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<document ID-DOI="10.37520/aemnp.2023.003" ID-ISSN="1804-6487" ID-Zenodo-Dep="7887332" ID-ZooBank="937DF12A-BC2F-4406-802F-63838312B0C1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683048435184" checkinUser="felipe" docAuthor="Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long &amp; Hendrich, Lars" docDate="2023" docId="03975B72FFC7760C9682CD0D5126FA2C" docLanguage="en" docName="ActaEntMusNatPra.63.1.75-102.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.003" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Cybister fumatus Sharp 1882" docType="treatment" docVersion="2" lastPageNumber="80" masterDocId="FFAE230AFFC376099545CC10505AFFDA" masterDocTitle="Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae)" masterLastPageNumber="102" masterPageNumber="75" pageNumber="79" updateTime="1683048975263" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Zhuo-Yin</mods:namePart>
<mods:affiliation>Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou 510275, China; e-mail: zhuoyin _ jiang @ 126. com; ORCID ID https: // orcid. org / 0000 - 0001 - 6825 - 4283</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Shuang</mods:namePart>
<mods:affiliation>Guangzhou Customs Technology Center, Tianhe District, Guangzhou, China; e-mail: topzs @ 163. com;</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mai, Zu-Qi</mods:namePart>
<mods:affiliation>School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, China; e-mail: maizq</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jia, Feng-Long</mods:namePart>
<mods:affiliation>Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou 510275, China; &amp; corresponding author</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>@ mail &amp; corresponding author</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>63</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>102</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.37520/aemnp.2023.003</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.37520/aemnp.2023.003</mods:identifier>
<mods:identifier type="ISSN">1804-6487</mods:identifier>
<mods:identifier type="Zenodo-Dep">7887332</mods:identifier>
<mods:identifier type="ZooBank">937DF12A-BC2F-4406-802F-63838312B0C1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03975B72FFC7760C9682CD0D5126FA2C" httpUri="http://treatment.plazi.org/id/03975B72FFC7760C9682CD0D5126FA2C" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
<subSubSection box="[967,1297,285,309]" pageId="4" pageNumber="79" type="nomenclature">
<paragraph blockId="4.[967,1297,285,309]" box="[967,1297,285,309]" pageId="4" pageNumber="79">
<heading box="[967,1297,285,309]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="79" reason="3">
<taxonomicName ID-CoL="7SV6D" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[967,1297,285,309]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="fumatus">
<emphasis bold="true" box="[967,1297,285,309]" pageId="4" pageNumber="79">
<emphasis bold="true" box="[967,1154,285,309]" italics="true" pageId="4" pageNumber="79">Cybister fumatus</emphasis>
<bibRefCitation author="SHARP D." box="[1161,1297,286,309]" pageId="4" pageNumber="79" pagination="179 - 1003" refId="ref18695" refString="SHARP D. 1882: On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series II 2: 179 - 1003 + pls 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1055,1209,316,335]" pageId="4" pageNumber="79" type="description">
<paragraph blockId="4.[828,1436,316,396]" box="[1055,1209,316,335]" pageId="4" pageNumber="79">
(
<figureCitation box="[1061,1156,316,335]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="4" pageNumber="79">Figs 3AE</figureCitation>
;
<figureCitation box="[1165,1203,316,335]" captionStart="Fig" captionStartId="25.[151,182,1084,1103]" captionTargetBox="[162,1430,197,1062]" captionTargetPageId="25" captionText="Fig. 20. Map of distribution of Cybister in China." figureDoi="http://doi.org/10.5281/zenodo.7887379" httpUri="https://zenodo.org/record/7887379/files/figure.png" pageId="4" pageNumber="79">20A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="reference_group">
<paragraph blockId="4.[828,1436,316,396]" pageId="4" pageNumber="79">
<treatmentCitationGroup pageId="4" pageNumber="79">
<treatmentCitation author="SHARP D." box="[828,1146,350,369]" page="731" pageId="4" pageNumber="79" year="1882">
<taxonomicName ID-CoL="7SV6D" authority="Sharp, 1882: 731" authorityName="Sharp" authorityPageNumber="731" authorityYear="1882" box="[828,1146,350,369]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="fumatus">
<emphasis box="[828,981,350,369]" italics="true" pageId="4" pageNumber="79">Cybister fumatus</emphasis>
<bibRefCitation author="SHARP D." box="[989,1146,350,369]" pageId="4" pageNumber="79" pagination="179 - 1003" refId="ref18695" refString="SHARP D. 1882: On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series II 2: 179 - 1003 + pls 7 - 18." type="journal article" year="1882">Sharp, 1882: 731</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(orig. descr.);
<treatmentCitation author="FENG H. T." box="[1285,1436,350,369]" page="13" pageId="4" pageNumber="79" year="1936">
<bibRefCitation author="FENG H. T." box="[1285,1403,350,369]" pageId="4" pageNumber="79" pagination="1 - 15" refId="ref17446" refString="FENG H. T. 1936: Notes on some Dytiscidae from Musee Hoang Ho Pai Ho, Tientsin with descriptions of eleven new species. Peking Natural History Bulletin 11 (1936 - 37): 1 - 15." type="journal article" year="1936">FENG (1936)</bibRefCitation>
: 13
</treatmentCitation>
(
<collectingRegion box="[864,948,377,396]" country="China" name="Shanghai" pageId="4" pageNumber="79">Shanghai</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="materials_examined">
<paragraph blockId="4.[828,1437,419,598]" lastBlockId="5.[150,760,190,422]" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
<emphasis bold="true" box="[828,1006,419,438]" pageId="4" pageNumber="79">Material examined.</emphasis>
None from China but
<specimenCount box="[1197,1301,419,438]" pageId="4" pageNumber="79" type="generic">9 specimens</specimenCount>
from
<emphasis bold="true" box="[1353,1420,419,438]" pageId="4" pageNumber="79">
<collectingCountry box="[1353,1416,419,438]" name="Laos" pageId="4" pageNumber="79">LAOS</collectingCountry>
:
</emphasis>
<specimenCount pageId="4" pageNumber="79" type="generic">4 spec.</specimenCount>
, Kham Mouan prov., Ban Khoun Ngeun,
<geoCoordinate box="[1236,1306,445,464]" degrees="18" direction="north" minutes="07" orientation="latitude" pageId="4" pageNumber="79" precision="925" value="18.116667">18°07N</geoCoordinate>
<geoCoordinate box="[1313,1395,445,464]" degrees="104" direction="east" minutes="29" orientation="longitude" pageId="4" pageNumber="79" precision="925" value="104.48333">104°29E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="79" unit="m" value="200.0">200 m</quantity>
,
<date box="[856,979,472,491]" pageId="4" pageNumber="79" value="2001-05-19" valueMax="2001-05-31" valueMin="2001-05-19">19.31.v.2001</date>
, P. Pachol
<emphasis bold="true" box="[1069,1080,472,491]" pageId="4" pageNumber="79">á</emphasis>
tko leg. (
<collectionCode box="[1158,1225,472,491]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="4" pageNumber="79">NHMB</collectionCode>
,
<collectionCode box="[1235,1301,472,491]" pageId="4" pageNumber="79">LHCM</collectionCode>
);
<specimenCount box="[1317,1380,472,491]" pageId="4" pageNumber="79" type="generic">1 spec.</specimenCount>
,
<collectingRegion country="Laos" name="Viangchan" pageId="4" pageNumber="79">Vientiane prov.</collectingRegion>
, Phou Khao Khouay,
<geoCoordinate box="[1101,1209,499,518]" degrees="18" direction="north" minutes="20.369" orientation="latitude" pageId="4" pageNumber="79" precision="1" value="18.339483">18°20.369N</geoCoordinate>
<geoCoordinate box="[1214,1332,499,518]" degrees="102" direction="east" minutes="48.523" orientation="longitude" pageId="4" pageNumber="79" precision="1" value="102.808716">102°48.523E</geoCoordinate>
, nr. strongly disturbed primary rainforest on light,
<date box="[1157,1280,525,544]" pageId="4" pageNumber="79" value="2008-05-25" valueMax="2008-05-30" valueMin="2008-05-25">25.30.v.2008</date>
, A. Solodovnikov &amp; J. Pedersen leg. (
<collectionCode box="[992,1059,552,571]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="79">ZMUC</collectionCode>
);
<specimenCount box="[1072,1134,552,571]" pageId="4" pageNumber="79" type="generic">1 spec.</specimenCount>
,
<collectingRegion box="[1141,1258,552,571]" country="Laos" name="Xekong" pageId="4" pageNumber="79">Sekong Prov.</collectingRegion>
, Tad Faek waterfalls,
<geoCoordinate box="[828,914,579,598]" degrees="15" direction="north" minutes="14.7" orientation="latitude" pageId="4" pageNumber="79" precision="92" value="15.245">15°14.7N</geoCoordinate>
<geoCoordinate box="[918,1015,579,598]" degrees="106" direction="east" minutes="42.1" orientation="longitude" pageId="4" pageNumber="79" precision="92" value="106.70167">106°42.1E</geoCoordinate>
,
<quantity box="[1022,1075,579,598]" metricMagnitude="2" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="79" unit="m" value="118.0">118 m</quantity>
, light trap,
<date box="[1167,1242,579,598]" pageId="4" pageNumber="79" value="2010-05-08">8.v.2010</date>
, J. Hájek leg.(
<collectionCode box="[1362,1427,579,598]" country="Czech Republic" name="National Museum Prague" pageId="4" pageNumber="79">NMPC</collectionCode>
);
<specimenCount box="[150,217,190,209]" pageId="5" pageNumber="80" type="generic">1 spec.</specimenCount>
, Bolikhamsay prov., Nam Kading NPA research center near
<collectingRegion box="[150,246,216,235]" country="Laos" name="Xekong" pageId="5" pageNumber="80">Ban Phone</collectingRegion>
Kham,
<geoCoordinate box="[319,390,216,235]" degrees="18" direction="north" minutes="20" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="18.333334">18°20N</geoCoordinate>
<geoCoordinate box="[399,480,216,235]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="104.13333">104°08E</geoCoordinate>
,
<quantity box="[491,550,216,235]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="80" unit="m" value="250.0">250 m</quantity>
,
<date box="[561,684,216,235]" pageId="5" pageNumber="80" value="2011-05-23" valueMax="2011-05-29" valueMin="2011-05-23">23.29.v.2011</date>
, NHMB Basel Laos 2011 Expedition M. Brancucci, M. Geiser, D. Hauck, Z. Kraus, A. Phantala &amp; E. Vongphachan (
<collectionCode box="[491,560,270,289]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="80">NHMB</collectionCode>
);
<specimenCount box="[575,638,270,289]" pageId="5" pageNumber="80" type="generic">1 spec.</specimenCount>
, Bolikhamsay prov., Pakkading
<collectingRegion box="[300,392,296,315]" country="Laos" name="Xekong" pageId="5" pageNumber="80">Ban Phone</collectingRegion>
Kham env.
<geoCoordinate box="[496,566,296,315]" degrees="18" direction="north" minutes="19" orientation="latitude" pageId="5" pageNumber="80" precision="925" value="18.316668">18°19N</geoCoordinate>
<geoCoordinate box="[572,652,296,315]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="80" precision="925" value="104.13333">104°08E</geoCoordinate>
<quantity box="[657,756,296,315]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="80" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</quantity>
,
<date box="[150,273,323,342]" pageId="5" pageNumber="80" value="2011-05-23" valueMax="2011-05-29" valueMin="2011-05-23">23.29.v.2011</date>
, NHMB Basel Laos 2011 Expedition M. Brancucci, M. Geiser, D. Hauck, Z. Kraus, A. Phantala &amp; E. Vongphachan (
<collectionCode box="[665,734,350,369]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="5" pageNumber="80">NHMB</collectionCode>
);
<specimenCount pageId="5" pageNumber="80" type="generic">1 spec.</specimenCount>
,
<collectingRegion box="[204,365,376,395]" country="Laos" name="Savannakhet" pageId="5" pageNumber="80">Savannakhet prov.</collectingRegion>
, 10 E of
<collectingRegion box="[440,551,376,395]" country="Laos" name="Savannakhet" pageId="5" pageNumber="80">Savannakhet</collectingRegion>
, Dong Natad forest,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="80" unit="m" value="180.0">180 m</quantity>
,
<date box="[178,265,403,422]" pageId="5" pageNumber="80" value="2011-12-04">4.xii.2011</date>
, J. Šťastný leg. (
<collectionCode box="[407,459,403,422]" pageId="5" pageNumber="80">JSCL</collectionCode>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7887339" ID-Zenodo-Dep="7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="4" pageNumber="79" startId="4.[150,181,2025,2044]" targetBox="[319,1259,662,1992]" targetPageId="4" targetType="figure">
<paragraph blockId="4.[150,1431,2019,2071]" pageId="4" pageNumber="79">
Fig. 3.
<taxonomicName authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[211,465,2025,2044]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="fumatus">
<emphasis box="[211,357,2025,2044]" italics="true" pageId="4" pageNumber="79">Cybister fumatus</emphasis>
<bibRefCitation author="SHARP D." box="[362,465,2025,2044]" pageId="4" pageNumber="79" pagination="179 - 1003" refId="ref18695" refString="SHARP D. 1882: On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series II 2: 179 - 1003 + pls 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
(J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE).
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="80" type="diagnosis">
<paragraph blockId="5.[150,760,447,1526]" pageId="5" pageNumber="80">
<emphasis bold="true" box="[150,264,447,470]" pageId="5" pageNumber="80">Diagnosis.</emphasis>
TL
<quantity box="[304,418,447,470]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="5" pageNumber="80" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">2224 mm</quantity>
, elongate, oblong-oval (
<figureCitation box="[665,747,447,470]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="5" pageNumber="80">Fig. 3A</figureCitation>
). Head dark green, clypeus yellow. Pronotum and elytra dark green, with distinct yellow margins laterally; apex of yellow margin on elytra slightly hooked. Ventral surface black; with lateral part of metaventrite and metacoxae yellow; abdominal ventrites IIIV with a pale spot on each side (
<figureCitation box="[263,347,639,662]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="5" pageNumber="80">Fig. 3B</figureCitation>
). Dorsal surface smooth in both sexes. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 13 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; metatarsal claws single in both sexes. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (
<figureCitation box="[450,533,991,1014]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="5" pageNumber="80">Fig. 3C</figureCitation>
); gradually narrowed from base to near apex in ventral view, apex furcate, with a deep acute-angled notch medially, lateral margins of apex straight and parallel (
<figureCitation box="[425,507,1087,1110]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="5" pageNumber="80">Fig. 3D</figureCitation>
). Parameres moderately broad, approximately the same width apically (
<figureCitation box="[652,735,1119,1142]" captionStart="Fig" captionStartId="4.[150,181,2025,2044]" captionTargetBox="[319,1259,662,1992]" captionTargetId="figure-235@4.[307,1274,649,2008]" captionTargetPageId="4" captionText="Fig. 3. Cybister fumatus Sharp, 1882 (J♀, Laos). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887339" httpUri="https://zenodo.org/record/7887339/files/figure.png" pageId="5" pageNumber="80">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph blockId="5.[150,760,447,1526]" pageId="5" pageNumber="80">
<emphasis bold="true" box="[150,245,1151,1174]" pageId="5" pageNumber="80">Habitat.</emphasis>
In
<collectingCountry box="[285,337,1151,1174]" name="Laos" pageId="5" pageNumber="80">Laos</collectingCountry>
all specimens were collected at lower altitudes between 180 and
<quantity box="[450,521,1183,1206]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="80" unit="m" value="300.0">300 m</quantity>
above sea level. The specimen from
<collectingRegion box="[313,393,1215,1238]" country="Laos" name="Xekong" pageId="5" pageNumber="80">Sekong</collectingRegion>
was attracted by a light trap placed near the border of a lowland river. The single specimen from Dong Natad was collected in a shallow swamp in lowland monsoon primary forest, associated with
<taxonomicName authority="(Swartz, 1808)" baseAuthorityName="Swartz" baseAuthorityYear="1808" class="Insecta" family="Dytiscidae" genus="Hyphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="lyratus">
<emphasis italics="true" pageId="5" pageNumber="80">Hyphydrus lyratus</emphasis>
(Swartz, 1808)
</taxonomicName>
,
<taxonomicName box="[394,545,1343,1366]" class="Insecta" family="Dytiscidae" genus="Hyphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[394,510,1343,1366]" italics="true" pageId="5" pageNumber="80">Hyphydrus</emphasis>
sp.
</taxonomicName>
and
<taxonomicName box="[592,759,1343,1366]" class="Insecta" family="Dytiscidae" genus="Lacconectus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[592,724,1343,1366]" italics="true" pageId="5" pageNumber="80">Lacconectus</emphasis>
sp.
</taxonomicName>
For habitat photos see
<bibRefCitation author="HENDRICH L. &amp; BRANCUCCI M." box="[392,740,1375,1398]" pageId="5" pageNumber="80" pagination="75 - 88" refId="ref17693" refString="HENDRICH L. &amp; BRANCUCCI M. 2013: The genus Cybister Curtis, 1827 in Laos (Coleoptera: Dytiscidae, Cybistrini). Entomologica Basiliensia et Collectionis Frey 34: 75 - 88." type="journal article" year="2013">HENDRICH &amp; BRANCUCCI (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="80" type="distribution">
<paragraph blockId="5.[150,760,447,1526]" pageId="5" pageNumber="80">
<emphasis bold="true" box="[150,299,1407,1430]" pageId="5" pageNumber="80">Distribution.</emphasis>
<collectingCountry box="[309,373,1407,1430]" name="China" pageId="5" pageNumber="80">China</collectingCountry>
(
<collectingRegion box="[391,495,1407,1430]" country="China" name="Shanghai" pageId="5" pageNumber="80">Shanghai</collectingRegion>
) (
<figureCitation box="[520,624,1407,1430]" captionStart="Fig" captionStartId="25.[151,182,1084,1103]" captionTargetBox="[162,1430,197,1062]" captionTargetPageId="25" captionText="Fig. 20. Map of distribution of Cybister in China." figureDoi="http://doi.org/10.5281/zenodo.7887379" httpUri="https://zenodo.org/record/7887379/files/figure.png" pageId="5" pageNumber="80">Fig. 20A</figureCitation>
),
<collectingCountry box="[647,755,1407,1430]" name="Indonesia" pageId="5" pageNumber="80">Indonesia</collectingCountry>
,
<collectingCountry box="[150,206,1439,1462]" name="Laos" pageId="5" pageNumber="80">Laos</collectingCountry>
,
<collectingCountry box="[221,324,1439,1462]" name="Malaysia" pageId="5" pageNumber="80">Malaysia</collectingCountry>
,
<collectingCountry box="[338,438,1439,1462]" name="Thailand" pageId="5" pageNumber="80">Thailand</collectingCountry>
,
<collectingCountry box="[452,544,1439,1462]" name="Vietnam" pageId="5" pageNumber="80">Vietnam</collectingCountry>
(RḖGIMBART 1899,
<bibRefCitation author="ZIMMERMANN A." box="[150,354,1471,1494]" pageId="5" pageNumber="80" refId="ref19188" refString="ZIMMERMANN A. 1920: Pars 71. Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Volumen IV. Junk, Berlin, 326 pp." type="book" year="1920">ZIMMERMANN 1920</bibRefCitation>
,
<bibRefCitation author="FENG H. T." box="[363,480,1471,1494]" pageId="5" pageNumber="80" pagination="1 - 15" refId="ref17446" refString="FENG H. T. 1936: Notes on some Dytiscidae from Musee Hoang Ho Pai Ho, Tientsin with descriptions of eleven new species. Peking Natural History Bulletin 11 (1936 - 37): 1 - 15." type="journal article" year="1936">FENG 1936</bibRefCitation>
,
<bibRefCitation author="NILSSON A. N." box="[490,638,1471,1494]" pageId="5" pageNumber="80" refId="ref18306" refString="NILSSON A. N. 1995: Noteridae and Dytiscidae: Annotated check list of the Noteridae and Dytiscidae of China (Coleoptera). Pp. 35 - 96. In: JACH M. A. &amp; JI L. (eds): Water beetles of China, Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, 410 pp." type="book" year="1995">NILSSON 1995</bibRefCitation>
,
<bibRefCitation author="HENDRICH L. &amp; BRANCUCCI M." pageId="5" pageNumber="80" pagination="75 - 88" refId="ref17693" refString="HENDRICH L. &amp; BRANCUCCI M. 2013: The genus Cybister Curtis, 1827 in Laos (Coleoptera: Dytiscidae, Cybistrini). Entomologica Basiliensia et Collectionis Frey 34: 75 - 88." type="journal article" year="2013">HENDRICH &amp; BRANCUCCI 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>