689BAF3A74F6C2E99">62</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="76E0FADA44F8A237F805E477B9402049">journal article</mods:classification>
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<mods:identifier id="31D994254161EC3E742FCE72C1499002" type="DOI">10.5281/zenodo.193804</mods:identifier>
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<mods:identifier id="C10851C4357F397E0B95C6BF93AE1E2D" type="GBIF-Dataset">844d15b3-9a8e-4b1e-b667-8c0cdf9ee6f9</mods:identifier>
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<mods:identifier id="07295BDAEA589B31E439A76D14D70F12" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="326E8AF36084BE23E86CBD0D6259A1C5" type="Zenodo-Dep">193804</mods:identifier>
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</mods:mods>
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<treatment id="0397D95FFFF3867750F3FE0A998C0FA9" ID-DOI="http://doi.org/10.5281/zenodo.6196497" ID-GBIF-Taxon="119383501" ID-Zenodo-Dep="6196497" LSID="urn:lsid:plazi:treatment:0397D95FFFF3867750F3FE0A998C0FA9" httpUri="http://treatment.plazi.org/id/0397D95FFFF3867750F3FE0A998C0FA9" lastPageId="4" lastPageNumber="61" pageId="1" pageNumber="58">
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<subSubSection id="C3243BC2FFF3867250F3FE0A99790AD1" pageId="1" pageNumber="58" type="nomenclature">
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<paragraph id="8B816849FFF3867250F3FE0A9AC50ABC" blockId="1.[151,683,413,474]" box="[151,683,413,439]" pageId="1" pageNumber="58">
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<heading id="D0C9DF25FFF3867250F3FE0A9AC50ABC" bold="true" box="[151,683,413,439]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
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<emphasis id="B94AB45BFFF3867250F3FE0A9AC50ABC" bold="true" box="[151,683,413,439]" pageId="1" pageNumber="58">
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<taxonomicName id="4C3E13CAFFF3867250F3FE0A99A40ABC" box="[151,458,413,439]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="wangi" status="gen. nov.">
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<emphasis id="B94AB45BFFF3867250F3FE0A99A40ABC" bold="true" box="[151,458,413,439]" italics="true" pageId="1" pageNumber="58">Protoliupanshania wangi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2790920FFF3867251B5FE0A9A2E0ABC" box="[465,576,413,439]" pageId="1" pageNumber="58" rank="genus">gen. nov.</taxonomicNameLabel>
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, sp. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B816849FFF3867250F3FE5799790AD1" blockId="1.[151,683,413,474]" box="[151,279,448,474]" pageId="1" pageNumber="58">
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(
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<figureCitation id="130574CCFFF3867250FBFE5799600AD1" box="[159,270,448,474]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 3 – 6" captionStartId-0="2.[151,255,1917,1941]" captionStartId-1="3.[151,275,1592,1616]" captionTargetBox-0="[295,1274,216,1897]" captionTargetBox-1="[384,1195,206,1548]" captionTargetId-0="figure@2.[273,1288,197,1902]" captionTargetId-1="figure@3.[366,1220,185,1571]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Protoliupanshania wangi gen. nov., sp. nov., holotype NIGP 148199, photograph (scale bar 5 mm). FIGURE 2. Protoliupanshania wangi gen. nov., sp. nov., paratype specimen NIGP 148200, photograph (scale bar 5 mm)." captionText-1="FIGURES 3 – 6. Protoliupanshania wangi gen. nov., sp. nov., 3. holotype NIGP 148199, drawing. 4 – 6. Paratype specimen NIGP 148200. 4, left hind wing. 5, left forewing. 6, right forewing, standard abbreviations: AA anal vein, Ax 1 Ax 2 primary antenodal cross-veins, CuAa distal branch of cubitus anterior, CuAb proximal branch of cubitus anterior, IRi intercalary radial veins, MAa distal branch of median anterior, MAb posterior branch of median anterior, MP median posterior, N nodus, ‘ O’ lestine oblique vein, PsA pseudo-anal vein, Pt pterostigma, RA radius anterior, RP radius posterior, tp trigonal planate, T discoidal triangle, al anal loop (scale bars 5 mm)." httpUri-0="https://zenodo.org/record/193805/files/figure.png" httpUri-1="https://zenodo.org/record/193806/files/figure.png" pageId="1" pageNumber="58">Figs. 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3243BC2FFF3867250F3FD9D9D60092F" box="[151,1294,522,548]" pageId="1" pageNumber="58" type="description">
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<paragraph id="8B816849FFF3867250F3FD9D9D60092F" blockId="1.[151,1437,522,2004]" box="[151,1294,522,548]" pageId="1" pageNumber="58">
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<emphasis id="B94AB45BFFF3867250F3FD9D99E6092F" bold="true" box="[151,392,522,548]" pageId="1" pageNumber="58">Derivation of name</emphasis>
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. Named for Mr. Wang Xiang-Dong, who collected and gave us the
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<typeStatus id="5485D6EBFFF3867254C4FD9D9D64092F" box="[1184,1290,522,548]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3243BC2FFF3867250A0FDB8988B09D4" pageId="1" pageNumber="58" type="materials_examined">
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<paragraph id="8B816849FFF3867250A0FDB89D60099F" blockId="1.[151,1437,522,2004]" pageId="1" pageNumber="58">
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<emphasis id="B94AB45BFFF3867250A0FDB8995D0942" bold="true" box="[196,307,559,585]" pageId="1" pageNumber="58">Material</emphasis>
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.
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<typeStatus id="5485D6EBFFF386725127FDA799DC0941" box="[323,434,560,586]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED2FF08CFFF3867251DFFDA79A6F0941" box="[443,513,560,586]" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="58">NIGP</collectionCode>
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148199, a print of a well-preserved complete hind wing;
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<typeStatus id="5485D6EBFFF3867254D8FDA79D4D0941" box="[1212,1315,560,586]" pageId="1" pageNumber="58" type="paratype">paratype</typeStatus>
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specimen
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<collectionCode id="ED2FF08CFFF3867250F3FDC298B20964" box="[151,220,597,623]" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="58">NIGP</collectionCode>
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148200, a poorly-preserved specimen with part and counterpart with head trunk and four wings.
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<typeStatus id="5485D6EBFFF386725532FDC29DF50964" box="[1366,1435,597,623]" pageId="1" pageNumber="58">Types</typeStatus>
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housed in Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences,
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<collectingCountry id="F32928D9FFF3867254A4FDED9D64099F" box="[1216,1290,634,660]" name="China" pageId="1" pageNumber="58">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B816849FFF3867250A0FD08988B09D4" blockId="1.[151,1437,522,2004]" pageId="1" pageNumber="58">
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<emphasis id="B94AB45BFFF3867250A0FD08990B09B2" bold="true" box="[196,357,671,697]" pageId="1" pageNumber="58">
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<typeStatus id="5485D6EBFFF3867250A0FD08996C09B2" box="[196,258,671,697]" pageId="1" pageNumber="58">Type</typeStatus>
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locality
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</emphasis>
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. Lower Cretaceous Yixian Formation (ca. 125 Ma); Beipiao City, Liaoning Province, NE
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<collectingCountry id="F32928D9FFF3867250F3FD52988F09D4" box="[151,225,709,735]" name="China" pageId="1" pageNumber="58">China</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3243BC2FFF3867250A0FD7D9A71080F" box="[196,543,746,772]" pageId="1" pageNumber="58" type="diagnosis">
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<paragraph id="8B816849FFF3867250A0FD7D9A71080F" blockId="1.[151,1437,522,2004]" box="[196,543,746,772]" pageId="1" pageNumber="58">
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<emphasis id="B94AB45BFFF3867250A0FD7D9952080F" bold="true" box="[196,316,746,772]" pageId="1" pageNumber="58">Diagnosis</emphasis>
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. That of the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3243BC2FFF3867250A0FC9899250CDF" pageId="1" pageNumber="58" type="description">
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<paragraph id="8B816849FFF3867250A0FC989D7C0D54" blockId="1.[151,1437,522,2004]" pageId="1" pageNumber="58">
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<emphasis id="B94AB45BFFF3867250A0FC98993B0822" bold="true" box="[196,341,783,809]" pageId="1" pageNumber="58">Description</emphasis>
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.
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<typeStatus id="5485D6EBFFF386725100FC8799B80821" box="[356,470,784,810]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
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: print of a complete hind wing,
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<quantity id="4CC6C5ACFFF386725329FC879BD80821" box="[845,950,784,810]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="1" pageNumber="58" unit="mm" value="45.5">45.5 mm</quantity>
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long, wing
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<quantity id="4CC6C5ACFFF386725426FC879CC50821" box="[1090,1195,784,810]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="1" pageNumber="58" unit="mm" value="16.1">16.1 mm</quantity>
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wide; distance from base to arculus 5.0 mm; from arculus to
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<geoCoordinate id="EE0A0E8EFFF38672520AFCA29AD30844" box="[622,701,821,847]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="5555" value="15.1">
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N
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<quantity id="4CC6C5ACFFF3867252EEFCA29AD30844" box="[650,701,821,847]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="1" pageNumber="58" unit="mm" value="15.1">15.1</quantity>
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</geoCoordinate>
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mm; from arculus to base of
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<collectingCountry id="F32928D9FFF386725472FCA29C550844" box="[1046,1083,821,847]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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3/4
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<quantity id="4CC6C5ACFFF386725402FCA29CAD0844" box="[1126,1219,821,847]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="58" unit="mm" value="9.2">9.2 mm</quantity>
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; from N to Pt
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<quantity id="4CC6C5ACFFF38672550DFCA298A6087F" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="1" pageNumber="58" unit="mm" value="14.6">14.6 mm</quantity>
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, from N to wing apex
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<quantity id="4CC6C5ACFFF3867251B2FCCD9A2D087F" box="[470,579,858,884]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="1" pageNumber="58" unit="mm" value="25.7">25.7 mm</quantity>
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, from Pt to wing apex
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<quantity id="4CC6C5ACFFF386725330FCCD9BDD087F" box="[852,947,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="58" unit="mm" value="6.4">6.4 mm</quantity>
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; Pt
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<quantity id="4CC6C5ACFFF386725384FCCD9C55087F" box="[992,1083,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="58" unit="mm" value="5.8">5.8 mm</quantity>
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long,
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<quantity id="4CC6C5ACFFF3867254ECFCCD9C8D087F" box="[1160,1251,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="58" unit="mm" value="1.1">1.1 mm</quantity>
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wide, covering five cells; pterostigmal brace oblique and aligned with basal side of Pt; no secondary antenodal cross-vein basal to Ax1, two antenodal cross-veins of first row and one of second row between Ax1 and Ax2, six of first row and six of second row distal of Ax2; distance from wing base to Ax1
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||
<quantity id="4CC6C5ACFFF38672539BFC5D9C3208EF" box="[1023,1116,970,996]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="58" unit="mm" value="3.9">3.9 mm</quantity>
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, between Ax1 and Ax2
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<quantity id="4CC6C5ACFFF38672551CFC5D98A60F01" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="58" unit="mm" value="6.1">6.1 mm</quantity>
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; Ax2 aligned with apex of T;
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<collectingCountry id="F32928D9FFF38672524FFC679A210F01" box="[555,591,1008,1034]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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||
and MA strongly separated at arculus; hypertriangle
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<quantity id="4CC6C5ACFFF3867254ACFC679D4D0F01" box="[1224,1315,1008,1034]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="58" unit="mm" value="5.3">5.3 mm</quantity>
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||
long, free of cross-veins; T
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||
<quantity id="4CC6C5ACFFF38672510DFB8299AA0F24" box="[361,452,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="58" unit="mm" value="1.8">1.8 mm</quantity>
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||
distal of arculus, free of cross-veins, broad, with basal side
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||
<quantity id="4CC6C5ACFFF3867254F3FB829C9C0F24" box="[1175,1266,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="58" unit="mm" value="1.9">1.9 mm</quantity>
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long, anterior side
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||
<quantity id="4CC6C5ACFFF3867250A9FBAD99460F5F" box="[205,296,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="58" unit="mm" value="3.3">3.3 mm</quantity>
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||
long, and MAb
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||
<quantity id="4CC6C5ACFFF38672518DFBAD9A2D0F5F" box="[489,579,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="58" unit="mm" value="3.7">3.7 mm</quantity>
|
||
long; a distinct angle in MAb at base of convex tp; part of MAb distal to tp long,
|
||
<quantity id="4CC6C5ACFFF38672509DFBF799390F71" box="[249,343,1120,1146]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="58" unit="mm" value="2.8">2.8 mm</quantity>
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||
; postdiscoidal area with two rows of cells; a strongly developed convex tp, long and slightly zigzagged; no Mspl; basal part of area between RA and
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||
<collectingCountry id="F32928D9FFF386725341FB129B270F94" box="[805,841,1157,1183]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
with six cross-veins basal to
|
||
<collectingCountry id="F32928D9FFF3867254FBFB129CA80F94" box="[1183,1222,1157,1183]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
3/4, and five distal of base of
|
||
<collectingCountry id="F32928D9FFF386725177FB3D99560FCF" box="[275,312,1194,1220]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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||
3/4 and basal of subnodus; area between
|
||
<collectingCountry id="F32928D9FFF38672537DFB3D9B530FCF" box="[793,829,1194,1220]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
and MAa with five cross-veins basal to
|
||
<collectingCountry id="F32928D9FFF38672557DFB3D9D2E0FCF" box="[1305,1344,1194,1220]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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||
3/4; one visible oblique vein ‘O’ three cells distal to base of
|
||
<collectingCountry id="F32928D9FFF386725365FB479B470FE1" box="[769,809,1232,1258]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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||
2; one row of cells between IR2 and
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||
<collectingCountry id="F32928D9FFF3867254BBFB479D6B0FE1" box="[1247,1285,1232,1258]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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3/4 two cells distal to subnodus; MAa and
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<collectingCountry id="F32928D9FFF386725190FB629A740E04" box="[500,538,1269,1295]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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||
3/4 parallel, nearly straight; one row of cells between
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<collectingCountry id="F32928D9FFF3867254F3FB629CD30E04" box="[1175,1213,1269,1295]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
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3/4 and MAa basal of level of vein ‘O’, two rows distally; one row of cells between MP and CuAa, with no narrowing of area between these veins opposite base of
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<collectingCountry id="F32928D9FFF38672523DFAD79AEF0E51" box="[601,641,1344,1370]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
3/4; median and submedian area, and subdiscoidal space free; CuP and PsA strongly curved; posterior wing margin rounded at base (indicating a female specimen); anal area long and broad, with three posterior branches of AA directed towards posterior wing margin, and 9-10 rows of cells between AA and posterior wing margin; al small, opened posteriorly with no strong cross-vein between AA and CuAb, composed of five cells; cubito-anal area broad, with seven rows of cells between CuAa and posterior wing margin; CuAa with six posterior branches; no Rspl, but four posterior longitudinal secondary veins between IR2 and posterior wing margin; base of
|
||
<collectingCountry id="F32928D9FFF38672537BF9B79B2A0D31" box="[799,836,1568,1594]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
2 aligned with subnodus; no well-defined IR1; one row of cells between
|
||
<collectingCountry id="F32928D9FFF3867251F5F9D299D40D54" box="[401,442,1605,1631]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
1 and
|
||
<collectingCountry id="F32928D9FFF386725199F9D29A4C0D54" box="[509,546,1605,1631]" name="Poland" pageId="1" pageNumber="58">RP</collectingCountry>
|
||
2 between subnodus and two cells basal of Pt, two rows distally.
|
||
</paragraph>
|
||
<paragraph id="8B816849FFF3867250A2F9FD99250CDF" blockId="1.[151,1437,522,2004]" pageId="1" pageNumber="58">
|
||
<typeStatus id="5485D6EBFFF3867250A2F9FD995F0D8F" box="[198,305,1642,1668]" pageId="1" pageNumber="58" type="paratype">Paratype</typeStatus>
|
||
: part and counterpart of a body with the four wings in connection. Wings and body are strongly deformed and their dimensions cannot be measured with accuracy. Eyes separated; abdomen with long cerci. All preserved structures of hind wings identical to those of
|
||
<typeStatus id="5485D6EBFFF386725323F9229BC30DC4" box="[839,941,1717,1743]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
hind wing, including absence of Mspl and Rspl, al posteriorly opened, shape of T and tp, etc. Fore wing venational details largely undecipherable or not visible: hypertriangle longer than T, free of cross-veins; T free of cross-veins, broad, equilateral; median, submedian, and subdiscoidal spaces free of cross-veins; PsA strong; postdiscoidal area with four rows of cells; a strong convex tp, long and slightly zigzagged; no Mspl; no Rspl; Pt elongate, covering four cells, pterostigmal brace oblique and aligned with basal side of Pt; two rows of cells in anal area; four rows of cells in cubito-anal area; at least one secondary antenodal cross-vein between Ax1 and Ax2; Ax2 nearly opposite distal side of T.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4138C1FFF0867150F3F8EA98BF0CDF" httpUri="https://zenodo.org/record/193805/files/figure.png" pageId="2" pageNumber="59" targetBox="[295,1274,216,1897]" targetPageId="2">
|
||
<paragraph id="8B816849FFF0867150F3F8EA98BF0CDF" blockId="2.[151,1436,1917,2004]" pageId="2" pageNumber="59">
|
||
<emphasis id="B94AB45BFFF0867150F3F8EA99770C9E" bold="true" box="[151,281,1917,1941]" pageId="2" pageNumber="59">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C3E13CAFFF086715144F8EA9A400C9F" box="[288,558,1917,1940]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="wangi" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF086715144F8EA9A400C9F" box="[288,558,1917,1940]" italics="true" pageId="2" pageNumber="59">Protoliupanshania wangi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2790920FFF086715251F8EA9AF70C9E" box="[565,665,1917,1941]" pageId="2" pageNumber="59" rank="genus">
|
||
<emphasis id="B94AB45BFFF086715251F8EA9AF70C9E" bold="true" box="[565,665,1917,1941]" pageId="2" pageNumber="59">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94AB45BFFF0867152C2F8EA9A920C9E" bold="true" box="[678,764,1917,1941]" pageId="2" pageNumber="59">sp. nov.</emphasis>
|
||
, holotype NIGP 148199, photograph (scale bar 5 mm).
|
||
<emphasis id="B94AB45BFFF0867150F3F80A99720CBE" bold="true" box="[151,284,1949,1973]" pageId="2" pageNumber="59">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3E13CAFFF086715140F80A9A540CBF" box="[292,570,1949,1972]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="wangi" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF086715140F80A9A540CBF" box="[292,570,1949,1972]" italics="true" pageId="2" pageNumber="59">Protoliupanshania wangi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2790920FFF086715225F80A9AC70CBE" box="[577,681,1949,1973]" pageId="2" pageNumber="59" rank="genus">
|
||
<emphasis id="B94AB45BFFF086715225F80A9AC70CBE" bold="true" box="[577,681,1949,1973]" pageId="2" pageNumber="59">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94AB45BFFF0867152D3F80A9B610CBE" bold="true" box="[695,783,1949,1973]" pageId="2" pageNumber="59">sp. nov.</emphasis>
|
||
, paratype specimen NIGP 148200, photograph (scale bar 5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4138C1FFF1867050F3F9AF9B980DE4" httpUri="https://zenodo.org/record/193806/files/figure.png" pageId="3" pageNumber="60" targetBox="[384,1195,206,1548]" targetPageId="3">
|
||
<paragraph id="8B816849FFF1867050F3F9AF9B980DE4" blockId="3.[151,1437,1592,1775]" pageId="3" pageNumber="60">
|
||
<emphasis id="B94AB45BFFF1867050F3F9AF99250D5B" bold="true" box="[151,331,1592,1616]" pageId="3" pageNumber="60">FIGURES 3–6.</emphasis>
|
||
<taxonomicName id="4C3E13CAFFF186705130F9AF9A1D0D44" box="[340,627,1592,1615]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wangi" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF186705130F9AF9A1D0D44" box="[340,627,1592,1615]" italics="true" pageId="3" pageNumber="60">Protoliupanshania wangi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2790920FFF18670521FF9AF9A880D5B" box="[635,742,1592,1616]" pageId="3" pageNumber="60" rank="genus">
|
||
<emphasis id="B94AB45BFFF18670521FF9AF9A880D5B" bold="true" box="[635,742,1592,1616]" pageId="3" pageNumber="60">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B94AB45BFFF186705291F9AF9B3F0D5B" bold="true" box="[757,849,1592,1616]" pageId="3" pageNumber="60">sp. nov.</emphasis>
|
||
, 3. holotype NIGP 148199, drawing. 4–6. Paratype specimen NIGP 148200. 4, left hind wing. 5, left forewing. 6, right forewing, standard abbreviations: AA anal vein, Ax1 Ax2 primary antenodal cross-veins, CuAa distal branch of cubitus anterior, CuAb proximal branch of cubitus anterior, IRi intercalary radial veins, MAa distal branch of median anterior, MAb posterior branch of median anterior, MP median posterior, N nodus, ‘O’ lestine oblique vein, PsA pseudo-anal vein, Pt pterostigma, RA radius anterior, RP radius posterior, tp trigonal planate, T discoidal triangle, al anal loop (scale bars 5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3243BC2FFF1867750A2F88A9C770859" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="discussion">
|
||
<paragraph id="8B816849FFF1867050A2F88A9B6D0CF9" blockId="3.[151,1436,1821,2034]" pageId="3" pageNumber="60">
|
||
<emphasis id="B94AB45BFFF1867050A2F88A993F0C3C" bold="true" box="[198,337,1821,1847]" pageId="3" pageNumber="60">Discussion.</emphasis>
|
||
<taxonomicName id="4C3E13CAFFF18670513AF8899AE30C3C" box="[350,653,1822,1847]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="wangi">
|
||
<emphasis id="B94AB45BFFF18670513AF8899AE30C3C" box="[350,653,1822,1847]" italics="true" pageId="3" pageNumber="60">Protoliupanshania wangi</emphasis>
|
||
</taxonomicName>
|
||
shares with Liupanshaniidae
|
||
<bibRefCitation id="EFAF15B8FFF18670539BF88A9C830C3C" author="Bechly" box="[1023,1261,1821,1847]" pageId="3" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF18670543AF8899CF00C3C" box="[1118,1182,1822,1847]" italics="true" pageId="3" pageNumber="60">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
the following characters: both pairs of wings with a strong convex secondary longitudinal vein (tp) in the postdiscoidal area originating from a strongly angulate MAb (convergent to several other groups of Aeshnoptera and to the clade Gomphides
|
||
<bibRefCitation id="EFAF15B8FFF186705141F81A9A660CAC" author="Bechly" box="[293,520,1933,1959]" pageId="3" pageNumber="62" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, Milano, (N. S.), 10, 1 - 68." type="journal article" year="1998">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF18670511BF81999D40CAC" box="[383,442,1934,1959]" italics="true" pageId="3" pageNumber="60">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
); unique shape of elongate and narrow hind wing T (anterior side of T distally curved and ending on anterior side MA of hypertriangle); forewing T broad and nearly equilateral; distal accessory oblique vein between
|
||
<collectingCountry id="F32928D9FFF186705276F84F9A590CF9" box="[530,567,2008,2034]" name="Poland" pageId="3" pageNumber="60">RP</collectingCountry>
|
||
2 and IR2 absent.
|
||
</paragraph>
|
||
<paragraph id="8B816849FFF6867750A2FF0F9B550979" blockId="4.[151,1437,152,1186]" pageId="4" pageNumber="61">
|
||
Similar shape tp and T also occur in the petalurid family Cretapetaluridae
|
||
<bibRefCitation id="EFAF15B8FFF6867754F5FF0F9D0C0BB9" author="Nel" box="[1169,1378,152,178]" pageId="4" pageNumber="62" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, Milano, (N. S.), 10, 1 - 68." type="journal article" year="1998">
|
||
Nel
|
||
<emphasis id="B94AB45BFFF6867754AFFF0F9D600BBA" box="[1227,1294,152,177]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
, but
|
||
<emphasis id="B94AB45BFFF6867750F3FF2A99520BDD" box="[151,316,189,214]" italics="true" pageId="4" pageNumber="61">Cretapetalura</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775127FF2A99940BDC" author="Nel" box="[323,506,189,215]" pageId="4" pageNumber="62" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, Milano, (N. S.), 10, 1 - 68." type="journal article" year="1998">
|
||
Nel
|
||
<emphasis id="B94AB45BFFF686775111FF2A99DE0BDD" box="[373,432,189,214]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
has its pterostigmal brace strongly recessed basally (
|
||
<bibRefCitation id="EFAF15B8FFF68677540DFF2A9D4B0BDC" author="Nel" box="[1129,1317,189,215]" pageId="4" pageNumber="62" refString="Nel, A., Bechly, G., Jarzembowski, E. A. & Martinez-Delclos, X. (1998) A revision of the fossil petalurid dragonflies (Insecta: Odonata: Anisoptera: Petalurida). Paleontologia Lombarda, Milano, (N. S.), 10, 1 - 68." type="journal article" year="1998">
|
||
Nel
|
||
<emphasis id="B94AB45BFFF6867754F8FF2A9CB90BDD" box="[1180,1239,189,214]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
). Also the aeshnopteran
|
||
<emphasis id="B94AB45BFFF686775125FF7599A40BF0" box="[321,458,226,251]" italics="true" pageId="4" pageNumber="61">Va l d a e s h n a</emphasis>
|
||
Jarzembowski, 1988 and the gomphid Hageniidae
|
||
<bibRefCitation id="EFAF15B8FFF686775421FF759D060BF7" author="Tillyard" box="[1093,1384,226,252]" pageId="4" pageNumber="62" refString="Tillyard, R. J. & Fraser, F. C. (1940) A reclassification of the order Odonata based on some new interpretations of the venation of the dragonfly wing. The Australian Zoologist, 9, 124 - 396." type="journal article" year="1940">Tillyard & Fraser, 1940</bibRefCitation>
|
||
and
|
||
<taxonomicName id="4C3E13CAFFF6867750F3FE9F9A0F0A29" authority="Jacobson & Bianchi, 1905" authorityName="Jacobson & Bianchi" authorityYear="1905" box="[151,609,264,290]" class="Insecta" family="Lindeniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="family">
|
||
Lindeniidae
|
||
<bibRefCitation id="EFAF15B8FFF68677514FFE9F9A0F0A29" author="Jacobson" box="[299,609,264,290]" pageId="4" pageNumber="62" refString="Jacobson, G. & Bianchi, V. (1905) Die Orthopteren und Pseudoneuropteren des Russischen Reiches und der angrenzenden Gebiete, pp. 635 - 846 (A. F. Dewrien, St. Petersburg). [In Russian]." type="book chapter" year="1905">Jacobson & Bianchi, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
(sensu Bechly 1991) have tp, but their T of both fore- and hind wings have the same elongate shape, unlike
|
||
<taxonomicName id="4C3E13CAFFF686775206FEBA9B2D0A4D" box="[610,835,301,326]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF686775206FEBA9B2D0A4D" box="[610,835,301,326]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B94AB45BFFF686775337FEBA9B980A4D" box="[851,1014,301,326]" italics="true" pageId="4" pageNumber="61">Enigmaeshna</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF68677539AFEBA9CD20A4C" author="Nel" box="[1022,1212,301,327]" pageId="4" pageNumber="62" refString="Nel, A., Neraudeau, D., Perrichot, V., Girard, V. & Gomez, B. (2008) A new dragonfly family in the Upper Cretaceous of France (Insecta: Odonata: Aeshnoptera). Acta Palaeontologica Polonica, 53, 165 - 168." type="journal article" year="2008">
|
||
Nel
|
||
<emphasis id="B94AB45BFFF686775456FEBA9C010A4D" box="[1074,1135,301,326]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
(Enigmaeshnidae, only hind wing known) has a strong tp but it differs from
|
||
<taxonomicName id="4C3E13CAFFF68677532BFEC59C430A60" box="[847,1069,338,363]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF68677532BFEC59C430A60" box="[847,1069,338,363]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
by the longer anal area with a posteriorly closed al, and hypertriangle, T and subdiscoidal space crossed.
|
||
<bibRefCitation id="EFAF15B8FFF686775474FEEF9CA10A99" author="Lin" box="[1040,1231,376,402]" pageId="4" pageNumber="62" refString="Lin, Q. - b., Zhang, S. & Huang, D. - y. (2004) Fuxiaeschna hsiufunia gen. nov., spec. nov., a new Lower Cretaceous dragonfly from Northwestern China (Aeshnoptera: Rudiaeschnidae). Odonatologica, 33, 437 - 442." type="journal article" year="2004">
|
||
Lin
|
||
<emphasis id="B94AB45BFFF686775425FEEF9C130A9A" box="[1089,1149,376,401]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
(2004
|
||
</bibRefCitation>
|
||
: Fig. 2) figured a zigzagged tp for the Chinese Cretaceous
|
||
<taxonomicName id="4C3E13CAFFF6867752E0FE0A9C3A0ABC" authority="Lin et al., 2004" authorityName="Lin et al." authorityYear="2004" box="[644,1108,413,439]" class="Insecta" family="Rudiaeschnidae" genus="Fuxiaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="hsuifunia">
|
||
<emphasis id="B94AB45BFFF6867752E0FE0A9BFD0ABD" box="[644,915,413,438]" italics="true" pageId="4" pageNumber="61">Fuxiaeschna hsuifunia</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF6867753FFFE0A9C3A0ABC" author="Lin" box="[923,1108,413,439]" pageId="4" pageNumber="62" refString="Lin, Q. - b., Zhang, S. & Huang, D. - y. (2004) Fuxiaeschna hsiufunia gen. nov., spec. nov., a new Lower Cretaceous dragonfly from Northwestern China (Aeshnoptera: Rudiaeschnidae). Odonatologica, 33, 437 - 442." type="journal article" year="2004">
|
||
Lin
|
||
<emphasis id="B94AB45BFFF6867753A9FE0A9C670ABD" box="[973,1033,413,438]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
which they included in the aeshnopteran family
|
||
<taxonomicName id="4C3E13CAFFF6867751F7FE559B2C0AD7" authority="Bechly et al., 2001" authorityName="Bechly et al." authorityYear="2001" box="[403,834,450,476]" class="Insecta" family="Rudiaeschnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="family">
|
||
Rudiaeschnidae
|
||
<bibRefCitation id="EFAF15B8FFF68677523DFE559B2C0AD7" author="Bechly" box="[601,834,450,476]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF6867752D1FE559A9C0AD0" box="[693,754,450,475]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, but this character is not shared by the
|
||
<typeStatus id="5485D6EBFFF68677557FFE559D210AD7" box="[1307,1359,450,476]" pageId="4" pageNumber="61">type</typeStatus>
|
||
genus
|
||
<emphasis id="B94AB45BFFF6867750F3FE7F995E090A" box="[151,304,488,513]" italics="true" pageId="4" pageNumber="61">Rudiaeschna</emphasis>
|
||
Ren & Guo, 1996.
|
||
<taxonomicName id="4C3E13CAFFF686775279FE7F9ADB090A" box="[541,693,488,513]" class="Insecta" family="Rudiaeschnidae" genus="Fuxiaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94AB45BFFF686775279FE7F9ADB090A" box="[541,693,488,513]" italics="true" pageId="4" pageNumber="61">Fuxiaeschna</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3E13CAFFF686775330FE7F9C5F090A" box="[852,1073,488,513]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF686775330FE7F9C5F090A" box="[852,1073,488,513]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a closed al and broader hind wing T. Lastly the representatives of the clade Neoaeshnida
|
||
<bibRefCitation id="EFAF15B8FFF686775482FD9A9DF2092C" author="Bechly" box="[1254,1436,525,551]" pageId="4" pageNumber="61" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der * Odonata. Petalura, Special Volume 2, 402 pp." type="book" year="1996">Bechly, 1996</bibRefCitation>
|
||
(
|
||
<taxonomicName id="4C3E13CAFFF6867750FBFDA599E90947" box="[159,391,562,588]" class="Insecta" family="Gomphaeschnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="family">Gomphaeschnidae</taxonomicName>
|
||
+ Aeshnodea) have developed tp but they have well defined Rspl and Mpsl, and a posteriorly closed hind wing al unlike
|
||
<taxonomicName id="4C3E13CAFFF68677523EFDCF9B5B097A" box="[602,821,600,625]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF68677523EFDCF9B5B097A" box="[602,821,600,625]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B816849FFF6867750A0FDEA9C770859" blockId="4.[151,1437,152,1186]" pageId="4" pageNumber="61">
|
||
Other liupanshaniid synapomorphies are less pronounced or absent in
|
||
<taxonomicName id="4C3E13CAFFF686775474FDEA9C80099D" box="[1040,1262,637,662]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF686775474FDEA9C80099D" box="[1040,1262,637,662]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
; its hind wing MAb is sigmoidally curved, but distinctly less so than in other Liupanshaniidae. The hind wing T of
|
||
<taxonomicName id="4C3E13CAFFF6867750F3FD5F991C09EA" box="[151,370,712,737]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF6867750F3FD5F991C09EA" box="[151,370,712,737]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
is free instead of being divided into several cells by parallel cross-veins; its forewing T is free instead of being divided into cells (forewing T only known in
|
||
<emphasis id="B94AB45BFFF6867753DDFD7A9CDE080D" box="[953,1200,749,774]" italics="true" pageId="4" pageNumber="61">Araripeliupanshania</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF6867754DCFD7A9DF2080C" author="Bechly" box="[1208,1436,749,775]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF686775570FD7A9D3F080D" box="[1300,1361,749,774]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
and
|
||
<emphasis id="B94AB45BFFF6867750AEFC8599C80820" box="[202,422,786,811]" italics="true" pageId="4" pageNumber="61">Paramesuropetala</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF6867751CAFC859AF90827" author="Bechly" box="[430,663,786,812]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF68677526FFC859A290820" box="[523,583,786,811]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
). Also the subdiscoidal space of
|
||
<taxonomicName id="4C3E13CAFFF686775446FC859D6E0820" box="[1058,1280,786,811]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF686775446FC859D6E0820" box="[1058,1280,786,811]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
is larger and broader than in other Liupanshaniidae (
|
||
<bibRefCitation id="EFAF15B8FFF686775200FCAF9B290859" author="Bechly" box="[612,839,824,850]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF6867752A4FCAF9A95085A" box="[704,763,824,849]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAF15B8FFF686775337FCAF9C630859" author="Lin" box="[851,1037,824,850]" pageId="4" pageNumber="62" refString="Lin, Q. - b., Nel, A. & Huang, D. - y. (2002) Phylogenetic analysis of the Mesozoic dragonfly family Liupanshaniidae (Insecta: Aeshnoptera: Odonata). Cretaceous Research, 23, 439 - 444." type="journal article" year="2002">
|
||
Lin
|
||
<emphasis id="B94AB45BFFF6867753E7FCAF9BD0085A" box="[899,958,824,849]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2002
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3243BC2FFF6867750A0FCCA998C0FA9" pageId="4" pageNumber="61" type="reference_group">
|
||
<paragraph id="8B816849FFF6867750A0FCCA998C0FA9" blockId="4.[151,1437,152,1186]" pageId="4" pageNumber="61">
|
||
<taxonomicName id="4C3E13CAFFF6867750A0FCCA99CF087D" box="[196,417,861,886]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF6867750A0FCCA99CF087D" box="[196,417,861,886]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
can be considered a member of the family Liupanshaniidae, but it could be the sister genus of all other taxa in this family, because of the plesiomorphic character state ‘tp basally zigzagged’ (hind wing tp not basally zigzagged in
|
||
<emphasis id="B94AB45BFFF68677524BFC3F9B4308CA" box="[559,813,936,961]" italics="true" pageId="4" pageNumber="61">Araripeliupanshania</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775352FC3F9C4A08C9" author="Bechly" box="[822,1060,936,962]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF6867753F1FC3F9BBD08CA" box="[917,979,936,961]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
,
|
||
<emphasis id="B94AB45BFFF686775455FC3F9D7B08CA" box="[1073,1301,936,961]" italics="true" pageId="4" pageNumber="61">Guyuanaeschnidia</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775579FC3F9DF608C9" author="Lin" box="[1309,1432,936,962]" pageId="4" pageNumber="62" refString="Lin, Q. - B. (1982) Insecta. In Palaeontological atlas of northwest China, Shaanxi, Gansu, Ningxi volume, part 3, Mesozoic and Cenozoic, pp. 70 - 83. (Xi'an Institute of Geology and Mineral Resources, Geological Publishing House, Beijing). [In Chinese]." type="journal article" year="1982">Lin, 1982</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C3E13CAFFF6867750F3FC5A99BF08EC" authority="Hong, 1982" authorityName="Hong" authorityYear="1982" box="[151,465,973,999]" class="Insecta" family="Liupanshaniidae" genus="Liupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94AB45BFFF6867750F3FC5A995408ED" box="[151,314,973,998]" italics="true" pageId="4" pageNumber="61">Liupanshania</emphasis>
|
||
Hong, 1982
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B94AB45BFFF686775274FC5A9A8B08ED" box="[528,741,973,998]" italics="true" pageId="4" pageNumber="61">Paraliupanshania</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF68677528AFC5A9BB808EC" author="Bechly" box="[750,982,973,999]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF68677532EFC5A9BE808ED" box="[842,902,973,998]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
); but some uncertainty remains due to the uncertain phylogenetic position of
|
||
<emphasis id="B94AB45BFFF6867752E0FC659B020F00" box="[644,876,1010,1035]" italics="true" pageId="4" pageNumber="61">Paramesuropetala</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775312FC659C000F07" author="Bechly" box="[886,1134,1010,1036]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF6867753BCFC659C770F00" box="[984,1049,1010,1035]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
. If the basal position of
|
||
<taxonomicName id="4C3E13CAFFF6867750F3FB8F99160F3A" box="[151,376,1048,1073]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF6867750F3FB8F99160F3A" box="[151,376,1048,1073]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
is confirmed after a better knowledge of
|
||
<emphasis id="B94AB45BFFF686775314FB8F9C200F3A" box="[880,1102,1048,1073]" italics="true" pageId="4" pageNumber="61">Paramesuropetala</emphasis>
|
||
, this might suggest Central Asia as an origin for this group. The absence of Rspl and Mspl in
|
||
<taxonomicName id="4C3E13CAFFF686775380FBAA9CA10F5D" box="[996,1231,1085,1110]" class="Insecta" family="Liupanshaniidae" genus="Protoliupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="57" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B94AB45BFFF686775380FBAA9CA10F5D" box="[996,1231,1085,1110]" italics="true" pageId="4" pageNumber="61">Protoliupanshania</emphasis>
|
||
</taxonomicName>
|
||
is also found in
|
||
<emphasis id="B94AB45BFFF6867750F3FBF499040F77" box="[151,362,1123,1148]" italics="true" pageId="4" pageNumber="61">Guyuanaeschidia</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775116FBF599850F77" author="Lin" box="[370,491,1122,1148]" pageId="4" pageNumber="62" refString="Lin, Q. - B. (1982) Insecta. In Palaeontological atlas of northwest China, Shaanxi, Gansu, Ningxi volume, part 3, Mesozoic and Cenozoic, pp. 70 - 83. (Xi'an Institute of Geology and Mineral Resources, Geological Publishing House, Beijing). [In Chinese]." type="journal article" year="1982">Lin, 1982</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C3E13CAFFF68677519CFBF49B560F77" authority="Hong, 1982" authorityName="Hong" authorityYear="1982" box="[504,824,1122,1148]" class="Insecta" family="Liupanshaniidae" genus="Liupanshania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94AB45BFFF68677519CFBF49AF00F77" box="[504,670,1123,1148]" italics="true" pageId="4" pageNumber="61">Liupanshania</emphasis>
|
||
Hong, 1982
|
||
</taxonomicName>
|
||
, while
|
||
<emphasis id="B94AB45BFFF6867753F5FBF49C050F77" box="[913,1131,1123,1148]" italics="true" pageId="4" pageNumber="61">Paraliupanshania</emphasis>
|
||
<bibRefCitation id="EFAF15B8FFF686775417FBF59D350F77" author="Bechly" box="[1139,1371,1122,1148]" pageId="4" pageNumber="61" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D., Fleck, G., Escuillie, F., Wisshak, M. M. & Maisch, M. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of several new families, genera and species (Insecta: Odonata: Anisoptera). Neue Palaontologische Abhandlungen, 4, 219 pp." type="book" year="2001">
|
||
Bechly
|
||
<emphasis id="B94AB45BFFF6867754B5FBF49D610F77" box="[1233,1295,1123,1148]" italics="true" pageId="4" pageNumber="61">et al.</emphasis>
|
||
, 2001
|
||
</bibRefCitation>
|
||
have well defined Rspl and Mspl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |