treatments-xml/data/03/8A/87/038A87A9FFA8FFFDE107FDD3FC693F29.xml
2024-06-21 12:22:17 +02:00

158 lines
19 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="7016091DA30956674173517701CB81C9" ID-DOI="10.11646/zootaxa.1030.1.2" ID-ISSN="1175­5334" ID-Zenodo-Dep="5050339" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625107904989" checkinUser="felipe" docAuthor="Pesic, Vladimir, Saboori, Alireza, Asadi, Mahdieh &amp; Vafaei, Reza" docDate="2005" docId="038A87A9FFA8FFFDE107FDD3FC693F29" docLanguage="en" docName="zt01030p060.pdf" docOrigin="Zootaxa 1030 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1030.1.2" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Atractides denticulatus" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="FFB3FFD1FFABFFFEE00FFFADFF863952" masterDocTitle="Water mites (Acari: Hydrachnidia) from interstitial waters of Iran, with the description of one new species" masterLastPageNumber="60" masterPageNumber="49" pageNumber="52" updateTime="1698980962083" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F6F44A521983CC9678970BCB83117E9C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="233D9D9885DF7213361F0FA12ACF5A68">
<mods:title id="B242EC22A4CB93686B22BE44383613DE">Water mites (Acari: Hydrachnidia) from interstitial waters of Iran, with the description of one new species</mods:title>
</mods:titleInfo>
<mods:name id="A255B1D480A3F72E4E3A3E8C724342A3" type="personal">
<mods:role id="DE2FBE3FFCB960BA70FB01C995EA6CC4">
<mods:roleTerm id="CB1ACAF86655B379E60AFC44E30F6677">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E442FFC742B10C5F72EC2474F6ACDF2">Pesic, Vladimir</mods:namePart>
<mods:affiliation id="39A091A6F6950C77263B5B582D5EF2B7">Department of Biology, Faculty of Sciences, University of Crna Gora, Cetinjski put b. b., 81000 Podgorica, Montenegro, Serbia and Montenegro (pesicv @ cg. yu) Department of Plant Protection, College of Agriculture, University of Tehran, Karaj, Iran Department of Entomology, College of Agriculture, Islamic Azad University, Arak, Iran</mods:affiliation>
</mods:name>
<mods:name id="4FD391FEAC2E8CC1E70E6F7419F23417" type="personal">
<mods:role id="35EF455AC7AB2A2F5BFB4C88726AE6D4">
<mods:roleTerm id="191C6897B506ECD761522473673D1E16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="45621D74955914135FF5CE3DFF5F4828">Saboori, Alireza</mods:namePart>
</mods:name>
<mods:name id="CA283FBCBA062E4EB21B476404196BAC" type="personal">
<mods:role id="103B3C6120FC26B9E0B22C63DAE4F63D">
<mods:roleTerm id="6451B1A5953F1701A2FF61CEC81379A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="106805F0A0049B4C557931AFF22B6920">Asadi, Mahdieh</mods:namePart>
</mods:name>
<mods:name id="DD82910CE7FE1F188FF532F8C337E47A" type="personal">
<mods:role id="BE7FE9C0F5BB19CB9E9B6B40F295E2B4">
<mods:roleTerm id="F432681995207A04E4463D1996DC9AA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31EAF2DC35698F05983ADFAB1CF6DC08">Vafaei, Reza</mods:namePart>
</mods:name>
<mods:typeOfResource id="2904BCE6BA4097BFE4644E78E68E274C">text</mods:typeOfResource>
<mods:relatedItem id="6949AD7729C5BF5C247CDA659F5EB7B5" type="host">
<mods:titleInfo id="B328CCB184ADB751E9C1C76A0E085E6A">
<mods:title id="B00D33CADEBB4F4E7238AC974280023B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="846BAEAD7F43639B758F9E36893ED3B1">
<mods:date id="D63B73196A863ABB8518845177A9C965">2005</mods:date>
<mods:detail id="B1FA0F743606F35DBF7E61D8BB83C4C1" type="pubDate">
<mods:number id="2F73F9DC7321F3F64428BB25144AD3B4">2005-08-08</mods:number>
</mods:detail>
<mods:detail id="D6DCF62B4CA2658FA566BCB102BF5FF1" type="volume">
<mods:number id="8E19E0092D09A39B93E13D22481DC0C5">1030</mods:number>
</mods:detail>
<mods:detail id="451C443464B732EE20EA381443B4909E" type="issue">
<mods:number id="AFC39A88D053421E40A0E9379AD45BC5">1</mods:number>
</mods:detail>
<mods:extent id="1AABE09970F8EDDBA80C41B75C5FB9FC" unit="page">
<mods:start id="CFCFCAC2C117842BBE9C0BF5FE60A8E3">49</mods:start>
<mods:end id="8EF5C00E521DC9731DB7B40F296D096E">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="55335B5CE832D2C5044155BD42D376C7">
<mods:url id="0E253A9DDAB560F0373E62850BBB495D">https://biotaxa.org/Zootaxa/article/view/zootaxa.1030.1.2</mods:url>
</mods:location>
<mods:classification id="D07F7B14F952C825040EED4878C97D0E">journal article</mods:classification>
<mods:identifier id="F7285FF296B858507BCB6A9C1AF15D97" type="DOI">10.11646/zootaxa.1030.1.2</mods:identifier>
<mods:identifier id="BF1246BDA363EA638ADD7F9A0F7050CA" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="115A3B4EF3FFAF3C42C54AEE7234D22E" type="Zenodo-Dep">5050339</mods:identifier>
<mods:identifier id="C8873368E8499CE187A56E5ABB1EB805" type="ZooBank">72C9EB4C-068B-42EA-8B90-9DA5024E4CD1</mods:identifier>
</mods:mods>
<treatment id="038A87A9FFA8FFFDE107FDD3FC693F29" ID-DOI="http://doi.org/10.5281/zenodo.5052943" ID-Zenodo-Dep="5052943" LSID="urn:lsid:plazi:treatment:038A87A9FFA8FFFDE107FDD3FC693F29" httpUri="http://treatment.plazi.org/id/038A87A9FFA8FFFDE107FDD3FC693F29" lastPageNumber="52" pageId="3" pageNumber="52">
<subSubSection id="C3396534FFA8FFFDE107FDD3FD7D3BCA" box="[264,763,638,664]" pageId="3" pageNumber="52" type="nomenclature">
<paragraph id="8B9C36BFFFA8FFFDE107FDD3FD7D3BCA" blockId="3.[264,763,638,699]" box="[264,763,638,664]" pageId="3" pageNumber="52">
<heading id="D0D481D3FFA8FFFDE107FDD3FD7D3BCA" bold="true" box="[264,763,638,664]" fontSize="11" level="1" pageId="3" pageNumber="52" reason="1">
<taxonomicName id="4C234D3CFFA8FFFDE107FDD3FD7D3BCA" authority="(Walter, 1947)" baseAuthorityName="Walter" baseAuthorityYear="1947" box="[264,763,638,664]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B957EAADFFA8FFFDE107FDD3FD7D3BCA" bold="true" box="[264,763,638,664]" pageId="3" pageNumber="52">
<emphasis id="B957EAADFFA8FFFDE107FDD3FDC53BCA" bold="true" box="[264,579,638,664]" italics="true" pageId="3" pageNumber="52">Atractides cf. denticulatus</emphasis>
(
<bibRefCitation id="EFB24B4EFFA8FFFDE25DFDD3FD743BCA" author="Walter, C." box="[594,754,638,664]" pageId="3" pageNumber="52" pagination="146 - 238" refId="ref4486" refString="Walter, C. (1947) Neue Acari (Hydrachnellae, Porohalacaridae, Trombidiidae) aus subterranen Gewassern der Schweiz und Rumaniens. Verhandlungen Naturforschende Gesellschaft, Basel, 58, 146 - 238." type="journal article" year="1947">Walter, 1947</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396534FFA8FFFDE107FD0CFC693F29" pageId="3" pageNumber="52" type="description">
<paragraph id="8B9C36BFFFA8FFFDE107FD0CFE0E3BE9" blockId="3.[264,763,638,699]" box="[264,392,673,699]" pageId="3" pageNumber="52">
(
<figureCitation id="13182A3AFFA8FFFDE11EFD0CFEF93BE9" box="[273,383,673,699]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Figs. 16</figureCitation>
)
</paragraph>
<paragraph id="8B9C36BFFFA8FFFDE107FD5CFC863A61" blockId="3.[264,1325,753,1659]" pageId="3" pageNumber="52">
<emphasis id="B957EAADFFA8FFFDE107FD5CFE6C3A58" box="[264,490,753,778]" italics="true" pageId="3" pageNumber="52">Material examined</emphasis>
:
<materialsCitation id="3B4B3CE2FFA8FFFDE1F2FD5CFD273A61" collectingDate="2003-07-27" collectorName="Pesic" elevation="2370" latitude="30.5" location="Simk village" longLatPrecision="1221" longitude="57.2" pageId="3" pageNumber="52" specimenCount="1" stateProvince="Kerman">
IR24
<collectingRegion id="49E7F85DFFA8FFFDE24CFD5CFC913A59" box="[579,791,753,779]" country="Iran" name="Kerman" pageId="3" pageNumber="52">Kerman Province</collectingRegion>
, stream in
<location id="8EFC6064FFA8FFFDE3ADFD5CFBBD3A59" LSID="urn:lsid:plazi:treatment:038A87A9FFA8FFFDE107FDD3FC693F29:8EFC6064FFA8FFFDE3ADFD5CFBBD3A59" box="[930,1083,753,779]" latitude="30.5" longLatPrecision="1221" longitude="57.2" name="Simk village" pageId="3" pageNumber="52" stateProvince="Kerman">Simk village</location>
(
<geoCoordinate id="EE175078FFA8FFFDE440FD5CFB363A59" box="[1103,1200,753,779]" degrees="30" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="52" precision="925" value="30.5">30°30'N</geoCoordinate>
<geoCoordinate id="EE175078FFA8FFFDE4B4FD5CFA9A3A59" box="[1211,1308,753,779]" degrees="57" direction="east" minutes="12" orientation="longitude" pageId="3" pageNumber="52" precision="925" value="57.2">57°12'E</geoCoordinate>
),
<elevation id="000ED18CFFA8FFFDE107FCB4FE083A61" box="[264,398,793,819]" metricMagnitude="3" metricUnit="m" metricValue="2.37" pageId="3" pageNumber="52" unit="m" value="2370.0">
<quantity id="4CDB9B5AFFA8FFFDE107FCB4FEE73A61" box="[264,353,793,819]" metricMagnitude="3" metricUnit="m" metricValue="2.37" pageId="3" pageNumber="52" unit="m" value="2370.0">2370 m</quantity>
asl
</elevation>
.,
<date id="FF9D107FFFA8FFFDE190FCB4FDA33A61" box="[415,549,793,819]" pageId="3" pageNumber="52" value="2003-07-27">
<collectingDate id="EFD9E997FFA8FFFDE190FCB4FDA33A61" box="[415,549,793,819]" pageId="3" pageNumber="52" value="2003-07-27">27.07.2003</collectingDate>
</date>
, leg.
<collectorName id="26D65369FFA8FFFDE26BFCB4FD273A61" box="[612,673,793,819]" pageId="3" pageNumber="52">Pesic</collectorName>
</materialsCitation>
(0/1/0).
</paragraph>
<paragraph id="8B9C36BFFFA8FFFDE137FCECFDA63D19" blockId="3.[264,1325,753,1659]" pageId="3" pageNumber="52">
<emphasis id="B957EAADFFA8FFFDE137FCECFDA83A08" box="[312,558,833,858]" italics="true" pageId="3" pageNumber="52">Description: Female</emphasis>
: Idiosoma L/W 806/594; Dgl­3 not fused with postoc and D­1; coxal field L 285; Cx­3 W 362; Cx­1+2 mL 78; I­L­5 dL 169, H 35, dL/H ratio 4.83; S­1 L 62, L/W ratio 10.6, S­2 L 71, L/W ratio 13.7, distance between sword setae 3, L ratio S­1/ 2 0.87; I­L­6 L 146, H 28, dL/H ratio 5.2; L ratio I­L­5/6 1.16; genital field L/W 177/144, Ac­1­3 27, 24, 24; chelicera total L 291; palp total L 459, dL and relative L (in parentheses given as % of total length): P­1 34 (7), P­2 116 (25), P­3 109 (24), P­4 150 (33), P­5 50 (11); L ratio P­2/4 0.77.
</paragraph>
<paragraph id="8B9C36BFFFA8FFFDE137FBF4FC6D3F01" blockId="3.[264,1325,753,1659]" pageId="3" pageNumber="52">
<emphasis id="B957EAADFFA8FFFDE137FBF4FE443D20" box="[312,450,1113,1138]" italics="true" pageId="3" pageNumber="52">Discussion:</emphasis>
The Iranian specimen shows a general conformity with
<taxonomicName id="4C234D3CFFA8FFFDE456FBF4FEC93DC8" baseAuthorityName="Walter" baseAuthorityYear="1947" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B957EAADFFA8FFFDE456FBF4FEC93DC8" italics="true" pageId="3" pageNumber="52">Atractides denticulatus</emphasis>
</taxonomicName>
in the combination of a striated integument, presence of muscle attachment sclerites, sclerotized excretory pore, fused Vgl­1/2, similar morphology of I­L­5 and 6 (
<figureCitation id="13182A3AFFA8FFFDE4DBFB04FA9C3D91" box="[1236,1306,1193,1219]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Fig. 3</figureCitation>
), IV­L­5 with a long, fine, distal hair (
<figureCitation id="13182A3AFFA8FFFDE2CEFB7CFC8D3DB9" box="[705,779,1233,1259]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Fig. 6</figureCitation>
), and the shape of palp with denticulation on ventral surface of P­2 and P­3. However, clear morphological differences can be observed between specimens from
<collectingCountry id="F334762FFFA8FFFDE237FA8CFDE13C69" box="[568,615,1313,1339]" name="Iran" pageId="3" pageNumber="52">Iran</collectingCountry>
and Central Europe (
<bibRefCitation id="EFB24B4EFFA8FFFDE366FA8CFB933C69" author="Gerecke, R." box="[873,1045,1313,1339]" pageId="3" pageNumber="52" pagination="141 - 378" refId="ref3650" refString="Gerecke, R. (2003) Water mite of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke 2003</bibRefCitation>
). In the specimen from Simk a Dgl­3 is not fused with postoc and D­1 (
<figureCitation id="13182A3AFFA8FFFDE342FAE4FC153C31" box="[845,915,1353,1379]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
), the gonopore is more elongated (
<figureCitation id="13182A3AFFA8FFFDE11EFADCFEDA3CD9" box="[273,348,1393,1419]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Fig. 2</figureCitation>
), and the ventral surface of P­2 and P­3 has a weaker denticulation (
<figureCitation id="13182A3AFFA8FFFDE4ABFADCFA9D3CD9" box="[1188,1307,1393,1419]" captionStart="FIGURES 16" captionStartId="4.[264,382,1492,1516]" captionTargetBox="[308,1302,335,1465]" captionTargetId="figure-85@4.[285,1302,329,1466]" captionTargetPageId="4" captionText="FIGURES 16. Atractides cf. denticulatus (Walter, 1947), female: 1 = idiosoma, dorsal view; 2 = genital field; 3 = I­L­5/6; 4 = palp, medial view; 5 = palp, lateral view; 6 = I­L­5. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050341" httpUri="https://zenodo.org/record/5050341/files/figure.png" pageId="3" pageNumber="52">Figs. 4, 5</figureCitation>
). Specimens described from
<collectingCountry id="F334762FFFA8FFFDE244FA34FD363CE1" box="[587,688,1433,1459]" name="Bulgaria" pageId="3" pageNumber="52">Bulgaria</collectingCountry>
(
<bibRefCitation id="EFB24B4EFFA8FFFDE2CFFA34FCE43CE1" author="Petrova, A." box="[704,866,1433,1459]" pageId="3" pageNumber="52" pagination="47 - 93" refId="ref4147" refString="Petrova, A. (1968) Hydracariens souterrains de Bulgarie III. (Hydrachnellae, Acari). Bulletin de l'Institut de Zoolgie et Musee, Academie Bulgare des Sciences, 28, 47 - 93." type="journal article" year="1968">Petrova 1968</bibRefCitation>
), based on the shape and arrangement of dorsal sclerites (Dgl­3 not fused with postoc and D­1), more closely resemble the specimens from
<collectingCountry id="F334762FFFA8FFFDE19FFA44FE393F51" box="[400,447,1513,1539]" name="Iran" pageId="3" pageNumber="52">Iran</collectingCountry>
than the orginal description. According to
<bibRefCitation id="EFB24B4EFFA8FFFDE3CEFA44FBFE3F51" author="Gerecke, R." box="[961,1144,1513,1539]" pageId="3" pageNumber="52" pagination="141 - 378" refId="ref3650" refString="Gerecke, R. (2003) Water mite of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke (2003)</bibRefCitation>
future investigations on the geographical variation in
<taxonomicName id="4C234D3CFFA8FFFDE302F9BCFC433F78" baseAuthorityName="Walter" baseAuthorityYear="1947" box="[781,965,1553,1578]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B957EAADFFA8FFFDE302F9BCFC433F78" box="[781,965,1553,1578]" italics="true" pageId="3" pageNumber="52">A. denticulatus</emphasis>
</taxonomicName>
could prove that this taxon represents a species complex. This is the first record from
<collectingCountry id="F334762FFFA8FFFDE3BAF994FC6E3F01" box="[949,1000,1593,1619]" name="Iran" pageId="3" pageNumber="52">Iran</collectingCountry>
.
</paragraph>
<paragraph id="8B9C36BFFFA8FFFDE137F9CCFC693F29" blockId="3.[264,1325,753,1659]" box="[312,1007,1633,1659]" pageId="3" pageNumber="52">
<emphasis id="B957EAADFFA8FFFDE137F9CCFE413F28" box="[312,455,1633,1658]" italics="true" pageId="3" pageNumber="52">Distribution</emphasis>
: Central and south­east Europe,
<collectingCountry id="F334762FFFA8FFFDE348F9CCFC2A3F29" box="[839,940,1633,1659]" name="United Kingdom" pageId="3" pageNumber="52">England</collectingCountry>
,
<collectingCountry id="F334762FFFA8FFFDE3B7F9CCFC6D3F29" box="[952,1003,1633,1659]" name="Iran" pageId="3" pageNumber="52">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>