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<mods:title id="7555EF55620C626B31074AB0F7CFE2E6">Three new synonyms within the flower chafer genus Goliathopsis Janson, 1881 (Coleoptera: Scarabaeidae: Cetoniinae) from China</mods:title>
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<paragraph id="8B9A36D1CE03FFB9FF0BF99BFD9FF970" blockId="4.[151,708,1649,1712]" box="[151,557,1649,1676]" pageId="4" pageNumber="95">
<heading id="D0D281BDCE03FFB9FF0BF99BFD9FF970" bold="true" box="[151,557,1649,1676]" fontSize="11" level="1" pageId="4" pageNumber="95" reason="1">
<taxonomicName id="4C254D52CE03FFB9FF0BF99BFD9FF970" authority="Janson, 1881" authorityName="in Janson" authorityYear="1881" box="[151,557,1649,1676]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE03FFB9FF0BF99BFD9FF970" bold="true" box="[151,557,1649,1676]" pageId="4" pageNumber="95">
<emphasis id="B951EAC3CE03FFB9FF0BF99BFE36F970" bold="true" box="[151,388,1649,1676]" italics="true" pageId="4" pageNumber="95">Goliathopsis cervus</emphasis>
<bibRefCitation id="EFB44B20CE03FFB9FE10F99BFD9FF970" author="Janson, O. E." box="[396,557,1649,1675]" pageId="4" pageNumber="95" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
</emphasis>
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<paragraph id="8B9A36D1CE03FFB9FF0BF97FFD76F94B" blockId="4.[151,708,1649,1712]" box="[151,708,1684,1712]" pageId="4" pageNumber="95">
(
<figureCitation id="131E2A54CE03FFB9FF03F97FFF5DF954" box="[159,239,1684,1711]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="4" pageNumber="95">Figs. 5</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FF67F97FFEABF954" box="[251,281,1685,1711]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="4" pageNumber="95">13</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FEB9F97FFEDCF954" box="[293,366,1685,1711]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="4" pageNumber="95">2223</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FEE5F97EFE73F954" box="[377,449,1684,1711]" captionStart-0="FIGURES 3853" captionStart-1="FIGURES 5457" captionStartId-0="7.[151,264,1590,1615]" captionStartId-1="8.[151,264,1902,1927]" captionTargetBox-0="[151,1435,463,1559]" captionTargetBox-1="[186,1367,278,1868]" captionTargetId-0="figure@7.[151,1435,453,1565]" captionTargetId-1="figure@8.[151,1435,266,1878]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." captionText-1="FIGURES 5457. Habitus of Goliathopsis cervus Janson, 1881 (dorsal and ventral view). 5455, male; 5657, female." figureDoi-0="http://doi.org/10.5281/zenodo.3884733" figureDoi-1="http://doi.org/10.5281/zenodo.3884735" httpUri-0="https://zenodo.org/record/3884733/files/figure.png" httpUri-1="https://zenodo.org/record/3884735/files/figure.png" pageId="4" pageNumber="95">5257</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FE51F97FFDA7F954" box="[461,533,1685,1711]" captionStart="FIGURES 89102" captionStartId="23.[151,264,1958,1983]" captionTargetBox="[222,1369,191,1931]" captionTargetId="figure@23.[200,1387,181,1940]" captionTargetPageId="23" captionText="FIGURES 89102. Habitus of Goliathopsis Janson, 1881 species (male in left, female in right; oblique view). 8990, G. cervus Janson, 1881; 9192, G. duponti Antoine, 1991; 9394, G. esquiroli Pouillaude, 1913; 9596, G. ferreroi Antoine, 1991; 9798, G. gressitti Ruter, 1978; 99100, G. lameyi Fairmaire, 1893; 101102, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884753" httpUri="https://zenodo.org/record/3884753/files/figure.png" pageId="4" pageNumber="95">8990</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FDBDF97FFD34F954" box="[545,646,1685,1711]" captionStart="FIGURES 103108" captionStartId="24.[151,264,1020,1045]" captionTargetBox="[154,1430,184,982]" captionTargetId="figure@24.[151,1435,181,994]" captionTargetPageId="24" captionText="FIGURES 103108. Male lectotype of Goliathopsis cervus Janson, 1881 in RMNH (photograph by Jan Krikken). 103, dorsal view; 104, ventral view; 105, lateral view; 106, oblique view; 107, pygidium; 108, labels." figureDoi="http://doi.org/10.5281/zenodo.3884755" httpUri="https://zenodo.org/record/3884755/files/figure.png" pageId="4" pageNumber="95">103108</figureCitation>
,
<figureCitation id="131E2A54CE03FFB9FD0DF97FFD0EF94B" box="[657,700,1685,1712]" captionStart="FIGURE 164" captionStartId="33.[151,250,1616,1641]" captionTargetBox="[159,1427,209,1579]" captionTargetId="figure@33.[151,1436,181,1591]" captionTargetPageId="33" captionText="FIGURE 164. Known distribution of Goliathopsis Janson, 1881 species. Syncretic markers indicate identical or very close localities; type localities of the known species are marked, that of G. despectus (Westwood, 1873), G. cervus Janson, 1881, and G. esquiroli Pouillaude, 1913 are missing because the precise type localities are unknown." figureDoi="http://doi.org/10.5281/zenodo.3884777" httpUri="https://zenodo.org/record/3884777/files/figure.png" pageId="4" pageNumber="95">164</figureCitation>
)
</paragraph>
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<treatmentCitation id="0A8410C0CE03FFB9FF0BF933FDAFF90A" author="Janson, O. E." box="[151,541,1752,1777]" page="610" pageId="4" pageNumber="95" year="1881">
<taxonomicName id="4C254D52CE03FFB9FF0BF933FDAFF90A" authority="Janson, 1881: 610" authorityName="in Janson" authorityPageNumber="610" authorityYear="1881" box="[151,541,1752,1777]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE03FFB9FF0BF933FED2F90A" box="[151,352,1753,1777]" italics="true" pageId="4" pageNumber="95">Goliathopsis cervus</emphasis>
<bibRefCitation id="EFB44B20CE03FFB9FEFAF930FDAFF90A" author="Janson, O. E." box="[358,541,1752,1777]" pageId="4" pageNumber="95" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881: 610</bibRefCitation>
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(type locality:
<collectingCountry id="F3327641CE03FFB9FD29F932FD5BF90A" box="[693,745,1752,1777]" name="Thailand" pageId="4" pageNumber="95">Siam</collectingCountry>
[=
<collectingCountry id="F3327641CE03FFB9FC9FF932FCD7F90A" box="[771,869,1752,1777]" name="Thailand" pageId="4" pageNumber="95">Thailand</collectingCountry>
]), plate XI, fig.
<specimenCount id="9D23FD58CE03FFB9FB9EF933FB9BF90A" box="[1026,1065,1753,1777]" pageId="4" pageNumber="95" type="male">4 ♂</specimenCount>
,
<specimenCount id="9D23FD58CE03FFB9FBAFF932FBE8F90A" box="[1075,1114,1752,1777]" pageId="4" pageNumber="95" type="female">5 ♀</specimenCount>
;
<treatmentCitation id="0A8410C0CE03FFB9FBF9F933FAA2F90A" author="Berge, A." box="[1125,1296,1752,1777]" page="158" pageId="4" pageNumber="95" year="1884">
<bibRefCitation id="EFB44B20CE03FFB9FBF9F933FAA2F90A" author="Berge, A." box="[1125,1296,1752,1777]" pageId="4" pageNumber="95" pagination="113 - 163" refId="ref19490" refString="Berge, A. (1884) Enumeration des Cetonides decrits depuis la publication du Catalogue de MM. Gemminger &amp; de Harold. An- nales de la Societe Entomologique de Belgique, 28, 113 - 163." type="journal article" year="1884">Bergé 1884: 158</bibRefCitation>
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;
<treatmentCitation id="0A8410C0CE03FFB9FA85F933FF5CF8EB" author="Gestro, R." page="119" pageId="4" pageNumber="95" year="1888">
<bibRefCitation id="EFB44B20CE03FFB9FA85F933FF5CF8EB" author="Gestro, R." pageId="4" pageNumber="95" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro 1888: 119</bibRefCitation>
</treatmentCitation>
, fig.
<specimenCount id="9D23FD58CE03FFB9FEB9F912FE8AF8F4" box="[293,312,1784,1807]" pageId="4" pageNumber="95" type="male"></specimenCount>
(horn);
<treatmentCitation id="0A8410C0CE03FFB9FE13F91DFDC3F8EB" author="Fairmaire, L." box="[399,625,1783,1808]" page="316" pageId="4" pageNumber="95" year="1893">
<bibRefCitation id="EFB44B20CE03FFB9FE13F91DFDC3F8EB" author="Fairmaire, L." box="[399,625,1783,1808]" pageId="4" pageNumber="95" pagination="303 - 325" refId="ref20102" refString="Fairmaire, L. (1893 b) Coleopteres du Haut Tokin. Annales de la Societe Entomologique de Belgique, 37, 303 - 325." type="journal article" year="1893">Fairmaire 1893b: 316</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410C0CE03FFB9FDE0F912FC9CF8EB" author="Arrow, G. J." box="[636,814,1783,1808]" page="206" pageId="4" pageNumber="95" year="1910">
<bibRefCitation id="EFB44B20CE03FFB9FDE0F912FC9CF8EB" author="Arrow, G. J." box="[636,814,1783,1808]" pageId="4" pageNumber="95" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow 1910: 206</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="4C254D52CE03FFB9FB9FF912FB34F8EB" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[1027,1158,1784,1808]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE03FFB9FB9FF912FB34F8EB" box="[1027,1158,1784,1808]" italics="true" pageId="4" pageNumber="95">G. despectus</emphasis>
</taxonomicName>
);
<treatmentCitation id="0A8410C0CE03FFB9FB02F912FAE0F8EB" author="Janson, O. E." box="[1182,1362,1784,1808]" page="110" pageId="4" pageNumber="95" year="1917">
<bibRefCitation id="EFB44B20CE03FFB9FB02F912FAE0F8EB" author="Janson, O. E." box="[1182,1362,1784,1808]" pageId="4" pageNumber="95" pagination="102 - 111" refId="ref20489" refString="Janson, O. E. (1917) Additions to the knowledge of the Cetoniinae of British India. Transactions of the Entomological Society of London, 65 (1), 102 - 111. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01404. x" type="journal article" year="1917">Janson 1917: 110</bibRefCitation>
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(status revalidated);
<treatmentCitation id="0A8410C0CE03FFB9FED1F8FCFD99F8D4" author="Schenkling, S." box="[333,555,1814,1839]" page="364" pageId="4" pageNumber="95" year="1921">
<bibRefCitation id="EFB44B20CE03FFB9FED1F8FCFD99F8D4" author="Schenkling, S." box="[333,555,1814,1839]" pageId="4" pageNumber="95" pagination="1 - 431" refId="ref21671" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, pp. 1 - 431 pp. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1921">Schenkling 1921: 364</bibRefCitation>
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(in synonymy
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<emphasis id="B951EAC3CE03FFB9FD5FF8FDFCF6F8D5" box="[707,836,1814,1838]" italics="true" pageId="4" pageNumber="95">G. despectus</emphasis>
</taxonomicName>
);
<treatmentCitation id="0A8410C0CE03FFB9FCC6F8FCFBABF8D4" author="Krikken, J." box="[858,1049,1814,1839]" page="314" pageId="4" pageNumber="95" year="1977">
<bibRefCitation id="EFB44B20CE03FFB9FCC6F8FCFBABF8D4" author="Krikken, J." box="[858,1049,1814,1839]" pageId="4" pageNumber="95" pagination="309 - 320" refId="ref20674" refString="Krikken, J. (1977). Notes on Asian cremastochiliform genera, with descriptions of two new species (Coleoptera: Cetoniidae). Zoologische Mededelingen, 50 (21), 309 - 320." type="journal article" year="1977">Krikken 1977: 314</bibRefCitation>
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(
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designated);
<treatmentCitation id="0A8410C0CE03FFB9FA97F8FCFF66F8B6" author="Antoine, P." page="9" pageId="4" pageNumber="95" year="1991">
<bibRefCitation id="EFB44B20CE03FFB9FA97F8FCFF66F8B6" author="Antoine, P." pageId="4" pageNumber="95" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine 1991: 9</bibRefCitation>
</treatmentCitation>
(status resurrected);
<treatmentCitation id="0A8410C0CE03FFB9FE35F8DEFDE5F8B6" author="Krajcik, M." box="[425,599,1844,1869]" page="39" pageId="4" pageNumber="95" year="1999">
<bibRefCitation id="EFB44B20CE03FFB9FE35F8DEFDE5F8B6" author="Krajcik, M." box="[425,599,1844,1869]" pageId="4" pageNumber="95" refId="ref20545" refString="Krajcik, M. (1999) Cetoniidae of the World Catalogue. Part II. Typos Studio, Most, 72 pp." type="book" year="1999">Krajčík 1999: 39</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A8410C0CE03FFB9FDFEF8DEFCAEF8B6" author="Krajcik, M." box="[610,796,1844,1869]" page="116" pageId="4" pageNumber="95" year="2012">
<bibRefCitation id="EFB44B20CE03FFB9FDFEF8DEFCAEF8B6" author="Krajcik, M." box="[610,796,1844,1869]" pageId="4" pageNumber="95" pagination="1 - 278" refId="ref20603" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 116</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE03FFB8FF0BF896FCABF8BD" lastPageId="5" lastPageNumber="96" pageId="4" pageNumber="95" type="materials_examined">
<paragraph id="8B9A36D1CE03FFB9FF0BF896FA2FF825" blockId="4.[151,1437,1915,2014]" pageId="4" pageNumber="95">
<emphasis id="B951EAC3CE03FFB9FF0BF896FEF6F86D" bold="true" box="[151,324,1915,1942]" pageId="4" pageNumber="95">Type material.</emphasis>
As
<bibRefCitation id="EFB44B20CE03FFB9FEF2F896FDB9F86D" author="Janson, O. E." box="[366,523,1916,1942]" pageId="4" pageNumber="95" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson (1881)</bibRefCitation>
stated in the original publication, this species was originally described from
<collectingCountry id="F3327641CE03FFB9FAFCF891FF75F842" name="Thailand" pageId="4" pageNumber="95">Thailand</collectingCountry>
based on a pair of specimens. The type specimens now are preserved in
<collectionCode id="ED34AE14CE03FFB9FC65F84AFBE2F841" box="[1017,1104,1952,1978]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="95" type="Museum">RMNH</collectionCode>
, and the male was designated as
<typeStatus id="549E8873CE03FFB9FF29F829FEAEF826" box="[181,284,1987,2013]" pageId="4" pageNumber="95" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EFB44B20CE03FFB9FED9F829FE46F825" author="Krikken, J." box="[325,500,1987,2014]" pageId="4" pageNumber="95" pagination="309 - 320" refId="ref20674" refString="Krikken, J. (1977). Notes on Asian cremastochiliform genera, with descriptions of two new species (Coleoptera: Cetoniidae). Zoologische Mededelingen, 50 (21), 309 - 320." type="journal article" year="1977">Krikken (1977)</bibRefCitation>
. Due to
<collectionCode id="ED34AE14CE03FFB9FDCDF82EFD17F825" box="[593,677,1988,2014]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="95" type="Museum">RMNH</collectionCode>
is still closed for collection integration (Jan Krikken, personal com-
</paragraph>
<paragraph id="8B9A36D1CE02FFB8FF0BFF73FB05FF2C" blockId="5.[151,1436,153,215]" pageId="5" pageNumber="96">
<materialsCitation id="3B4D3C8CCE02FFB8FF0BFF73FB06FF2C" country="Thailand" location="Type" pageId="5" pageNumber="96" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
munication, 2019), the female
<typeStatus id="549E8873CE02FFB8FE70FF73FD31FF48" box="[492,643,153,179]" pageId="5" pageNumber="96" type="paralectotype">paralectotype</typeStatus>
is unavailable for the present work, and the male
<emphasis id="B951EAC3CE02FFB8FB30FF73FAA5FF48" bold="true" box="[1196,1303,153,179]" pageId="5" pageNumber="96">
<typeStatus id="549E8873CE02FFB8FB30FF73FAA5FF48" box="[1196,1303,153,179]" pageId="5" pageNumber="96" type="lectotype">lectotype</typeStatus>
</emphasis>
was simply examined by photographs (
<figureCitation id="131E2A54CE02FFB8FE54FF57FDDDFF2C" box="[456,623,189,215]" captionStart="FIGURES 103108" captionStartId="24.[151,264,1020,1045]" captionTargetBox="[154,1430,184,982]" captionTargetId="figure@24.[151,1435,181,994]" captionTargetPageId="24" captionText="FIGURES 103108. Male lectotype of Goliathopsis cervus Janson, 1881 in RMNH (photograph by Jan Krikken). 103, dorsal view; 104, ventral view; 105, lateral view; 106, oblique view; 107, pygidium; 108, labels." figureDoi="http://doi.org/10.5281/zenodo.3884755" httpUri="https://zenodo.org/record/3884755/files/figure.png" pageId="5" pageNumber="96">Figs. 103108</figureCitation>
):
<taxonomicName id="4C254D52CE02FFB8FD1DFF57FC15FF2C" authority=", Jans." authorityName="Jans." box="[641,935,189,215]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="cervus">Goliathopsis cervus, Jans.</taxonomicName>
,
<location id="8EFA600ACE02FFB8FC2DFF57FC59FF2C" LSID="urn:lsid:plazi:treatment:038C87C7CE03FFBAFF0BF99BFAECFE54:8EFA600ACE02FFB8FC2DFF57FC59FF2C" box="[945,1003,189,215]" country="Thailand" name="Type" pageId="5" pageNumber="96">Type</location>
,
<collectingCountry id="F3327641CE02FFB8FC6BFF57FB83FF2C" box="[1015,1073,189,215]" name="Thailand" pageId="5" pageNumber="96">Siam</collectingCountry>
//
<specimenCount id="9D23FD58CE02FFB8FBD2FF57FBD1FF2C" box="[1102,1123,189,215]" pageId="5" pageNumber="96" type="male"></specimenCount>
TYPE
</materialsCitation>
.
</paragraph>
<caption id="DF5A6659CE02FFB8FF0BF9E7FE1EF969" ID-DOI="http://doi.org/10.5281/zenodo.3884729" ID-Zenodo-Dep="3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="5" pageNumber="96" startId="5.[151,264,1549,1574]" targetBox="[177,1420,257,1500]" targetPageId="5">
<paragraph id="8B9A36D1CE02FFB8FF0BF9E7FE1EF969" blockId="5.[151,1437,1549,1682]" pageId="5" pageNumber="96">
<emphasis id="B951EAC3CE02FFB8FF0BF9E7FEE5F9DD" bold="true" box="[151,343,1549,1574]" pageId="5" pageNumber="96">FIGURES 1321.</emphasis>
Cephalic horns of
<taxonomicName id="4C254D52CE02FFB8FDB8F9E4FC85F9DD" authority="Janson, 1881" authorityName="Janson" authorityYear="1881" box="[548,823,1550,1574]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAC3CE02FFB8FDB8F9E4FD14F9DD" box="[548,678,1550,1574]" italics="true" pageId="5" pageNumber="96">Goliathopsis</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FD33F9E4FC85F9DD" author="Janson, O. E." box="[687,823,1550,1574]" pageId="5" pageNumber="96" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
</taxonomicName>
species (dorsal view). 13,
<taxonomicName id="4C254D52CE02FFB8FBC9F9E4FAFFF9DD" authority="Janson, 1881" authorityName="in Janson" authorityYear="1881" box="[1109,1357,1550,1574]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE02FFB8FBC9F9E4FB0BF9DE" box="[1109,1209,1550,1573]" italics="true" pageId="5" pageNumber="96">G. cervus</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FB5FF9E4FAFFF9DD" author="Janson, O. E." box="[1219,1357,1550,1574]" pageId="5" pageNumber="96" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
</taxonomicName>
; 14,
<taxonomicName id="4C254D52CE02FFB8FA18F9E4FE0DF9B1" authority="(Westwood, 1873)" baseAuthorityName="Westwood" baseAuthorityYear="1873" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE02FFB8FA18F9E4FF4BF9B1" italics="true" pageId="5" pageNumber="96">G. despectus</emphasis>
(
<bibRefCitation id="EFB44B20CE02FFB8FE9BF9DBFE05F9B1" author="Westwood, J. O." box="[263,439,1585,1610]" pageId="5" pageNumber="96" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
)
</taxonomicName>
, paralectotype of
<taxonomicName id="4C254D52CE02FFB8FDE9F9D8FC30F9B1" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" box="[629,898,1586,1610]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE02FFB8FDE9F9D8FD45F9B1" box="[629,759,1586,1610]" italics="true" pageId="5" pageNumber="96">G. capreolus</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FD62F9D8FC30F9B1" author="Gestro, R." box="[766,898,1586,1610]" pageId="5" pageNumber="96" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro, 1888</bibRefCitation>
</taxonomicName>
in MNHN; 15,
<taxonomicName id="4C254D52CE02FFB8FBBAF9D8FA9CF9B1" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[1062,1326,1585,1610]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="duponti">
<emphasis id="B951EAC3CE02FFB8FBBAF9D8FB21F9B1" box="[1062,1171,1586,1610]" italics="true" pageId="5" pageNumber="96">G. duponti</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FB05F9DBFA9CF9B1" author="Antoine, P." box="[1177,1326,1585,1610]" pageId="5" pageNumber="96" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
</taxonomicName>
; 16,
<emphasis id="B951EAC3CE02FFB8FAC3F9D8FF69F995" italics="true" pageId="5" pageNumber="96">G. es- quiroli</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FF7EF9BFFE20F995" author="Pouillaude, I." box="[226,402,1621,1646]" pageId="5" pageNumber="96" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
; 1718,
<taxonomicName id="4C254D52CE02FFB8FE75F9BCFD5DF995" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[489,751,1621,1646]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE02FFB8FE75F9BCFDE7F995" box="[489,597,1622,1646]" italics="true" pageId="5" pageNumber="96">G. ferreroi</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FDC7F9BFFD5DF995" author="Antoine, P." box="[603,751,1621,1646]" pageId="5" pageNumber="96" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
</taxonomicName>
; 19,
<taxonomicName id="4C254D52CE02FFB8FCBCF9BCFBA0F995" authority="Ruter, 1978" authorityName="Ruter" authorityYear="1978" box="[800,1042,1622,1646]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B951EAC3CE02FFB8FCBCF9BCFC23F996" box="[800,913,1622,1645]" italics="true" pageId="5" pageNumber="96">G. gressitti</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FC0BF9BCFBA0F995" author="Ruter, G." box="[919,1042,1622,1646]" pageId="5" pageNumber="96" pagination="285 - 287" refId="ref21505" refString="Ruter, G. (1978) Description de Cetonides nouveaux. Nouvelle Revue d'Entomologie, 8 (3), 285 - 287." type="journal article" year="1978">Ruter, 1978</bibRefCitation>
</taxonomicName>
; 20,
<taxonomicName id="4C254D52CE02FFB8FBDFF9BCFAE1F995" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[1091,1363,1621,1646]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="lameyi">
<emphasis id="B951EAC3CE02FFB8FBDFF9BCFB14F995" box="[1091,1190,1622,1646]" italics="true" pageId="5" pageNumber="96">G. lameyi</emphasis>
Fairmaire, 1893
</taxonomicName>
; 21,
<taxonomicName id="4C254D52CE02FFB8FA18F9BCFE1AF969" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE02FFB8FA18F9BCFF40F969" italics="true" pageId="5" pageNumber="96">G. velutinus</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FF65F993FE1AF969" author="Pouillaude, I." box="[249,424,1657,1682]" pageId="5" pageNumber="96" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9A36D1CE02FFB8FF5BF955FCABF8BD" blockId="5.[151,1437,1727,2042]" pageId="5" pageNumber="96">
<emphasis id="B951EAC3CE02FFB8FF5BF955FE49F921" bold="true" box="[199,507,1727,1754]" pageId="5" pageNumber="96">Other material examined</emphasis>
(
<specimenCount id="9D23FD58CE02FFB8FD92F92AFDFBF921" box="[526,585,1728,1754]" pageId="5" pageNumber="96" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D23FD58CE02FFB8FDCBF92AFD3CF921" box="[599,654,1728,1754]" pageId="5" pageNumber="96" type="female">1♀♀</specimenCount>
).
<emphasis id="B951EAC3CE02FFB8FD35F955FCF2F921" bold="true" box="[681,832,1727,1754]" pageId="5" pageNumber="96">
<collectingCountry id="F3327641CE02FFB8FD35F955FCF2F921" box="[681,832,1727,1754]" name="Thailand" pageId="5" pageNumber="96">THAILAND</collectingCountry>
</emphasis>
:
<materialsCitation id="3B4D3C8CCE02FFB8FCCEF92AFF7DF8D9" collectingDate="2012-05-22" collectionCode="KSCJ" collectorName="Petchaburi Prov. C. Thailand &amp; S. Tsuyuki" country="Thailand" determinerName="K. Sakai" location="Petchaburi Prov." municipality="Khao Bandai It" pageId="5" pageNumber="96" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D23FD58CE02FFB8FCCEF92AFC39F921" box="[850,907,1728,1754]" pageId="5" pageNumber="96" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED34AE14CE02FFB8FC01F92AFC56F921" box="[925,996,1728,1754]" pageId="5" pageNumber="96">KSCJ</collectionCode>
),
<collectingMunicipality id="6BFEACABCE02FFB8FC66F92AFB1CF921" box="[1018,1198,1728,1754]" pageId="5" pageNumber="96">Khao Bandai It</collectingMunicipality>
,
<collectorName id="26D05307CE02FFB8FB20F92AFF4FF905" pageId="5" pageNumber="96">
<location id="8EFA600ACE02FFB8FB20F92AFACBF921" LSID="urn:lsid:plazi:treatment:038C87C7CE03FFBAFF0BF99BFAECFE54:8EFA600ACE02FFB8FB20F92AFACBF921" box="[1212,1401,1728,1754]" country="Thailand" municipality="Khao Bandai It" name="Petchaburi Prov." pageId="5" pageNumber="96">Petchaburi Prov.</location>
C. Thailand
</collectorName>
,
<date id="FF9B1011CE02FFB8FE95F90EFECCF905" box="[265,382,1764,1790]" pageId="5" pageNumber="96" value="2012-05-22">
<collectingDate id="EFDFE9F9CE02FFB8FE95F90EFECCF905" box="[265,382,1764,1790]" pageId="5" pageNumber="96" value="2012-05-22">2012.V.22</collectingDate>
</date>
,
<collectorName id="26D05307CE02FFB8FE16F90EFE49F905" box="[394,507,1764,1790]" pageId="5" pageNumber="96">S.Tsuyuki</collectorName>
leg. //
<taxonomicName id="4C254D52CE02FFB8FDD5F90EFBBFF905" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" box="[585,1037,1764,1790]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE02FFB8FDD5F90EFCF7F905" box="[585,837,1764,1790]" italics="true" pageId="5" pageNumber="96">Goliathopsis velutinus</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FCD0F90EFBBFF905" author="Pouillaude, I." box="[844,1037,1764,1790]" pageId="5" pageNumber="96" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
</taxonomicName>
, det.
<determinerName id="7E46DA15CE02FFB8FBD5F90EFB1FF905" box="[1097,1197,1764,1790]" pageId="5" pageNumber="96">K. Sakai</determinerName>
, 2015 [misidentification]
</materialsCitation>
;
<materialsCitation id="3B4D3C8CCE02FFB8FF47F8E2FCA7F8BD" collectingDate="2012-05-17" collectingDateMax="2012-05-21" collectingDateMin="2012-05-17" collectionCode="KSCJ" collectorName="K. Takahashi" country="Thailand" determinerName="K. Sakai" location="Thailand" municipality="Khao Soi Dao" pageId="5" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="Chanthaburi">
<specimenCount id="9D23FD58CE02FFB8FF47F8E2FF4CF8D9" box="[219,254,1800,1826]" pageId="5" pageNumber="96" type="female">1♀</specimenCount>
(
<collectionCode id="ED34AE14CE02FFB8FE91F8E2FEE6F8D9" box="[269,340,1800,1826]" pageId="5" pageNumber="96">KSCJ</collectionCode>
),
<collectingMunicipality id="6BFEACABCE02FFB8FEFBF8E2FDBAF8D9" box="[359,520,1800,1826]" pageId="5" pageNumber="96">Khao Soi Dao</collectingMunicipality>
,
<collectingRegion id="49E1F833CE02FFB8FD8FF8E2FD12F8D9" box="[531,672,1800,1826]" country="Thailand" name="Chanthaburi" pageId="5" pageNumber="96">Chanthaburi</collectingRegion>
,
<location id="8EFA600ACE02FFB8FD37F8E2FCA3F8D9" LSID="urn:lsid:plazi:treatment:038C87C7CE03FFBAFF0BF99BFAECFE54:8EFA600ACE02FFB8FD37F8E2FCA3F8D9" box="[683,785,1800,1826]" country="Thailand" municipality="Khao Soi Dao" name="Thailand" pageId="5" pageNumber="96" stateProvince="Chanthaburi">Thailand</location>
,
<date id="FF9B1011CE02FFB8FC81F8E2FC0EF8D9" box="[797,956,1800,1826]" pageId="5" pageNumber="96" value="2012-05-17" valueMax="2012-05-21" valueMin="2012-05-17">
<collectingDate id="EFDFE9F9CE02FFB8FC81F8E2FC0EF8D9" box="[797,956,1800,1826]" pageId="5" pageNumber="96" value="2012-05-17" valueMax="2012-05-21" valueMin="2012-05-17">2012.V.1721</collectingDate>
</date>
,
<collectorName id="26D05307CE02FFB8FC5BF8E2FBE1F8D9" box="[967,1107,1800,1826]" pageId="5" pageNumber="96">K.Takahashi</collectorName>
leg. //
<taxonomicName id="4C254D52CE02FFB8FB3DF8E2FEEAF8BD" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="96" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE02FFB8FB3DF8E2FA2EF8D9" box="[1185,1436,1800,1826]" italics="true" pageId="5" pageNumber="96">Goliathopsis velutinus</emphasis>
<bibRefCitation id="EFB44B20CE02FFB8FF0BF8C6FEEAF8BD" author="Pouillaude, I." box="[151,344,1836,1862]" pageId="5" pageNumber="96" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
</taxonomicName>
, det.
<determinerName id="7E46DA15CE02FFB8FE0FF8C6FE44F8BD" box="[403,502,1836,1862]" pageId="5" pageNumber="96">K. Sakai</determinerName>
, 2015 [misidentification]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE02FFBBFF5BF8A5FC6EF8BA" lastPageId="6" lastPageNumber="97" pageId="5" pageNumber="96" type="diagnosis">
<paragraph id="8B9A36D1CE02FFBBFF5BF8A5FE67FEE4" blockId="5.[151,1437,1727,2042]" lastBlockId="6.[151,1436,153,287]" lastPageId="6" lastPageNumber="97" pageId="5" pageNumber="96">
<emphasis id="B951EAC3CE02FFB8FF5BF8A5FEF0F891" bold="true" box="[199,322,1871,1898]" pageId="5" pageNumber="96">Diagnosis.</emphasis>
Smallest species of the genus, body length 9.510.5 mm; width 5.35.5 mm. Dorsal surface and pygidium clad with brown tomentum (
<figureCitation id="131E2A54CE02FFB8FDDAF89EFD7FF875" box="[582,717,1907,1934]" captionStart="FIGURES 5457" captionStartId="8.[151,264,1902,1927]" captionTargetBox="[186,1367,278,1868]" captionTargetId="figure@8.[151,1435,266,1878]" captionTargetPageId="8" captionText="FIGURES 5457. Habitus of Goliathopsis cervus Janson, 1881 (dorsal and ventral view). 5455, male; 5657, female." figureDoi="http://doi.org/10.5281/zenodo.3884735" httpUri="https://zenodo.org/record/3884735/files/figure.png" pageId="5" pageNumber="96">Figs. 54, 56</figureCitation>
,
<figureCitation id="131E2A54CE02FFB8FD45F89EFC8DF874" box="[729,831,1908,1935]" captionStart="FIGURES 103108" captionStartId="24.[151,264,1020,1045]" captionTargetBox="[154,1430,184,982]" captionTargetId="figure@24.[151,1435,181,994]" captionTargetPageId="24" captionText="FIGURES 103108. Male lectotype of Goliathopsis cervus Janson, 1881 in RMNH (photograph by Jan Krikken). 103, dorsal view; 104, ventral view; 105, lateral view; 106, oblique view; 107, pygidium; 108, labels." figureDoi="http://doi.org/10.5281/zenodo.3884755" httpUri="https://zenodo.org/record/3884755/files/figure.png" pageId="5" pageNumber="96">103, 106</figureCitation>
). Midline on pronotum and scutellum yellow, distinct. Elytron usually with two light brown maculae (not distinct): a small one on posthumeral area, sometimes absent; the other small one on the mediodiscal area near sutural costa. Disc of elytron with U-shaped (few drop-shaped), setiferous punctures (
<figureCitation id="131E2A54CE02FFB8FEF5F80AFE1FF801" box="[361,429,2015,2042]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="5" pageNumber="96">Fig. 5</figureCitation>
); setae short. Pygidium with small, rounded, setiferous punctures (
<figureCitation id="131E2A54CE02FFB8FB00F80AFA94F801" box="[1180,1318,2016,2042]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="5" pageNumber="96">Figs. 2223</figureCitation>
). Cephalic horns of male small and slender, inner side with tomentum, not constricted at base and not distinctly expanded at apex (
<figureCitation id="131E2A54CE01FFBBFF47FF57FE85FF2C" box="[219,311,189,215]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="6" pageNumber="97">Figs. 13</figureCitation>
,
<figureCitation id="131E2A54CE01FFBBFEDFFF57FEC3FF2C" box="[323,369,189,215]" captionStart="FIGURES 103108" captionStartId="24.[151,264,1020,1045]" captionTargetBox="[154,1430,184,982]" captionTargetId="figure@24.[151,1435,181,994]" captionTargetPageId="24" captionText="FIGURES 103108. Male lectotype of Goliathopsis cervus Janson, 1881 in RMNH (photograph by Jan Krikken). 103, dorsal view; 104, ventral view; 105, lateral view; 106, oblique view; 107, pygidium; 108, labels." figureDoi="http://doi.org/10.5281/zenodo.3884755" httpUri="https://zenodo.org/record/3884755/files/figure.png" pageId="6" pageNumber="97">103</figureCitation>
). Male genitalia small (
<figureCitation id="131E2A54CE01FFBBFDE9FF57FD4FFF2C" box="[629,765,188,215]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="6" pageNumber="97">Figs. 5253</figureCitation>
), proximal and distal parts of parameres gradually expanded in apical view, medially constricted, apex rounded and inner side slightly curved; parameres slightly curved in lateral view, basal piece broad.
</paragraph>
<caption id="DF5A6659CE01FFBBFF0BF9EDFC4DF977" ID-DOI="http://doi.org/10.5281/zenodo.3884731" ID-Zenodo-Dep="3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="6" pageNumber="97" startId="6.[151,264,1543,1568]" targetBox="[156,1430,332,1511]" targetPageId="6">
<paragraph id="8B9A36D1CE01FFBBFF0BF9EDFC4DF977" blockId="6.[151,1437,1543,1676]" pageId="6" pageNumber="97">
<emphasis id="B951EAC3CE01FFBBFF0BF9EDFEE6F9DB" bold="true" box="[151,340,1543,1568]" pageId="6" pageNumber="97">FIGURES 2237.</emphasis>
Pygidium of
<taxonomicName id="4C254D52CE01FFBBFE40F9E2FD5AF9DB" authority="Janson, 1881" authorityName="Janson" authorityYear="1881" box="[476,744,1544,1568]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="B951EAC3CE01FFBBFE40F9E2FDECF9DB" box="[476,606,1544,1568]" italics="true" pageId="6" pageNumber="97">Goliathopsis</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFDF8F9E3FD5AF9DB" author="Janson, O. E." box="[612,744,1544,1568]" pageId="6" pageNumber="97" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
</taxonomicName>
species (male in left, female in right; posterior view). 2223,
<emphasis id="B951EAC3CE01FFBBFAC9F9E3FF0BF9BF" italics="true" pageId="6" pageNumber="97">G. cer- vus</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFF23F9C7FEF4F9BF" author="Janson, O. E." box="[191,326,1580,1604]" pageId="6" pageNumber="97" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson, 1881</bibRefCitation>
; 2425,
<taxonomicName id="4C254D52CE01FFBBFE00F9C7FD52F9BF" authority="(Westwood, 1873)" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[412,736,1579,1604]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE01FFBBFE00F9C7FDAFF9BF" box="[412,541,1580,1604]" italics="true" pageId="6" pageNumber="97">G. despectus</emphasis>
(
<bibRefCitation id="EFB44B20CE01FFBBFDB6F9C1FD6BF9BF" author="Westwood, J. O." box="[554,729,1579,1604]" pageId="6" pageNumber="97" refId="ref21916" refString="Westwood, J. O. (1873) Thesaurus Entomologicus Oxoniensis; or, Illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope. Part I. Clarendon Press, Oxford, 56 pp., 10 pls. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1873">Westwood, 1873</bibRefCitation>
)
</taxonomicName>
, lectotype and paralectotype of
<taxonomicName id="4C254D52CE01FFBBFBBEF9C7FA9EF9BF" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" box="[1058,1324,1580,1604]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE01FFBBFBBEF9C7FB11F9BF" box="[1058,1187,1580,1604]" italics="true" pageId="6" pageNumber="97">G. capreolus</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFB35F9C6FA9EF9BF" author="Gestro, R." box="[1193,1324,1580,1604]" pageId="6" pageNumber="97" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro, 1888</bibRefCitation>
</taxonomicName>
in MSNG; 2627,
<taxonomicName id="4C254D52CE01FFBBFF78F9BBFE5EF993" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[228,492,1615,1640]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="duponti">
<emphasis id="B951EAC3CE01FFBBFF78F9BBFEE3F993" box="[228,337,1616,1640]" italics="true" pageId="6" pageNumber="97">G. duponti</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFECBF9A5FE5EF993" author="Antoine, P." box="[343,492,1615,1640]" pageId="6" pageNumber="97" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
</taxonomicName>
; 2829,
<taxonomicName id="4C254D52CE01FFBBFDD8F9BBFCC6F993" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" box="[580,884,1615,1640]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="esquiroli">
<emphasis id="B951EAC3CE01FFBBFDD8F9BBFD0FF993" box="[580,701,1616,1640]" italics="true" pageId="6" pageNumber="97">G. esquiroli</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFD58F9A5FCC6F993" author="Pouillaude, I." box="[708,884,1615,1640]" pageId="6" pageNumber="97" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
</taxonomicName>
; 3031,
<taxonomicName id="4C254D52CE01FFBBFC50F9BBFB66F993" authority="Antoine, 1991" authorityName="Antoine" authorityYear="1991" box="[972,1236,1615,1640]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="ferreroi">
<emphasis id="B951EAC3CE01FFBBFC50F9BBFB8BF993" box="[972,1081,1616,1640]" italics="true" pageId="6" pageNumber="97">G. ferreroi</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFBA3F9A5FB66F993" author="Antoine, P." box="[1087,1236,1615,1640]" pageId="6" pageNumber="97" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine, 1991</bibRefCitation>
</taxonomicName>
; 3233,
<taxonomicName id="4C254D52CE01FFBBFAB0F9BBFEA3F977" authority="Ruter, 1978" authorityName="Ruter" authorityYear="1978" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="B951EAC3CE01FFBBFAB0F9BBFA2FF993" box="[1324,1437,1617,1640]" italics="true" pageId="6" pageNumber="97">G. gressitti</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFF0BF99FFEA3F977" author="Ruter, G." box="[151,273,1652,1676]" pageId="6" pageNumber="97" pagination="285 - 287" refId="ref21505" refString="Ruter, G. (1978) Description de Cetonides nouveaux. Nouvelle Revue d'Entomologie, 8 (3), 285 - 287." type="journal article" year="1978">Ruter, 1978</bibRefCitation>
</taxonomicName>
; 3435,
<taxonomicName id="4C254D52CE01FFBBFEFBF99FFDC7F977" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[359,629,1651,1676]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="lameyi">
<emphasis id="B951EAC3CE01FFBBFEFBF99FFE78F977" box="[359,458,1652,1676]" italics="true" pageId="6" pageNumber="97">G. lameyi</emphasis>
Fairmaire, 1893
</taxonomicName>
; 3637,
<taxonomicName id="4C254D52CE01FFBBFD57F99FFC49F977" authority="Pouillaude, 1913" authorityName="Pouillaude" authorityYear="1913" box="[715,1019,1651,1676]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE01FFBBFD57F99FFCF7F977" box="[715,837,1652,1676]" italics="true" pageId="6" pageNumber="97">G. velutinus</emphasis>
<bibRefCitation id="EFB44B20CE01FFBBFCD0F999FC49F977" author="Pouillaude, I." box="[844,1019,1651,1676]" pageId="6" pageNumber="97" pagination="421 - 427" refId="ref21344" refString="Pouillaude, I. (1913) Description de trois nouvelles espece de Cetonides. Insecta, 3, 421 - 427." type="journal article" year="1913">Pouillaude, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9A36D1CE01FFBBFF5BF950FC6EF8BA" blockId="6.[151,1436,1722,2037]" pageId="6" pageNumber="97">
This species is closed to
<taxonomicName id="4C254D52CE01FFBBFE74F951FDC3F92F" box="[488,625,1722,1749]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE01FFBBFE74F951FDC3F92F" box="[488,625,1722,1749]" italics="true" pageId="6" pageNumber="97">G. velutinus</emphasis>
</taxonomicName>
, but body size distinctly smaller (
<figureCitation id="131E2A54CE01FFBBFC66F950FB34F92E" box="[1018,1158,1722,1749]" captionStart="FIGURES 89102" captionStartId="23.[151,264,1958,1983]" captionTargetBox="[222,1369,191,1931]" captionTargetId="figure@23.[200,1387,181,1940]" captionTargetPageId="23" captionText="FIGURES 89102. Habitus of Goliathopsis Janson, 1881 species (male in left, female in right; oblique view). 8990, G. cervus Janson, 1881; 9192, G. duponti Antoine, 1991; 9394, G. esquiroli Pouillaude, 1913; 9596, G. ferreroi Antoine, 1991; 9798, G. gressitti Ruter, 1978; 99100, G. lameyi Fairmaire, 1893; 101102, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884753" httpUri="https://zenodo.org/record/3884753/files/figure.png" pageId="6" pageNumber="97">Figs. 8990</figureCitation>
); cephalic horns smaller and not constricted at base (
<figureCitation id="131E2A54CE01FFBBFE51F934FDADF902" box="[461,543,1758,1785]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="6" pageNumber="97">Fig. 13</figureCitation>
; larger and constricted at base in
<taxonomicName id="4C254D52CE01FFBBFC0DF935FBAAF903" box="[913,1048,1758,1785]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE01FFBBFC0DF935FBAAF903" box="[913,1048,1758,1785]" italics="true" pageId="6" pageNumber="97">G. velutinus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE01FFBBFBB9F934FBC5F902" box="[1061,1143,1758,1785]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="6" pageNumber="97">Fig. 21</figureCitation>
); punctures on elytral disc usually U-shaped (
<figureCitation id="131E2A54CE01FFBBFEF4F8E8FE19F8E6" box="[360,427,1794,1821]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="6" pageNumber="97">Fig. 5</figureCitation>
; usually drop-shaped in
<taxonomicName id="4C254D52CE01FFBBFD26F8E9FCF3F8E7" box="[698,833,1794,1821]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE01FFBBFD26F8E9FCF3F8E7" box="[698,833,1794,1821]" italics="true" pageId="6" pageNumber="97">G. velutinus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE01FFBBFCD2F8E8FC2DF8E6" box="[846,927,1794,1821]" captionStart="FIGURES 112" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[151,1435,278,1670]" captionTargetId="figure@3.[151,1435,273,1680]" captionTargetPageId="3" captionText="FIGURES 112. Characters of Goliathopsis Janson, 1881 species. 14, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 512, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884727" httpUri="https://zenodo.org/record/3884727/files/figure.png" pageId="6" pageNumber="97">Fig. 12</figureCitation>
); parameres smaller and distal parts gradually expanded (
<figureCitation id="131E2A54CE01FFBBFE8EF8CCFED7F8BA" box="[274,357,1830,1857]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="6" pageNumber="97">Fig. 52</figureCitation>
; larger and distinctly expanded in
<taxonomicName id="4C254D52CE01FFBBFD7AF8CDFCDCF8BB" box="[742,878,1830,1857]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="velutinus">
<emphasis id="B951EAC3CE01FFBBFD7AF8CDFCDCF8BB" box="[742,878,1830,1857]" italics="true" pageId="6" pageNumber="97">G. velutinus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE01FFBBFCE0F8CCFC7DF8BA" box="[892,975,1830,1857]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="6" pageNumber="97">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE01FFBBFF5BF8A0FE61F89F" box="[199,467,1866,1893]" pageId="6" pageNumber="97" type="distribution">
<paragraph id="8B9A36D1CE01FFBBFF5BF8A0FE61F89F" blockId="6.[151,1436,1722,2037]" box="[199,467,1866,1893]" pageId="6" pageNumber="97">
<emphasis id="B951EAC3CE01FFBBFF5BF8A0FED3F89E" bold="true" box="[199,353,1866,1893]" pageId="6" pageNumber="97">Distribution.</emphasis>
<collectingCountry id="F3327641CE01FFBBFEFBF8A0FE7CF89F" box="[359,462,1866,1892]" name="Thailand" pageId="6" pageNumber="97">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F655ACE01FFBAFF5BF885FAECFE54" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97" type="discussion">
<paragraph id="8B9A36D1CE01FFBAFF5BF885FAECFE54" blockId="6.[151,1436,1722,2037]" lastBlockId="7.[151,1437,153,431]" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97">
<emphasis id="B951EAC3CE01FFBBFF5BF885FE89F872" bold="true" box="[199,315,1903,1929]" pageId="6" pageNumber="97">Remarks.</emphasis>
Based on the good illustrations of
<taxonomicName id="4C254D52CE01FFBBFD5AF884FBDEF872" authority="in Janson (1881)" authorityName="in Janson" authorityYear="1881" box="[710,1132,1902,1929]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE01FFBBFD5AF884FC14F873" box="[710,934,1902,1928]" italics="true" pageId="6" pageNumber="97">Goliathopsis cervus</emphasis>
in
<bibRefCitation id="EFB44B20CE01FFBBFC57F885FBDEF872" author="Janson, O. E." box="[971,1132,1903,1929]" pageId="6" pageNumber="97" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson (1881)</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFB44B20CE01FFBBFBE4F885FAA7F872" author="Gestro, R." box="[1144,1301,1903,1929]" pageId="6" pageNumber="97" pagination="87 - 132" refId="ref20159" refString="Gestro, R. (1888) Viaggio di Leonardo Fea in Birmania regione vicine. IV. Nuove specie di Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 6, 87 - 132." type="journal article" year="1888">Gestro (1888)</bibRefCitation>
pointed out several differences between
<taxonomicName id="4C254D52CE01FFBBFE40F879FDF8F857" authorityName="in Janson" authorityYear="1881" box="[476,586,1939,1965]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE01FFBBFE40F879FDF8F857" box="[476,586,1939,1965]" italics="true" pageId="6" pageNumber="97">G. cervus</emphasis>
</taxonomicName>
and his new species
<taxonomicName id="4C254D52CE01FFBBFCDEF879FC66F857" box="[834,980,1938,1965]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="capreolus">
<emphasis id="B951EAC3CE01FFBBFCDEF879FC66F857" box="[834,980,1938,1965]" italics="true" pageId="6" pageNumber="97">G. capreolus</emphasis>
</taxonomicName>
, especially in the shape of the cephalic horns. Nevertheless,
<bibRefCitation id="EFB44B20CE01FFBBFE1CF85DFDA8F82A" author="Arrow, G. J." box="[384,538,1975,2001]" pageId="6" pageNumber="97" refId="ref19451" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma) Coleoptera, Lamellicornia (Cetoniinae and Dy- nastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
regarded them as the same species and listed both as synonyms of
<taxonomicName id="4C254D52CE01FFBBFA9AF85DFA27F82B" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[1286,1429,1974,2001]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE01FFBBFA9AF85DFA27F82B" box="[1286,1429,1974,2001]" italics="true" pageId="6" pageNumber="97">G. despectus</emphasis>
</taxonomicName>
. In response to Arrows treatment,
<bibRefCitation id="EFB44B20CE01FFBBFD8EF831FD1DF80E" author="Janson, O. E." box="[530,687,2011,2037]" pageId="6" pageNumber="97" pagination="102 - 111" refId="ref20489" refString="Janson, O. E. (1917) Additions to the knowledge of the Cetoniinae of British India. Transactions of the Entomological Society of London, 65 (1), 102 - 111. https: // doi. org / 10.1111 / j. 1365 - 2311.1917. tb 01404. x" type="journal article" year="1917">Janson (1917)</bibRefCitation>
insisted on the validity of
<taxonomicName id="4C254D52CE01FFBBFC4AF831FBF3F80F" authorityName="in Janson" authorityYear="1881" box="[982,1089,2011,2037]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE01FFBBFC4AF831FBF3F80F" box="[982,1089,2011,2037]" italics="true" pageId="6" pageNumber="97">G. cervus</emphasis>
</taxonomicName>
and provided more differences between females based on a re-examination of the female type of
<taxonomicName id="4C254D52CE00FFBAFCF1FF73FC49FF48" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[877,1019,153,179]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFCF1FF73FC49FF48" box="[877,1019,153,179]" italics="true" pageId="7" pageNumber="98">G. despectus</emphasis>
</taxonomicName>
in OXUM. Unfortunately, this disputation was not mentioned by subsequent authors. In the most recent work,
<bibRefCitation id="EFB44B20CE00FFBAFC44FF57FB31FF2C" author="Antoine, P." box="[984,1155,189,215]" pageId="7" pageNumber="98" pagination="9 - 16" refId="ref19382" refString="Antoine, P. (1991) Quelques especes nouvelles ou peu connues de la famille des Cetoniidae II. Bulletin de la Societe Sciences Nat Compiegne, 69, 9 - 16." type="journal article" year="1991">Antoine (1991)</bibRefCitation>
also treated
<taxonomicName id="4C254D52CE00FFBAFA8EFF57FACCFF2C" authorityName="in Janson" authorityYear="1881" box="[1298,1406,189,215]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE00FFBAFA8EFF57FACCFF2C" box="[1298,1406,189,215]" italics="true" pageId="7" pageNumber="98">G. cervus</emphasis>
</taxonomicName>
as valid species mainly by the difference of the tomentose area on cephalic horns and elytra according to the figures in
<bibRefCitation id="EFB44B20CE00FFBAFF0BFEEFFE8AFEE4" author="Janson, O. E." box="[151,312,261,287]" pageId="7" pageNumber="98" pagination="603 - 611" refId="ref20457" refString="Janson, O. E. (1881) Notices of new or little known Cetoniidae No. 7. Cistula Entomologica, 2 (24), 603 - 611." type="journal article" year="1881">Janson (1881)</bibRefCitation>
. By examining the fresh specimens collected from southern
<collectingCountry id="F3327641CE00FFBAFC45FEEFFB8FFEE4" box="[985,1085,261,287]" name="Thailand" pageId="7" pageNumber="98">Thailand</collectingCountry>
and the male
<typeStatus id="549E8873CE00FFBAFB45FEEFFAF1FEE4" box="[1241,1347,261,287]" pageId="7" pageNumber="98" type="lectotype">lectotype</typeStatus>
, several distinct differences from
<taxonomicName id="4C254D52CE00FFBAFE2FFEC3FDF1FEB8" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[435,579,297,323]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFE2FFEC3FDF1FEB8" box="[435,579,297,323]" italics="true" pageId="7" pageNumber="98">G. despectus</emphasis>
</taxonomicName>
are observed: cephalic horns of male smaller and inner side with tomentum in
<taxonomicName id="4C254D52CE00FFBAFF28FEA7FE92FE9C" authorityName="in Janson" authorityYear="1881" box="[180,288,333,359]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE00FFBAFF28FEA7FE92FE9C" box="[180,288,333,359]" italics="true" pageId="7" pageNumber="98">G. cervus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A54CE00FFBAFEB3FEA7FE31FE9C" box="[303,387,333,359]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="7" pageNumber="98">Fig. 13</figureCitation>
; larger and glabrous in
<taxonomicName id="4C254D52CE00FFBAFD17FEA7FCA9FE9C" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[651,795,333,359]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFD17FEA7FCA9FE9C" box="[651,795,333,359]" italics="true" pageId="7" pageNumber="98">G. despectus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE00FFBAFCB5FEA7FCCEFE9C" box="[809,892,333,359]" captionStart="FIGURES 1321" captionStartId="5.[151,264,1549,1574]" captionTargetBox="[177,1420,257,1500]" captionTargetId="figure@5.[151,1435,234,1521]" captionTargetPageId="5" captionText="FIGURES 1321. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 1718, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884729" httpUri="https://zenodo.org/record/3884729/files/figure.png" pageId="7" pageNumber="98">Fig. 14</figureCitation>
); pygidium subtriangular in posterior view in
<taxonomicName id="4C254D52CE00FFBAFA1DFEA7FF52FE70" authorityName="in Janson" authorityYear="1881" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE00FFBAFA1DFEA7FF52FE70" italics="true" pageId="7" pageNumber="98">G. cervus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A54CE00FFBAFF73FE9BFEC5FE70" box="[239,375,369,395]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="7" pageNumber="98">Figs. 2223</figureCitation>
; oval in
<taxonomicName id="4C254D52CE00FFBAFE4AFE9BFDD7FE70" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[470,613,369,395]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFE4AFE9BFDD7FE70" box="[470,613,369,395]" italics="true" pageId="7" pageNumber="98">G. despectus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE00FFBAFDEEFE9BFD49FE70" box="[626,763,368,395]" captionStart="FIGURES 2237" captionStartId="6.[151,264,1543,1568]" captionTargetBox="[156,1430,332,1511]" captionTargetId="figure@6.[151,1435,309,1518]" captionTargetPageId="6" captionText="FIGURES 2237. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 2223, G. cer- vus Janson, 1881; 2425, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 2627, G. duponti Antoine, 1991; 2829, G. esquiroli Pouillaude, 1913; 3031, G. ferreroi Antoine, 1991; 3233, G. gressitti Ruter, 1978; 3435, G. lameyi Fairmaire, 1893; 3637, G. velutinus Pouillaude, 1913." figureDoi="http://doi.org/10.5281/zenodo.3884731" httpUri="https://zenodo.org/record/3884731/files/figure.png" pageId="7" pageNumber="98">Figs. 2425</figureCitation>
); parameres smaller and apex rounded in
<taxonomicName id="4C254D52CE00FFBAFB56FE9BFA87FE70" authorityName="in Janson" authorityYear="1881" box="[1226,1333,369,395]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE00FFBAFB56FE9BFA87FE70" box="[1226,1333,369,395]" italics="true" pageId="7" pageNumber="98">G. cervus</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A54CE00FFBAFAD8FE9BFA25FE70" box="[1348,1431,368,395]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="7" pageNumber="98">Fig. 52</figureCitation>
; larger and apex sharp in
<taxonomicName id="4C254D52CE00FFBAFE37FE7FFD88FE54" baseAuthorityName="Westwood" baseAuthorityYear="1873" box="[427,570,405,431]" class="Insecta" family="Cetoniidae" genus="Pilinurgus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="B951EAC3CE00FFBAFE37FE7FFD88FE54" box="[427,570,405,431]" italics="true" pageId="7" pageNumber="98">G. despectus</emphasis>
</taxonomicName>
,
<figureCitation id="131E2A54CE00FFBAFDD4FE7FFD29FE54" box="[584,667,405,431]" captionStart="FIGURES 3853" captionStartId="7.[151,264,1590,1615]" captionTargetBox="[151,1435,463,1559]" captionTargetId="figure@7.[151,1435,453,1565]" captionTargetPageId="7" captionText="FIGURES 3853. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 3839, G. lameyi Fairmaire, 1893; 4041, G. esquiroli Pouillaude, 1913; 4243, G. velutinus Pouillaude, 1913; 4445, G. duponti Antoine, 1991; 4647, G. ferreroi Antoine, 1991; 4849, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 5051, G. gressitti Ruter, 1978; 5253, G. cervus Janson, 1881." figureDoi="http://doi.org/10.5281/zenodo.3884733" httpUri="https://zenodo.org/record/3884733/files/figure.png" pageId="7" pageNumber="98">Fig. 48</figureCitation>
). Consequently,
<taxonomicName id="4C254D52CE00FFBAFCC9FE7FFB86FE54" authorityName="in Janson" authorityYear="1881" box="[853,1076,405,431]" class="Insecta" family="Cetoniidae" genus="Goliathopsis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="cervus">
<emphasis id="B951EAC3CE00FFBAFCC9FE7FFB86FE54" box="[853,1076,405,431]" italics="true" pageId="7" pageNumber="98">Goliathopsis cervus</emphasis>
</taxonomicName>
is an independent species.
</paragraph>
</subSubSection>
</treatment>
</document>