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<mods:title id="90EE232EECB34ED03A183A64716B9FF5">Two new records of hippolytoid shrimps, Lysmata hochi Baeza &amp; Anker, 2008 (Decapoda: Lysmatidae) and Lysmata amboinensis (de Man, 1888) from Lakshadweep Islands, India with taxonomic notes</mods:title>
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<emphasis id="B951EAECDF26FFD930C5FAA5F06AFA81" bold="true" box="[151,328,1341,1368]" italics="true" pageId="1" pageNumber="354">Lysmata hochi</emphasis>
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, figs 1–3, (
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locality: 
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, Caribbean coast of 
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. 
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LYSLHOC/ 
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, 
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(CL 10.0–12.0 mm), Arabian Sea, off Agatti Islands, 
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, 
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, 
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C, 
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35 ppt, 
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deep, 
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.
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<paragraph id="8B9A36FEDF26FFDA3095F994F06DFEBE" blockId="1.[151,1437,1479,2011]" lastBlockId="2.[151,1437,153,359]" lastPageId="2" lastPageNumber="355" pageId="1" pageNumber="354">
<emphasis id="B951EAECDF26FFD93095F994F066F9F3" bold="true" box="[199,324,1551,1578]" pageId="1" pageNumber="354">Diagnosis:</emphasis>
Rostrum (
<figureCitation id="131E2A7BDF26FFD931EFF98BF324F9F3" box="[445,518,1552,1578]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1a</figureCitation>
) moderately long, straight and tip is just exceeding the first antennule peduncle, 0.3 times as long as carapace, dorso-rostral carina curving downwards but recurved on tip; armed dorsally with four teeth and an epigastric tooth, stiff setae present in middle of dorso-rostral carina for each tooth (
<figureCitation id="131E2A7BDF26FFD93495F9C3F432F9AB" box="[1223,1296,1624,1650]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1a</figureCitation>
), third tooth starting from the level of orbital margin; ventral border bearing a small tooth near to the tip. Carapace glabrous with a strong and elongated antennal spine, pterygostominal angle somewhat rounded, ventral margin of carapace not rounded. Eyes are rounded and with large ocellus; orbital margin regularly concave with ventral end with setae. Stylocerite broad, sharply bifid and reaching near to distal margin of basal antennular peduncle. Antennule peduncle not exceeding the scaphocerite, disto-dorsally few spinules are present on each peduncle, lateral flagellum contains accessory branches with an unfuis-shaped segment (
<figureCitation id="131E2A7BDF26FFD93316F8ABF2B2F892" box="[836,912,1840,1867]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1b</figureCitation>
). First Pereopods (
<figureCitation id="131E2A7BDF26FFD93436F8ABF5EBF892" box="[1124,1225,1840,1867]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1c, d</figureCitation>
) short, moderately stout with simple chela and extended beyond the scaphocerite. Pereopod II (
<figureCitation id="131E2A7BDF26FFD933A8F8CFF561F8B7" box="[1018,1091,1876,1902]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1e</figureCitation>
) slender and slightly unequal; merus composed of 17–18 articles; carpus composed of 25–26 articles and chela is small with numerous long setae. Posterior 3 Pereopods are similar, overreaching scaphocerite by dactylus; merus bearing 1–4 spines on distally (
<figureCitation id="131E2A7BDF26FFD93527F807F183F802" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="1" pageNumber="354">Fig 1</figureCitation>
f–h); propodus with 7–9 spinules ventrally including a distal pair; dactylus biunguiculate, 0.15–0.17 times as long as propodus, accessory spine shorter than terminal spines, flexor margin armed with 2 distinct movable spines. Anterior three pleomeres arched round margin and without posterior spine; pleura IV and V moderately angled posteroventrally; pleura VI with a spine on posteroventrally. Telson (
<figureCitation id="131E2A7BDF25FFDA33C6FF7AF521FF25" box="[916,1027,225,252]" captionStart="FIGURE 1" captionStartId="2.[151,250,1646,1671]" captionTargetBox="[171,1414,389,1612]" captionTargetId="figure@2.[164,1423,381,1621]" captionTargetPageId="2" captionText="FIGURE 1. Lysmata hochi Baeza &amp; Anker, 2008 (CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1st pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson." figureDoi="http://doi.org/10.5281/zenodo.3733673" httpUri="https://zenodo.org/record/3733673/files/figure.png" pageId="2" pageNumber="355">Fig 1i &amp; j</figureCitation>
) 1.5 times long as abdominal somite VI and 2.9 times as long as the width of base, armed with 1 pair of dorsolateral spine appeared medially and 2 pairs of posterior spines (outer pair short and mesial pair larger) with long setae. Eggs are small and numerous, 
<quantity id="4CDD9B1BDF25FFDA3568FEB3F1E1FEBE" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="5.1" metricValueMin="4.0" pageId="2" pageNumber="355" unit="mm" value="0.455" valueMax="0.51" valueMin="0.4">0.4–0.51 mm</quantity>
in diameter.
</paragraph>
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<paragraph id="8B9A36FEDF25FFDA30C5F9F5F28BF916" blockId="2.[151,1437,1646,1743]" pageId="2" pageNumber="355">
<emphasis id="B951EAECDF25FFDA30C5F9F5F037F95E" bold="true" box="[151,277,1646,1671]" pageId="2" pageNumber="355">FIGURE 1.</emphasis>
<taxonomicName id="4C254D7DDF25FFDA314CF9F4F3BCF95E" authority="Baeza &amp; Anker, 2008" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[286,670,1646,1671]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="355" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF25FFDA314CF9F4F096F95F" box="[286,436,1646,1670]" italics="true" pageId="2" pageNumber="355">Lysmata hochi</emphasis>
<bibRefCitation id="EFB44B0FDF25FFDA31EFF9F4F3BCF95E" author="Baeza &amp; Anker" box="[445,670,1646,1671]" firstAuthor="Baeza" pageId="2" pageNumber="355" pagination="148 - 155" refId="ref4738" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the Southwestern Caribbean Sea (Caridea: Hippolytidae) Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker, 2008</bibRefCitation>
</taxonomicName>
(CL 10 mm). from Agatti Island, Lakshadweep, India. a, Lateral view of carapace; b, Antennule flagellum-accessory branches; c, Pereopod 1; d, fingers of 1 
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pereopod; e, Pereopod 2; f, Pereopod 3; g, Pereopod 4; h, pereopod 5; i, Dorsal view of telson; j, Posterior end of telson.
</paragraph>
</caption>
<paragraph id="8B9A36FEDF25FFDA3095F967F2BEF836" blockId="2.[151,1437,1788,2031]" pageId="2" pageNumber="355">
<emphasis id="B951EAECDF25FFDA3095F967F08BF8CE" bold="true" box="[199,425,1788,1815]" pageId="2" pageNumber="355">Colouration of life:</emphasis>
Body (
<figureCitation id="131E2A7BDF25FFDA31ABF966F317F8CE" box="[505,565,1789,1815]" captionStart="FIGURE 2" captionStartId="3.[151,250,1578,1603]" captionTargetBox="[189,1398,529,1553]" captionTargetId="figure@3.[189,1398,525,1553]" captionTargetPageId="3" captionText="FIGURE 2. Picture representation of Lysmata hochi Baeza &amp; Anker, 2008 from Agatti Island, Lakshadweep, India." figureDoi="http://doi.org/10.5281/zenodo.3733675" httpUri="https://zenodo.org/record/3733675/files/figure.png" pageId="2" pageNumber="355">Fig 2</figureCitation>
) semi-translucent and with several transverse red bands and patches. Carapace with four transverse red bands on dorsally and few irregular longitudinal red bands on laterally; anterior margin of carapace with fine red patches. Eyestalk semi-translucent and fine pale red patches on dorsally; cornea darken to black. Abdominal pleura I–VI with a broad transverse band on dorso-posterior end and a longitudinal red band on laterally; abdominal pleura III with a red dot on mid-dorsally. Anterior and posterior 1/3 of telson and uropods with red, middle with translucent. Maxilliped and 1 
<superScript id="7C509BB6DF25FFDA328EF82BF3C5F867" attach="left" box="[732,743,1968,1982]" fontSize="6" pageId="2" pageNumber="355">st</superScript>
two pereopods are semi-translucent with several red patches. Pereopod III–V are mostly red and white bands; dactylus with white.
</paragraph>
</subSubSection>
<subSubSection id="C33F6575DF24FFDB3095FF03F46AFF22" pageId="3" pageNumber="356" type="distribution">
<paragraph id="8B9A36FEDF24FFDB3095FF03F46AFF22" blockId="3.[151,1436,152,504]" pageId="3" pageNumber="356">
<emphasis id="B951EAECDF24FFDB3095FF03F041FF6A" bold="true" box="[199,355,152,179]" pageId="3" pageNumber="356">Distribution:</emphasis>
<collectingRegion id="49E1F81CDF24FFDB3138FF02F337FF6A" box="[362,533,153,179]" country="Panama" name="Bocas del Toro" pageId="3" pageNumber="356">Bocas del Toro</collectingRegion>
, 
<collectingCountry id="F332766EDF24FFDB3272FF02F35BFF6A" box="[544,633,153,179]" name="Panama" pageId="3" pageNumber="356">Panama</collectingCountry>
(
<typeStatus id="549E885CDF24FFDB32DAFF02F398FF6A" box="[648,698,153,179]" pageId="3" pageNumber="356">type</typeStatus>
locality), Cahuita, 
<collectingCountry id="F332766EDF24FFDB33C3FF02F529FF6A" box="[913,1035,153,179]" name="Costa Rica" pageId="3" pageNumber="356">Costa Rica</collectingCountry>
in the southwestern Caribbean Sea, Western Atlantic at depth of 
<quantity id="4CDD9B1BDF24FFDB3188FF27F314FF0E" box="[474,566,188,215]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="3" pageNumber="356" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 m</quantity>
(
<bibRefCitation id="EFB44B0FDF24FFDB3217FF26F21CFF0E" author="Baeza &amp; Anker" box="[581,830,189,215]" firstAuthor="Baeza" pageId="3" pageNumber="356" pagination="148 - 155" refId="ref4738" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the Southwestern Caribbean Sea (Caridea: Hippolytidae) Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker, 2008</bibRefCitation>
). Currently, we noticed the distribution of this species occurred in Agatti Island, 
<collectingCountry id="F332766EDF24FFDB31E9FF7AF37AFF22" box="[443,600,225,251]" name="India" pageId="3" pageNumber="356">Lakshadweep</collectingCountry>
, the Arabian Sea at depth of 
<quantity id="4CDD9B1BDF24FFDB33C8FF7BF2FEFF22" box="[922,988,224,251]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="356" unit="m" value="0.5">0.5 m</quantity>
, new record from Indian waters.
</paragraph>
</subSubSection>
<subSubSection id="C33F6575DF24FFDB3095FE9EF449F82A" pageId="3" pageNumber="356" type="discussion">
<paragraph id="8B9A36FEDF24FFDB3095FE9EF03DFE21" blockId="3.[151,1436,152,504]" pageId="3" pageNumber="356">
<emphasis id="B951EAECDF24FFDB3095FE9EF01FFEC6" bold="true" box="[199,317,261,287]" pageId="3" pageNumber="356">Remarks:</emphasis>
Originally, 
<taxonomicName id="4C254D7DDF24FFDB3197FE9DF34BFEC6" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[453,617,261,287]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB3197FE9DF34BFEC6" box="[453,617,261,287]" italics="true" pageId="3" pageNumber="356">Lysmata hochi</emphasis>
</taxonomicName>
was described based on three individuals from 
<collectingRegion id="49E1F81CDF24FFDB34D6FE9EF412FEC6" box="[1156,1328,261,287]" country="Panama" name="Bocas del Toro" pageId="3" pageNumber="356">Bocas del Toro</collectingRegion>
, 
<collectingCountry id="F332766EDF24FFDB356FFE9EF4BBFEC6" box="[1341,1433,261,287]" name="Panama" pageId="3" pageNumber="356">Panama</collectingCountry>
, Caribbean coast (
<bibRefCitation id="EFB44B0FDF24FFDB310EFEB2F37AFE9A" author="Baeza &amp; Anker" box="[348,600,297,323]" firstAuthor="Baeza" pageId="3" pageNumber="356" pagination="148 - 155" refId="ref4738" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the Southwestern Caribbean Sea (Caridea: Hippolytidae) Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker, 2008</bibRefCitation>
). In the present study, three specimens from 
<collectingCountry id="F332766EDF24FFDB3406FEB2F5ACFE9A" box="[1108,1166,297,323]" name="India" pageId="3" pageNumber="356">India</collectingCountry>
are closely agreed with the original description and illustrations of 
<bibRefCitation id="EFB44B0FDF24FFDB3229FED6F25BFEBE" author="Baeza &amp; Anker" box="[635,889,333,359]" firstAuthor="Baeza" pageId="3" pageNumber="356" pagination="148 - 155" refId="ref4738" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the Southwestern Caribbean Sea (Caridea: Hippolytidae) Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker (2008)</bibRefCitation>
and it represents the first report from 
<collectingCountry id="F332766EDF24FFDB3571FED6F47DFEBE" box="[1315,1375,333,359]" name="India" pageId="3" pageNumber="356">India</collectingCountry>
, Arabian Sea. Although some differences were noticed, this is intraspecific and noteworthy. For instance, in the present specimens of the carpus of 2 
<superScript id="7C509BB6DF24FFDB3187FE0FF0C7FE7B" attach="left" box="[469,485,404,418]" fontSize="6" pageId="3" pageNumber="356">nd</superScript>
pereopod composed of 25–26 articles, merus of posterior three pereopods were bearing 1–4 spines, dactylus 0.15–0.17 times as long as propodus and telson 2.9 times as long as the width of the base (fig. 1i &amp; j).
</paragraph>
<caption id="DF5A6676DF24FFDB30C5F9B1F40FF99A" ID-DOI="http://doi.org/10.5281/zenodo.3733675" ID-Zenodo-Dep="3733675" box="[151,1325,1578,1603]" httpUri="https://zenodo.org/record/3733675/files/figure.png" pageId="3" pageNumber="356" startId="3.[151,250,1578,1603]" targetBox="[189,1398,529,1553]" targetPageId="3">
<paragraph id="8B9A36FEDF24FFDB30C5F9B1F40FF99A" blockId="3.[151,1325,1578,1603]" box="[151,1325,1578,1603]" pageId="3" pageNumber="356">
<emphasis id="B951EAECDF24FFDB30C5F9B1F031F99A" bold="true" box="[151,275,1578,1603]" pageId="3" pageNumber="356">FIGURE 2.</emphasis>
Picture representation of 
<taxonomicName id="4C254D7DDF24FFDB324BF9B7F2ACF99A" authority="Baeza &amp; Anker, 2008" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[537,910,1578,1603]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB324BF9B7F38EF99A" box="[537,684,1579,1603]" italics="true" pageId="3" pageNumber="356">Lysmata hochi</emphasis>
<bibRefCitation id="EFB44B0FDF24FFDB32E1F9B0F2ACF99A" author="Baeza &amp; Anker" box="[691,910,1578,1603]" firstAuthor="Baeza" pageId="3" pageNumber="356" pagination="148 - 155" refId="ref4738" refString="Baeza, J. A. &amp; Anker, A. (2008) Lysmata hochi n. sp., a new hermaphroditic shrimp from the Southwestern Caribbean Sea (Caridea: Hippolytidae) Journal of Crustacean Biology, 28 (1), 148 - 155. https: // doi. org / 10.1651 / 07 - 2839 R. 1" type="journal article" year="2008">Baeza &amp; Anker, 2008</bibRefCitation>
</taxonomicName>
from Agatti Island, Lakshadweep, India.
</paragraph>
</caption>
<paragraph id="8B9A36FEDF24FFDB3095F9E9F541F8BA" blockId="3.[151,1437,1649,2036]" pageId="3" pageNumber="356">
In Indian waters, 
<taxonomicName id="4C254D7DDF24FFDB31DEF9E9F3B9F952" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[396,667,1649,1676]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="debelius">
<emphasis id="B951EAECDF24FFDB31DEF9E9F325F952" box="[396,519,1649,1675]" italics="true" pageId="3" pageNumber="356">L. debelius</emphasis>
<bibRefCitation id="EFB44B0FDF24FFDB325CF9E9F3B9F952" author="Bruce" box="[526,667,1649,1676]" firstAuthor="Bruce" pageId="3" pageNumber="356" pagination="115 - 120" refId="ref4942" refString="Bruce, A. J. (1983) Lysmata debelius new species, a new hippolytid shrimp from the Philippines. Revue Francaise d'Aquariologie, 9, 115 - 120." type="journal article" year="1983">Bruce, 1983</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C254D7DDF24FFDB32F4F9E9F525F952" authority="(de Man, 1888)" baseAuthorityName="de Man" baseAuthorityYear="1888" box="[678,1031,1649,1676]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B951EAECDF24FFDB32F4F9E9F272F952" box="[678,848,1649,1675]" italics="true" pageId="3" pageNumber="356">L. amboinensis</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB330DF9EAF2DDF952" author="de Man" box="[863,1023,1649,1676]" firstAuthor="de Man" pageId="3" pageNumber="356" pagination="289 - 600" refId="ref5466" refString="De Man, J. G. (1888) Bericht uber die von Herrn Dr. J. Brock imindischen Archipelges ammelten Decapoden und Stomatopoden. Archivfur Naturgeschichte, 53 (3), 289 - 600, pls. 11 - 22 a. https: // doi. org / 10.5962 / bhl. part. 4747" type="journal article" year="1888">de Man, 1888</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C254D7DDF24FFDB3441F9E9F46DF952" authority="de Man, 1902" authorityName="de Man" authorityYear="1902" box="[1043,1359,1649,1676]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="ternatensis">
<emphasis id="B951EAECDF24FFDB3441F9E9F58EF952" box="[1043,1196,1650,1675]" italics="true" pageId="3" pageNumber="356">L. ternatensis</emphasis>
<bibRefCitation id="EFB44B0FDF24FFDB34E1F9EAF46DF952" author="de Man" box="[1203,1359,1649,1676]" firstAuthor="de Man" pageId="3" pageNumber="356" pagination="466 - 929" refId="ref5526" refString="De Man, J. G. (1902) Die von Herrn Professor Kukenthal im Indiscen Archipel gesammelten Dekapoden und Stomatopoden. Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 25, 466 - 929. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">de Man, 1902</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C254D7DDF24FFDB35D4F9E9F08AF969" authority="(Stimpson, 1860)" baseAuthorityName="Stimpson" baseAuthorityYear="1860" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="B951EAECDF24FFDB35D4F9E9F1FDF976" italics="true" pageId="3" pageNumber="356">L. vittata</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB30BFF90EF082F969" author="Stimpson" box="[237,416,1685,1712]" firstAuthor="Stimpson" pageId="3" pageNumber="356" pagination="22 - 47" refId="ref7333" refString="Stimpson, W. (1860) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladore Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VIII, Crustacea Macrura. Proceedings of the Academy of Natural Sciences of Philadelphia, 1860, 22 - 47." type="journal article" year="1860">Stimpson, 1860</bibRefCitation>
)
</taxonomicName>
were observed from Bay of Bengal and Arabian Sea (
<bibRefCitation id="EFB44B0FDF24FFDB3454F90EF41BF976" author="Radhakrishnan" box="[1030,1337,1685,1711]" etAl="et al." firstAuthor="Radhakrishnan" pageId="3" pageNumber="356" pagination="50 - 72" refId="ref6928" refString="Radhakrishnan, E. V., Deshmukh, V. D., Maheswarudu, G., Josileen, J., Dineshbabu, A. P., Philipose, K. K., Sarada, P. T., Pillai, S., Lakshmi, S. K. N., Chakraborty, R. D., Dash, G., Sajeev, C. K., Thirumilu, P., Sridhara, B., Muniyappa, Y., Sawant, A. D., Vaidya, N. G., Johny, R. D. Verma, J. B., Baby, P. K., Unnikrishnan, C., Ramachandran, N. P., Vairamani, A., Palanichamy, A., Radhakrishnan, M. &amp; Raju, B. (2012) Prawn fauna (Crustacea: Decapoda) of India-An annotated checklist of the Penaeoid, Sergestoid, Stenopodid and Caridean prawns. Marine Biological Association of India, 54 (1), 50 - 72. https: // doi. org / 10.6024 / jmbai. 2012.54.1.01697 - 08" type="journal article" year="2012">
Radhakrishnan 
<emphasis id="B951EAECDF24FFDB34E5F90DF5D4F976" box="[1207,1270,1685,1711]" italics="true" pageId="3" pageNumber="356">et al.,</emphasis>
2012
</bibRefCitation>
; 
<bibRefCitation id="EFB44B0FDF24FFDB3516F90EF034F90A" author="Prakash" etAl="et al." firstAuthor="Prakash" pageId="3" pageNumber="356" pagination="576 - 582" refId="ref6593" refString="Prakash, S. &amp; Baeza, J. A. (2017) A new species of LysmataRisso, 1816 (Crustacea, Decapoda, Lysmatidae) from the Gulf of Mexico. Zootaxa, 4363 (4), 576 - 582. https: // doi. org / 10.11646 / zootaxa. 4363.4.10" type="journal article" year="2017">
Prakash 
<emphasis id="B951EAECDF24FFDB30C5F921F1F7F90A" box="[151,213,1721,1747]" italics="true" pageId="3" pageNumber="356">et al.,</emphasis>
2017
</bibRefCitation>
; 
<bibRefCitation id="EFB44B0FDF24FFDB3173F922F339F90A" author="Madhavan" box="[289,539,1721,1748]" etAl="et al." firstAuthor="Madhavan" pageId="3" pageNumber="356" pagination="351 - 364" refId="ref6371" refString="Madhavan, M., Purushothaman, P., Akash, S., Bharathi, S., Sheena, J., Dhinakaran, A., Charan, R., Kumar, A. &amp; Lal, K. K. (2019) New record of Thor hainanensis Xu &amp; Li, 2014 and taxonomical remarks on Lysmata ternatensis de Man, 1902 (Decapoda: Thoridae &amp; Lysmatidae) from the Lakshadweep Islands, India. Zootaxa, 4624 (3), 351 - 364. https: // doi. org / 10.11646 / zootaxa. 4624.3.4" type="journal article" year="2019">
Madhavan 
<emphasis id="B951EAECDF24FFDB31CFF921F0EFF90A" box="[413,461,1721,1747]" italics="true" pageId="3" pageNumber="356">et al</emphasis>
., 2019
</bibRefCitation>
) which were differed distinctively by the colour pattern and combination of various morphological traits even though most importantly, the dorsal rostral formula. For instance, the dorsal rostrum formula is 1-2+ 
<quantity id="4CDD9B1BDF24FFDB3116F89BF0AAF8C2" box="[324,392,1792,1819]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="3" pageNumber="356" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4-5 in</quantity>
<taxonomicName id="4C254D7DDF24FFDB31DDF899F23CF8C2" authority="(Chace et al., 1997)" baseAuthorityName="Chace" baseAuthorityYear="1997" box="[399,798,1793,1820]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B951EAECDF24FFDB31DDF899F31BF8C2" box="[399,569,1793,1819]" italics="true" pageId="3" pageNumber="356">L. amboinensis</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB3215F89AF237F8C2" author="Chace" box="[583,789,1793,1820]" etAl="et al." firstAuthor="Chace" pageId="3" pageNumber="356" pagination="1 - 106" refId="ref5085" refString="Chace, F. A. Jr. (1997) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 7: Families Atyidae, Eugonatonotidae, Rhynchocinetidae, Bathypalaemonellidae, Processidae, and Hippolytidae. Smithsonian Contributions to Zoology, 587, 1 - 106. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1997">
Chace 
<emphasis id="B951EAECDF24FFDB32C4F899F3EAF8C2" box="[662,712,1793,1819]" italics="true" pageId="3" pageNumber="356">et al</emphasis>
., 1997
</bibRefCitation>
)
</taxonomicName>
, 
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<taxonomicName id="4C254D7DDF24FFDB3328F899F40BF8C5" authority="(Madhavan et al., 2019)" baseAuthorityName="Madhavan" baseAuthorityYear="2019" box="[890,1321,1793,1820]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="ternatensis">
<emphasis id="B951EAECDF24FFDB3328F899F531F8C2" box="[890,1043,1794,1819]" italics="true" pageId="3" pageNumber="356">L. ternatensis</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB3473F89AF43CF8C2" author="Madhavan" box="[1057,1310,1793,1820]" etAl="et al." firstAuthor="Madhavan" pageId="3" pageNumber="356" pagination="351 - 364" refId="ref6371" refString="Madhavan, M., Purushothaman, P., Akash, S., Bharathi, S., Sheena, J., Dhinakaran, A., Charan, R., Kumar, A. &amp; Lal, K. K. (2019) New record of Thor hainanensis Xu &amp; Li, 2014 and taxonomical remarks on Lysmata ternatensis de Man, 1902 (Decapoda: Thoridae &amp; Lysmatidae) from the Lakshadweep Islands, India. Zootaxa, 4624 (3), 351 - 364. https: // doi. org / 10.11646 / zootaxa. 4624.3.4" type="journal article" year="2019">
Madhavan 
<emphasis id="B951EAECDF24FFDB34F2F899F5F0F8C2" box="[1184,1234,1793,1819]" italics="true" pageId="3" pageNumber="356">et al</emphasis>
., 2019
</bibRefCitation>
<emphasis id="B951EAECDF24FFDB354CF899F40BF8C5" box="[1310,1321,1794,1820]" italics="true" pageId="3" pageNumber="356">)</emphasis>
</taxonomicName>
<emphasis id="B951EAECDF24FFDB357BF899F40DF8C5" box="[1321,1327,1794,1820]" italics="true" pageId="3" pageNumber="356">,</emphasis>
1+ 
<quantity id="4CDD9B1BDF24FFDB3501F89BF45DF8C2" box="[1363,1407,1792,1819]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="356" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C254D7DDF24FFDB35D4F899F0CEF899" authority="(Prakash et al., 2016)" baseAuthorityName="Prakash" baseAuthorityYear="2016" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="debelius">
<emphasis id="B951EAECDF24FFDB35D4F899F1D6F8E6" italics="true" pageId="3" pageNumber="356">L. debelius</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB3151F8BEF0C7F899" author="Prakash" box="[259,485,1829,1856]" etAl="et al." firstAuthor="Prakash" pageId="3" pageNumber="356" pagination="1 - 6" refId="ref6530" refString="Prakash, S., Kumar, A. &amp; Subramoniam, T. (2016) New records of marine ornamental shrimps (Decapoda: Stenopodidea and Caridea) from the Gulf of Mannar, Tamil Nadu, India. Check List, 12 (6), 1 - 6. https: // doi. org / 10.15560 / 12.6.2010" type="journal article" year="2016">
Prakash 
<emphasis id="B951EAECDF24FFDB3137F8BDF0B4F8E6" box="[357,406,1829,1855]" italics="true" pageId="3" pageNumber="356">et al</emphasis>
., 2016
</bibRefCitation>
)
</taxonomicName>
and 1+ 
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<taxonomicName id="4C254D7DDF24FFDB3220F8BDF2EBF8E6" authority="(Wang &amp; Sha, 2018)" baseAuthorityName="Wang &amp; Sha" baseAuthorityYear="2018" box="[626,969,1829,1856]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="B951EAECDF24FFDB3220F8BDF3C5F899" box="[626,743,1830,1856]" italics="true" pageId="3" pageNumber="356">L. vittata (</emphasis>
<bibRefCitation id="EFB44B0FDF24FFDB32BAF8BDF2E3F8E6" author="Wang &amp; Sha" box="[744,961,1829,1856]" firstAuthor="Wang" pageId="3" pageNumber="356" pagination="28 - 40" refId="ref7451" refString="Wang, Y. R. &amp; Sha, Z. L. (2018) Description of two new species of Lysmata Risso, 1816 (Decapoda, Lysmatidae) from the China seas, with remarks on Lysmata vittata (Stimpson 1860). Zootaxa, 4392 (1), 28 - 40. https: // doi. org / 10.11646 / zootaxa. 4392.1.2" type="journal article" year="2018">Wang &amp; Sha, 2018</bibRefCitation>
)
</taxonomicName>
. Also, 
<taxonomicName id="4C254D7DDF24FFDB3447F8BDF552F8E6" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[1045,1136,1829,1855]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB3447F8BDF552F8E6" box="[1045,1136,1829,1855]" italics="true" pageId="3" pageNumber="356">L. hochi</emphasis>
</taxonomicName>
is very close to 
<taxonomicName id="4C254D7DDF24FFDB357AF8BDF0A3F8BA" authority="(de Man, 1902)" baseAuthorityName="de Man" baseAuthorityYear="1902" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="kuekenthali">
<emphasis id="B951EAECDF24FFDB357AF8BDF1E8F8BA" italics="true" pageId="3" pageNumber="356">L. kuekenthali</emphasis>
(
<bibRefCitation id="EFB44B0FDF24FFDB308BF8D2F058F8BA" author="de Man" box="[217,378,1865,1892]" firstAuthor="de Man" pageId="3" pageNumber="356" pagination="466 - 929" refId="ref5526" refString="De Man, J. G. (1902) Die von Herrn Professor Kukenthal im Indiscen Archipel gesammelten Dekapoden und Stomatopoden. Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 25, 466 - 929. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">de Man, 1902</bibRefCitation>
)
</taxonomicName>
, where it was differed by the presence of a bifid tip of stylocerite.
</paragraph>
<paragraph id="8B9A36FEDF24FFDB3095F8F6F449F82A" blockId="3.[151,1437,1649,2036]" pageId="3" pageNumber="356">
Three specimens of 
<taxonomicName id="4C254D7DDF24FFDB31F4F8F5F322F85E" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[422,512,1901,1927]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB31F4F8F5F322F85E" box="[422,512,1901,1927]" italics="true" pageId="3" pageNumber="356">L. hochi</emphasis>
</taxonomicName>
were sequenced for COI which are submitted to NCBI (Accession No: 
<accessionNumber id="9476AB1DDF24FFDB3545F8F6F4B7F85E" box="[1303,1429,1901,1927]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT079203" pageId="3" pageNumber="356">MT079203</accessionNumber>
, 
<accessionNumber id="9476AB1DDF24FFDB30C5F80AF037F872" box="[151,277,1937,1963]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT079204" pageId="3" pageNumber="356">MT079204</accessionNumber>
&amp; 
<accessionNumber id="9476AB1DDF24FFDB3110F80AF0E2F872" box="[322,448,1937,1963]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT079208" pageId="3" pageNumber="356">MT079208</accessionNumber>
). The ML analysis showed that the specimens from the Indian and 
<collectingRegion id="49E1F81CDF24FFDB34B8F809F4BEF875" box="[1258,1436,1937,1964]" country="Panama" name="Bocas del Toro" pageId="3" pageNumber="356">Bocas del Toro</collectingRegion>
(
<accessionNumber id="9476AB1DDF24FFDB30CDF82EF03FF809" box="[159,285,1973,2000]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC962202" pageId="3" pageNumber="356">KC962202</accessionNumber>
) materials of 
<taxonomicName id="4C254D7DDF24FFDB31E4F82DF330F816" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[438,530,1973,1999]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB31E4F82DF330F816" box="[438,530,1973,1999]" italics="true" pageId="3" pageNumber="356">L. hochi</emphasis>
</taxonomicName>
form a robust monophyletic clade with 
<taxonomicName id="4C254D7DDF24FFDB339DF82DF5A2F816" authorityName="Schmitt" authorityYear="1924" box="[975,1152,1973,1999]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="B951EAECDF24FFDB339DF82DF5A2F816" box="[975,1152,1973,1999]" italics="true" pageId="3" pageNumber="356">L. galapagensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C254D7DDF24FFDB34E4F82DF403F816" baseAuthorityName="M.J.Rathbun" baseAuthorityYear="1901" box="[1206,1313,1974,1999]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="moorei">
<emphasis id="B951EAECDF24FFDB34E4F82DF403F816" box="[1206,1313,1974,1999]" italics="true" pageId="3" pageNumber="356">L. moorei</emphasis>
</taxonomicName>
with interspecific distances ranged from 20.6%– 25.8%. Intraspecific divergence for 
<taxonomicName id="4C254D7DDF24FFDB33B1F841F51DF82A" authorityName="Baeza &amp; Anker" authorityYear="2008" box="[995,1087,2009,2035]" class="Malacostraca" family="Hippolytidae" genus="Lysmata" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="356" phylum="Arthropoda" rank="species" species="hochi">
<emphasis id="B951EAECDF24FFDB33B1F841F51DF82A" box="[995,1087,2009,2035]" italics="true" pageId="3" pageNumber="356">L. hochi</emphasis>
</taxonomicName>
was showed 0.0% (fig. 3).
</paragraph>
</subSubSection>
</treatment>
</document>