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<document id="ED5BF12EF19E8FF9F30FE4198498DF5F" ID-CLB-Dataset="133817" ID-DOI="10.1093/zoolinnean/zlac015" ID-GBIF-Dataset="372f8265-c53c-43fa-9448-07d5a58d311f" ID-ISSN="0024-4082" ID-Zenodo-Dep="7037958" ID-ZooBank="78C56409-FCCF-4116-8D8C-FF66B247C56C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1661767832469" checkinUser="plazi" docAuthor="Kajihara, Hiroshi, Abukawa, Shushi &amp; Chernyshev, Alexei V." docDate="2022" docId="038A8132EE76ED58FCBACFC9FB9EFB72" docLanguage="en" docName="zlac015.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Baseodiscus delineatus" docType="treatment" docVersion="10" lastPageNumber="533" masterDocId="FFB3F94AEE6BED46FFE9CE7DFFF7FFBC" masterDocTitle="Exploring the basal topology of the heteronemertean tree of life: establishment of a new family, along with turbotaxonomy of Valenciniidae (Nemertea: Pilidiophora: Heteronemertea)" masterLastPageNumber="548" masterPageNumber="503" pageNumber="532" updateTime="1699374125710" updateUser="ExternalLinkService">
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<mods:title id="B9359C59DCD39374B13B8137710A1F50">Exploring the basal topology of the heteronemertean tree of life: establishment of a new family, along with turbotaxonomy of Valenciniidae (Nemertea: Pilidiophora: Heteronemertea)</mods:title>
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<mods:namePart id="B2EB93311FD80A60FCB63718F8F969E2">Abukawa, Shushi</mods:namePart>
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<mods:namePart id="6C4D387766E462F331CFD0DC9B928B15">Chernyshev, Alexei V.</mods:namePart>
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<mods:date id="26B21A16E679C043D1A13A973243D4AA">2022</mods:date>
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<subSubSection id="C33963AFEE76ED5BFCBACFC9FA7DFE70" box="[851,1418,435,460]" pageId="29" pageNumber="532" type="nomenclature">
<paragraph id="8B9C3024EE76ED5BFCBACFC9FA7DFE70" blockId="29.[851,1418,435,460]" box="[851,1418,435,460]" pageId="29" pageNumber="532">
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<taxonomicName id="4C234BA7EE76ED5BFCBACFC9FA7DFE70" ID-CoL="5WCTV" ID-ENA="1519741" authority="(DELLE CHIAJE, 1822)" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1822" box="[851,1418,435,460]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE76ED5BFCBACFC9FB71FE77" box="[851,1158,436,459]" italics="true" pageId="29" pageNumber="532">BASEODISCUS DELINEATUS</emphasis>
(DELLE CHIAJE, 1822)
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<subSubSection id="C33963AFEE76ED5BFBC2CFA7FB45FE4F" box="[1067,1202,474,499]" pageId="29" pageNumber="532" type="description">
<paragraph id="8B9C3024EE76ED5BFBC2CFA7FB45FE4F" blockId="29.[1067,1202,474,499]" box="[1067,1202,474,499]" pageId="29" pageNumber="532">
(
<figureCitation id="13182CA1EE76ED5BFBDACFA7FB5DFE4F" box="[1075,1194,474,499]" captionStart="Figure 3" captionStartId="12.[142,222,1745,1767]" captionTargetBox="[222,1342,194,1706]" captionTargetId="figure-25@12.[222,1342,194,1706]" captionTargetPageId="12" captionText="Figure 3. A, Baseodiscus aff. marmoratus (Bürger, 1890) (voucher DNA, ICHUM 6322); B, Baseodiscus aff. maculosus (Bürger, 1895a) (voucher DNA, ICHUM 6324); CE, Baseodiscus delineatus (Delle Chiaje, 1822-1829), ICHUM 6326; F, Baseodiscus cf. curtus (Hubrecht, 1879), ICHUM 6328; G, H, Baseodiscus urgorrii sp. nov., holotype, MCZ IZ-135319, entire body (G) and protruding proboscis (indicated by an arrowhead, H); I, Baseodiscus punnetti (Coe, 1904), specimen different from any voucher specimens in this study; J, K, Baseodiscus kakuii sp. nov., holotype, ICHUM 6334, anterior body (J) and" figureDoi="http://doi.org/10.5281/zenodo.7037968" httpUri="https://zenodo.org/record/7037968/files/figure.png" pageId="29" pageNumber="532">FIG. 3CE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33963AFEE76ED5BFCD2CC7FFC1EF997" pageId="29" pageNumber="532" type="reference_group">
<paragraph id="8B9C3024EE76ED5BFCD2CC7FFB02FD6C" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<taxonomicName id="4C234BA7EE76ED5BFCD2CC7FFB2CFDA4" authority="Delle Chiaje, 1822" authorityName="Delle Chiaje" authorityYear="1825" box="[827,1243,514,536]" class="Pilidiophora" family="Lineidae" genus="Polia" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE76ED5BFCD2CC7FFC1BFDAB" box="[827,1004,514,535]" italics="true" pageId="29" pageNumber="532">Polia delineata</emphasis>
Delle Chiaje,
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in Delle Chiaje (182229), pl. XXVIII (28, but correctly
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position), fig. 4;
<bibRefCitation id="EFB24DD5EE76ED5BFBFFCC3DFB1DFDEA" author="Delle Chiaje S" box="[1046,1258,576,598]" pageId="29" pageNumber="532" pagination="185 - 444" refId="ref35980" refString="Delle Chiaje S. 1822 - 1829. Memorie sulla storia e notomia degli animali senza vertebre del regno di Napoli. Naples: Societa' Tipografica [pls I - CIX published in 1822; Vol. 1, xii + 184 pp., in 1825; Vol. 2, 260 pp. (pp. 185 - 444), in 1825; Vol. 3, xx + 232 pp., in 1828; Vol. 4, viii + 214 pp., in 1829]." type="book chapter" year="1825">
Delle Chiaje,
<quantity id="4CDB9DC1EE76ED5BFB5ACC3DFB1DFDEA" box="[1203,1258,576,598]" metricMagnitude="1" metricUnit="m" metricValue="4.6355" pageId="29" pageNumber="532" unit="in" value="1825.0">1825</quantity>
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in Delle Chiaje (182229): 427428, 444 (Naples,
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);
<bibRefCitation id="EFB24DD5EE76ED5BFABFCC23FC10FD2F" author="Grube AE" pageId="29" pageNumber="532" refId="ref37043" refString="Grube AE. 1840. Actinien, Echinodermen und Wurmer des Adriatischen- und Mittelmeers. Konigsberg [Kaliningrad]: J. H. Bon." type="book" year="1840">Grube, 1840: 5758</bibRefCitation>
, fig. 8a, b (Naples and Palermo,
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);
<bibRefCitation id="EFB24DD5EE76ED5BFCB5CCE1FBBEFD0D" author="Hubrecht AAW" box="[860,1097,668,690]" pageId="29" pageNumber="532" pagination="193 - 232" refId="ref37309" refString="Hubrecht AAW. 1879. The genera of European nemerteans critically revised, with description of several new species. Notes from the Leyden Museum 1: 193 - 232." type="journal article" year="1879">Hubrecht, 1879: 209</bibRefCitation>
(Naples,
<collectingCountry id="F33470B4EE76ED5BFB52CCE1FB04FD0E" box="[1211,1267,668,690]" name="Italy" pageId="29" pageNumber="532">Italy</collectingCountry>
);
<bibRefCitation id="EFB24DD5EE76ED5BFAE1CCE1FC4CFD6C" author="Joubin L" pageId="29" pageNumber="532" pagination="461 - 602" refId="ref37886" refString="Joubin L. 1890. Recherches sur les turbellaries des cotes de France (Nemertes). Archives de Zoologie Experimentale et Generale, Deuxieme Serie 8: 461 - 602, pls XXV - XXXI." type="journal article" year="1890">Joubin, 1890: 510511</bibRefCitation>
(Banyuls-sur-Mer,
<collectingCountry id="F33470B4EE76ED5BFB7FCCC6FB1EFD6C" box="[1174,1257,699,720]" name="France" pageId="29" pageNumber="532">France</collectingCountry>
).
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CCA7FA53FD53" blockId="29.[827,1444,514,1579]" box="[827,1444,729,751]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CCA7FA53FD53" box="[827,1444,729,751]" pageId="29" pageNumber="532">
<treatmentCitation id="0A821635EE76ED5BFCD2CCA7FB04FD53" author="Kolliker A" box="[827,1267,729,751]" page="95" pageId="29" pageNumber="532" year="1845">
<taxonomicName id="4C234BA7EE76ED5BFCD2CCA7FB04FD53" authority=": Kolliker, 1845: 95" authorityName="Kolliker" authorityPageNumber="95" authorityYear="1845" box="[827,1267,729,751]" class="Pilidiophora" family="Lineidae" genus="Nemertes" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE76ED5BFCD2CCA7FBD7FD52" box="[827,1056,729,751]" italics="true" pageId="29" pageNumber="532">Nemertes delineatus</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFBC4CCA4FB04FD53" author="Kolliker A" box="[1069,1267,729,751]" pageId="29" pageNumber="532" pagination="86 - 98" refId="ref38934" refString="Kolliker A. 1845. Drei neue Gattungen von Wurmern. Verhandlungen der Schweizerischen Naturforschenden Gesellschaft bei ihrer Versammlung zu Chur 29: 86 - 98." type="journal article" year="1845">Kölliker, 1845: 95</bibRefCitation>
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(Naples,
<collectingCountry id="F33470B4EE76ED5BFAB7CCA4FA60FD53" box="[1374,1431,729,751]" name="Italy" pageId="29" pageNumber="532">Italy</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CC85FAFCFC1B" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CC85FAFCFC1B" pageId="29" pageNumber="532">
<treatmentCitation id="0A821635EE76ED5BFCD2CC85FB1CFCB2" author="Burger O" box="[827,1259,760,782]" page="151" pageId="29" pageNumber="532" year="1892">
<taxonomicName id="4C234BA7EE76ED5BFCD2CC85FB1CFCB2" authority=": Burger, 1892: 151" authorityName="Burger" authorityPageNumber="151" authorityYear="1892" box="[827,1259,760,782]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE76ED5BFCD2CC85FBFEFCB1" box="[827,1033,760,781]" italics="true" pageId="29" pageNumber="532">Eupolia delineata</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFBF3CC85FB1CFCB2" author="Burger O" box="[1050,1259,760,782]" pageId="29" pageNumber="532" pagination="137 - 178" refId="ref34328" refString="Burger O. 1892. Zur Systematik der Nemertinenfauna des Golfs von Neapel. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften und der Georg-Augusts- Universitat zu Gottingen 5: 137 - 178." type="journal article" year="1892">Bürger, 1892: 151</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(Naples,
<collectingCountry id="F33470B4EE76ED5BFAB7CC85FA61FCB2" box="[1374,1430,760,782]" name="Italy" pageId="29" pageNumber="532">Italy</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFCB2CD6BFBB9FC90" author="Joubin L" box="[859,1102,790,812]" page="79" pageId="29" pageNumber="532" year="1894">
<bibRefCitation id="EFB24DD5EE76ED5BFCB2CD6BFBB9FC90" author="Joubin L" box="[859,1102,790,812]" pageId="29" pageNumber="532" refId="ref37923" refString="Joubin L. 1894. Faune Francaise, publiee par les soins de MM. Raphael Blanchard et Jules de Guerne; les Nemertiens. Paris: Societe d'Editions Scientifiques." type="book" year="1894">Joubin, 1894: 7980</bibRefCitation>
</treatmentCitation>
, pl. I, fig. 11 (Banyuls-surMer,
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);
<treatmentCitation id="0A821635EE76ED5BFC15CD4BFAF9FCF7" author="Burger O" box="[1020,1294,821,843]" page="600" pageId="29" pageNumber="532" year="1895">
<bibRefCitation id="EFB24DD5EE76ED5BFC15CD4BFAF9FCF7" author="Burger O" box="[1020,1294,821,843]" pageId="29" pageNumber="532" refId="ref34395" refString="Burger O. 1895 a. Die Nemertinen des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel und der Angrenzenden Meeres-Abschnitte," type="book" year="1895">Bürger, 1895a: 600601</bibRefCitation>
</treatmentCitation>
, pl. 4, figs 4, 6 (Naples,
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: the specimen in fig. 8 lacks longitudinal stripes in the anterior part of body and is bulkier than that depicted in fig. 6).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CDCDFB17FB06" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CDCDFB17FB06" pageId="29" pageNumber="532">
<treatmentCitation id="0A821635EE76ED5BFCD2CDCDFA68FC79" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" box="[827,1439,944,966]" page="3787" pageId="29" pageNumber="532" year="2005">
<taxonomicName id="4C234BA7EE76ED5BFCD2CDCDFA68FC79" ID-CoL="5WCTV" authority=": Strand et al., 2005: 3787" authorityName="Strand" authorityPageNumber="3787" authorityYear="2005" box="[827,1439,944,966]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE76ED5BFCD2CDCDFBBDFC79" box="[827,1098,944,965]" italics="true" pageId="29" pageNumber="532">Baseodiscus delineatus</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFB88CDCDFA68FC79" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" box="[1121,1439,944,966]" pageId="29" pageNumber="532" pagination="3785 - 3793" refId="ref41221" refString="Strand M, Hjelmgren A, Sundberg P. 2005. Genus Baseodiscus (Nemertea: Heteronemertea): molecular identification of a new species in a phylogenetic context. Journal of Natural History 39: 3785 - 3793." type="journal article" year="2005">
Strand
<emphasis id="B957EC36EE76ED5BFB29CDCCFAF5FC79" box="[1216,1282,944,966]" italics="true" pageId="29" pageNumber="532">et al.</emphasis>
, 2005: 3787
</bibRefCitation>
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</treatmentCitation>
, table II [Ischia,
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; Rottnest Island,
<collectingCountry id="F33470B4EE76ED5BFAC0CDB3FA60FC58" box="[1321,1431,974,996]" name="Australia" pageId="29" pageNumber="532">Australia</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFCB2CD90FBC4FBBF" author="Schwartz ML" box="[859,1075,1005,1027]" page="62" pageId="29" pageNumber="532" year="2009">
<bibRefCitation id="EFB24DD5EE76ED5BFCB2CD90FBC4FBBF" author="Schwartz ML" box="[859,1075,1005,1027]" pageId="29" pageNumber="532" refId="ref40578" refString="Schwartz ML. 2009. Untying a Gordian knot of worms: systematics and taxonomy of the Pilidiophora (phylum Nemertea) from multiple data sets. Unpublished D. Phil. Thesis, The George Washington University." type="book" year="2009">Schwartz, 2009: 62</bibRefCitation>
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, fig. 1B (Carrie Bow Cay,
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; Peanut Island,
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,
<collectingCountry id="F33470B4EE76ED5BFB79CA71FB30FB9D" box="[1168,1223,1036,1057]" name="United States of America" pageId="29" pageNumber="532">USA</collectingCountry>
;
<collectingRegion id="49E7FEC6EE76ED5BFB36CA71FA68FB9D" box="[1247,1439,1036,1058]" country="Panama" name="Bocas del Toro" pageId="29" pageNumber="532">Bocas del Toro</collectingRegion>
,
<collectingCountry id="F33470B4EE76ED5BFCB2CA56FC4CFBFC" box="[859,955,1067,1088]" name="Panama" pageId="29" pageNumber="532">Panama</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFC27CA57FAD9FBFC" author="Gonzalez-Cueto J &amp; Quiroga S &amp; Norenburg J" box="[974,1326,1066,1088]" httpUri="http://treatment.plazi.org/id/217F6DCE8A3CF6CFB0EF3BFFA852D823" page="92" pageId="29" pageNumber="532" year="2014">
<bibRefCitation id="EFB24DD5EE76ED5BFC27CA57FAD9FBFC" author="Gonzalez-Cueto J &amp; Quiroga S &amp; Norenburg J" box="[974,1326,1066,1088]" pageId="29" pageNumber="532" pagination="83 - 108" refId="ref36953" refString="Gonzalez-Cueto J, Quiroga S, Norenburg J. 2014. A shore-based preliminary survey of marine ribbon worms (Nemertea) from the Caribbean coast of Colombia. ZooKeys 439: 83 - 108." type="journal article" year="2014">
Gonzalez-Cueto
<emphasis id="B957EC36EE76ED5BFB61CA56FB36FB83" box="[1160,1217,1066,1088]" italics="true" pageId="29" pageNumber="532">et al.</emphasis>
, 2014: 92
</bibRefCitation>
</treatmentCitation>
, fig. 2CE (Inca-Inca and Taganga,
<collectingCountry id="F33470B4EE76ED5BFB94CA34FB1CFBE3" box="[1149,1259,1097,1119]" name="Colombia" pageId="29" pageNumber="532">Colombia</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFAE9CA34FC3EFBC1" author="Mendes CB &amp; Matthews-Cascon H &amp; Norenburg JL" page="149" pageId="29" pageNumber="532" year="2016">
<bibRefCitation id="EFB24DD5EE76ED5BFAE9CA34FC3EFBC1" author="Mendes CB &amp; Matthews-Cascon H &amp; Norenburg JL" pageId="29" pageNumber="532" pagination="146 - 156" refId="ref39466" refString="Mendes CB, Matthews-Cascon H, Norenburg JL. 2016. New records of ribbon worms (Nemertea) from Ceara, northeast Brazil. Zootaxa 4061: 146 - 156." type="journal article" year="2016">
Mendes
<emphasis id="B957EC36EE76ED5BFA8BCA37FA6AFBE2" box="[1378,1437,1097,1119]" italics="true" pageId="29" pageNumber="532">et al.</emphasis>
, 2016: 149
</bibRefCitation>
</treatmentCitation>
, fig. 2-3 (Caucaia,
<collectingCountry id="F33470B4EE76ED5BFB70CA15FB16FBC2" box="[1177,1249,1128,1150]" name="Brazil" pageId="29" pageNumber="532">Brazil</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFB1DCA15FC71FB20" author="Kajihara H" page="423" pageId="29" pageNumber="532" year="2017">
<bibRefCitation id="EFB24DD5EE76ED5BFB1DCA15FC71FB20" author="Kajihara H" pageId="29" pageNumber="532" pagination="419 - 444" refId="ref38160" refString="Kajihara H. 2017. Species diversity of Japanese ribbon worms (Nemertea). In: Motokawa M, Kajihara H, eds. Species diversity of animals in Japan. Tokyo: Springer Japan, 419 - 444." type="book chapter" year="2017">Kajihara, 2017: 423</bibRefCitation>
</treatmentCitation>
, fig. 16.2b (Misaki,
<collectingCountry id="F33470B4EE76ED5BFB8ACAFAFB5DFB20" box="[1123,1194,1159,1180]" name="Japan" pageId="29" pageNumber="532">Japan</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFB57CAFBFC4BFB07" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" page="348" pageId="29" pageNumber="532" year="2019">
<bibRefCitation id="EFB24DD5EE76ED5BFB57CAFBFC4BFB07" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" pageId="29" pageNumber="532" pagination="348 - 353" refId="ref37407" refString="Ikenaga J, Hookabe N, Kohtsuka H, Yoshida M, Kajihara H. 2019. A population without females: males of Baseodiscus delineatus (Nemertea: Heteronemertea) reproduce asexually by fragmentation. Zoological Science 36: 348 - 353." type="journal article" year="2019">
Ikenaga
<emphasis id="B957EC36EE76ED5BFAC8CAFAFAAEFB27" box="[1313,1369,1158,1180]" italics="true" pageId="29" pageNumber="532">et al.</emphasis>
, 2019: 348353
</bibRefCitation>
</treatmentCitation>
, figs 14 [Misaki,
<collectingCountry id="F33470B4EE76ED5BFB62CAD8FB23FB06" box="[1163,1236,1189,1210]" name="Japan" pageId="29" pageNumber="532">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CAB9FBC4FB44" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CAB9FBC4FB44" pageId="29" pageNumber="532">
<treatmentCitation id="0A821635EE76ED5BFCD2CAB9FA68FB65" author="Kvist S &amp; Laumer CE &amp; Junoy J &amp; Giribet G" box="[827,1439,1220,1242]" page="291" pageId="29" pageNumber="532" year="2014">
<taxonomicName id="4C234BA7EE76ED5BFCD2CAB9FA68FB65" ID-CoL="5WCTV" authority=": Kvist et al., 2014: 291" authorityName="Kvist" authorityPageNumber="291" authorityYear="2014" box="[827,1439,1220,1242]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" isUncertain="true" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE76ED5BFCD2CAB9FC33FB65" box="[827,964,1220,1241]" italics="true" pageId="29" pageNumber="532">Baseodiscus</emphasis>
cf.
<emphasis id="B957EC36EE76ED5BFC12CAB9FB87FB65" box="[1019,1136,1220,1241]" italics="true" pageId="29" pageNumber="532">delineatus</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFB6CCAB9FA68FB65" author="Kvist S &amp; Laumer CE &amp; Junoy J &amp; Giribet G" box="[1157,1439,1220,1242]" pageId="29" pageNumber="532" pagination="287 - 308" refId="ref39075" refString="Kvist S, Laumer CE, Junoy J, Giribet G. 2014. New insights into the phylogeny, systematics and DNA barcoding of Nemertea. Invertebrate Systematics 28: 287 - 308." type="journal article" year="2014">
Kvist
<emphasis id="B957EC36EE76ED5BFB3BCAB8FAE5FB65" box="[1234,1298,1220,1242]" italics="true" pageId="29" pageNumber="532">et al.</emphasis>
, 2014: 291
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, table 1 (
<collectingCountry id="F33470B4EE76ED5BFC5ECA9FFBD0FB44" box="[951,1063,1250,1272]" name="Australia" pageId="29" pageNumber="532">Australia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CB7CFA8CFACF" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CB7CFA8CFACF" pageId="29" pageNumber="532">
?
<treatmentCitation id="0A821635EE76ED5BFCAFCB7CFAB6FAAB" author="Coe WR" box="[838,1345,1281,1303]" page="105" pageId="29" pageNumber="532" year="1947">
<taxonomicName id="4C234BA7EE76ED5BFCAFCB7CFAB6FAAB" ID-CoL="5WCTV" authority=": Coe, 1947: 104 - 105" authorityName=": Coe" authorityPageNumber="104 - 105" authorityYear="1947" box="[838,1345,1281,1303]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE76ED5BFCAFCB7CFBB8FAAA" box="[838,1103,1281,1302]" italics="true" pageId="29" pageNumber="532">Baseodiscus delineatus</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFBB6CB7CFAB6FAAB" author="Coe WR" box="[1119,1345,1281,1303]" pageId="29" pageNumber="532" pagination="101 - 106" refId="ref35631" refString="Coe WR. 1947. Nemerteans of the Hawaiian and Marshall Islands. Occasional Papers of Bernice P. Bishop Museum 14: 101 - 106." type="journal article" year="1947">Coe, 1947: 104105</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(Bogoni Island,
<collectingCountry id="F33470B4EE76ED5BFC5CCB5DFB88FA8A" box="[949,1151,1312,1334]" name="Marshall Islands" pageId="29" pageNumber="532">Marshall Islands</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFB71CB5DFA55FA89" author="Correa DD" box="[1176,1442,1312,1334]" page="442" pageId="29" pageNumber="532" year="1958">
<bibRefCitation id="EFB24DD5EE76ED5BFB71CB5DFA55FA89" author="Correa DD" box="[1176,1442,1312,1334]" pageId="29" pageNumber="532" pagination="441 - 455" refId="ref35723" refString="Correa DD. 1958. Nemertinos do litoral Brasileiro (VII). Anais da Academia Brasileira de Siencias 29: 441 - 455, pls 1 - 4." type="journal article" year="1958">Corrêa, 1958: 442443</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F33470B4EE76ED5BFC8ACB43FC5CFAE8" box="[867,939,1342,1364]" name="Brazil" pageId="29" pageNumber="532">Brazil</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFC57CB43FB48FAE8" author="Gibson R" box="[958,1215,1342,1364]" page="139" pageId="29" pageNumber="532" year="1979">
<bibRefCitation id="EFB24DD5EE76ED5BFC57CB43FB48FAE8" author="Gibson R" box="[958,1215,1342,1364]" pageId="29" pageNumber="532" pagination="137 - 160" refId="ref36552" refString="Gibson R. 1979. Nemerteans of the Great Barrier Reef 2. Anopla Heteronemertea (Baseodiscidae). Zoological Journal of the Linnean Society 66: 137 - 160." type="journal article" year="1979">Gibson, 1979: 139146</bibRefCitation>
</treatmentCitation>
, figs 13 (Low Isles, off Port Douglas, Great Barrier Reef,
<collectingCountry id="F33470B4EE76ED5BFAE9CB20FA87FACF" box="[1280,1392,1373,1395]" name="Australia" pageId="29" pageNumber="532">Australia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CB01FC26FA0C" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CB01FC26FA0C" pageId="29" pageNumber="532">
?
<taxonomicName id="4C234BA7EE76ED5BFCAFCB01FC3AFA0C" ID-CoL="68D6P" authority=": Stiasny-Wijnhoff, 1925: 102" authorityName="Stiasny-Wijnhoff" authorityPageNumber="102" authorityYear="1925" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="curtus">
<treatmentCitation id="0A821635EE76ED5BFCAFCB01FA54FA2D" author="Stiasny-Wijnhoff G" box="[838,1443,1404,1426]" page="102" pageId="29" pageNumber="532" year="1925">
<emphasis id="B957EC36EE76ED5BFCAFCB01FBD9FA2E" box="[838,1070,1404,1426]" italics="true" pageId="29" pageNumber="532">Baseodiscus curtus</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFBA3CB01FA54FA2D" author="Stiasny-Wijnhoff G" box="[1098,1443,1404,1426]" pageId="29" pageNumber="532" pagination="97 - 120" refId="ref41047" refString="Stiasny-Wijnhoff G. 1925. On a collection of nemerteans from Curacao. Bijdragen tot de Dierkunde 24: 97 - 120, pls V - VIII." type="journal article" year="1925">Stiasny-Wijnhoff, 1925: 102</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F33470B4EE76ED5BFC8BCBE6FC32FA0C" box="[866,965,1435,1456]" name="Curaçao" pageId="29" pageNumber="532">Curaçao</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CBC4FA8AFA52" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CBC4FA8AFA52" pageId="29" pageNumber="532">
?
<treatmentCitation id="0A821635EE76ED5BFCAFCBC4FB17FA72" author="Burger O" box="[838,1248,1465,1487]" page="230" pageId="29" pageNumber="532" year="1893">
<taxonomicName id="4C234BA7EE76ED5BFCAFCBC4FB17FA72" authority=": Burger, 1893: 230" authorityName=": Burger" authorityPageNumber="230" authorityYear="1893" box="[838,1248,1465,1487]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE76ED5BFCAFCBC4FBF9FA72" box="[838,1038,1465,1486]" italics="true" pageId="29" pageNumber="532">Eupolia delineata</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFBF3CBC7FB17FA72" author="Burger O" box="[1050,1248,1465,1487]" pageId="29" pageNumber="532" pagination="207 - 240" refId="ref34361" refString="Burger O. 1893. Sudgeorgische und andere exotische Nemertinen. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Biologie der Thiere 7: 207 - 240, pls 8, 9." type="book chapter" year="1893">Bürger, 1893: 230</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 8, fig. 4 (
<collectingRegion id="49E7FEC6EE76ED5BFA8ECBC4FA69FA72" box="[1383,1438,1465,1486]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="532">Java</collectingRegion>
,
<collectingCountry id="F33470B4EE76ED5BFCB0CBA5FC3CFA52" box="[857,971,1496,1518]" name="Indonesia" pageId="29" pageNumber="532">Indonesia</collectingCountry>
);
<treatmentCitation id="0A821635EE76ED5BFC37CBA5FB37FA52" author="Punnett RC" box="[990,1216,1496,1518]" page="825" pageId="29" pageNumber="532" year="1900">
<bibRefCitation id="EFB24DD5EE76ED5BFC37CBA5FB37FA52" author="Punnett RC" box="[990,1216,1496,1518]" pageId="29" pageNumber="532" pagination="825 - 831" refId="ref40142" refString="Punnett RC. 1900 c. On some nemerteans from Torres Straits. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London 68: 825 - 831." type="journal article" year="1900">Punnett, 1900c: 825</bibRefCitation>
</treatmentCitation>
(Torres Straits).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9C3024EE76ED5BFCD2CB8AFC1EF997" blockId="29.[827,1444,514,1579]" pageId="29" pageNumber="532">
<treatmentCitationGroup id="AB33170AEE76ED5BFCD2CB8AFC1EF997" pageId="29" pageNumber="532">
?
<treatmentCitation id="0A821635EE76ED5BFCAFCB8AFC13F997" author="Coe WR" page="226" pageId="29" pageNumber="532" year="1901">
<taxonomicName id="4C234BA7EE76ED5BFCAFCB8AFC13F997" authority=": Coe, 1901 b: 226 (Ensenada)" authorityName=": Coe" authorityYear="1901" class="Pilidiophora" family="Valenciniidae" genus="Taeniosoma" kingdom="Animalia" order="Heteronemertea" pageId="29" pageNumber="532" phylum="Nemertea" rank="species" species="delineatum">
<emphasis id="B957EC36EE76ED5BFCAFCB8AFB89F9B7" box="[838,1150,1526,1548]" italics="true" pageId="29" pageNumber="532">Taeniosoma delineatum</emphasis>
:
<bibRefCitation id="EFB24DD5EE76ED5BFB45CB8AFA54F9B0" author="Coe WR" box="[1196,1443,1526,1548]" pageId="29" pageNumber="532" pagination="223 - 229" refId="ref35410" refString="Coe WR. 1901 b. The nemerteans of Porto Rico. Bulletin of the United States Commission of Fish and Fisheries for 1900 2: 223 - 229." type="journal article" year="1901">Coe, 1901b: 226</bibRefCitation>
(Ensenada)
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33963AFEE76ED58FCD2C814FD95FAD0" lastPageId="30" lastPageNumber="533" pageId="29" pageNumber="532" type="materials_examined">
<paragraph id="8B9C3024EE76ED5BFCD2C814FBA9F8C8" blockId="29.[827,1444,1641,1908]" pageId="29" pageNumber="532">
<emphasis id="B957EC36EE76ED5BFCD2C814FBD6F9C2" box="[827,1057,1641,1662]" italics="true" pageId="29" pageNumber="532">Material examined:</emphasis>
<materialsCitation id="3B4B3A79EE76ED5BFBC6C814FC0EF8A4" ID-GBIF-Occurrence="3897229313" collectingDate="2014-02-22" collectionCode="ICHUM" collectorName="H. Kajihara" latitude="35.158337" location="Misaki Marine Biological Station" longLatPrecision="19" longitude="139.6125" pageId="29" pageNumber="532" specimenCode="ICHUM 6325" specimenCount="1">
Five specimens (extracted total DNA and remaining body preserved in 99% EtOH):
<specimenCount id="9D25FBADEE76ED5BFCD2C8DBFC12F900" box="[827,997,1702,1724]" pageId="29" pageNumber="532" type="generic">four specimens</specimenCount>
(including
<specimenCode id="DB85985FEE76ED5BFB8EC8DAFAF4F900" box="[1127,1283,1702,1724]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="29" pageNumber="532">ICHUM 6325</specimenCode>
),
<date id="FF9D16E4EE76ED5BFAFEC8DAFC83F966" pageId="29" pageNumber="532" value="2014-02-22">
<collectingDate id="EFD9EF0CEE76ED5BFAFEC8DAFC83F966" pageId="29" pageNumber="532" value="2014-02-22">22 February 2014</collectingDate>
</date>
, rocky intertidal, under stone, near
<location id="8EFC66FFEE76ED5BFB15C8B8FBF2F946" LSID="urn:lsid:plazi:treatment:038A8132EE76ED58FCBACFC9FB9EFB72:8EFC66FFEE76ED5BFB15C8B8FBF2F946" latitude="35.158337" longLatPrecision="19" longitude="139.6125" name="Misaki Marine Biological Station" pageId="29" pageNumber="532">Misaki Marine Biological Station</location>
(
<geoCoordinate id="EE1756E3EE76ED5BFBFAC899FB62F945" box="[1043,1173,1763,1786]" degrees="35" direction="north" minutes="09" orientation="latitude" pageId="29" pageNumber="532" precision="15" seconds="30" value="35.158337">35°0930″N</geoCoordinate>
,
<geoCoordinate id="EE1756E3EE76ED5BFB49C899FADAF945" box="[1184,1325,1763,1786]" degrees="139" direction="east" minutes="36" orientation="longitude" pageId="29" pageNumber="532" precision="15" seconds="45" value="139.6125">139°3645″E</geoCoordinate>
), collected by
<collectorName id="26D655F2EE76ED5BFC8BC97EFC0EF8A4" box="[866,1017,1794,1816]" pageId="29" pageNumber="532">H. Kajihara</collectorName>
</materialsCitation>
;
<materialsCitation id="3B4B3A79EE76ED5BFBE0C97EFBA9F8C8" ID-GBIF-Occurrence="3897229477" collectingDate="2014-03-26" collectionCode="ICHUM" collectorName="Misaki &amp; H. Kohtsuka." country="Japan" latitude="35.15917" location="Araihama" longLatPrecision="19" longitude="139.61111" pageId="29" pageNumber="532" specimenCode="ICHUM 6326" specimenCount="1" stateProvince="Kanagawa">
<specimenCount id="9D25FBADEE76ED5BFBE0C97EFB47F8A4" box="[1033,1200,1794,1816]" pageId="29" pageNumber="532" type="generic">one specimen</specimenCount>
(
<specimenCode id="DB85985FEE76ED5BFB2AC97EFA99F8A4" box="[1219,1390,1794,1816]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="29" pageNumber="532">ICHUM 6326</specimenCode>
),
<date id="FF9D16E4EE76ED5BFA6FC97FFC30F88A" pageId="29" pageNumber="532" value="2014-03-26">
<collectingDate id="EFD9EF0CEE76ED5BFA6FC97FFC30F88A" pageId="29" pageNumber="532" value="2014-03-26">26 March 2014</collectingDate>
</date>
, rocky intertidal, under stone,
<location id="8EFC66FFEE76ED5BFAC7C95CFA55F88B" LSID="urn:lsid:plazi:treatment:038A8132EE76ED58FCBACFC9FB9EFB72:8EFC66FFEE76ED5BFAC7C95CFA55F88B" box="[1326,1442,1825,1847]" country="Japan" latitude="35.15917" longLatPrecision="19" longitude="139.61111" name="Araihama" pageId="29" pageNumber="532" stateProvince="Kanagawa">Araihama</location>
(
<geoCoordinate id="EE1756E3EE76ED5BFCAAC93DFC32F8E9" box="[835,965,1855,1878]" degrees="35" direction="north" minutes="09" orientation="latitude" pageId="29" pageNumber="532" precision="15" seconds="33" value="35.15917">35°0933″N</geoCoordinate>
,
<geoCoordinate id="EE1756E3EE76ED5BFC39C93DFBA9F8E9" box="[976,1118,1855,1878]" degrees="139" direction="east" minutes="36" orientation="longitude" pageId="29" pageNumber="532" precision="15" seconds="40" value="139.61111">139°3640″E</geoCoordinate>
),
<collectorName id="26D655F2EE76ED5BFB9AC93DFB33F8EA" box="[1139,1220,1856,1878]" pageId="29" pageNumber="532">Misaki</collectorName>
,
<collectingRegion id="49E7FEC6EE76ED5BFB39C93DFABCF8E9" box="[1232,1355,1856,1877]" country="Japan" name="Kanagawa" pageId="29" pageNumber="532">Kanagawa</collectingRegion>
,
<collectingCountry id="F33470B4EE76ED5BFABEC93DFA68F8E9" box="[1367,1439,1856,1877]" name="Japan" pageId="29" pageNumber="532">Japan</collectingCountry>
, collected by
<collectorName id="26D655F2EE76ED5BFC2FC922FBA9F8C8" box="[966,1118,1886,1908]" pageId="29" pageNumber="532">H. Kohtsuka.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9C3024EE75ED58FF78CEBBFD47FEA4" blockId="30.[144,761,197,280]" pageId="30" pageNumber="533">
<emphasis id="B957EC36EE75ED58FF78CEBBFEEEFF67" box="[145,281,198,219]" italics="true" pageId="30" pageNumber="533">Sequences:</emphasis>
<materialsCitation id="3B4B3A79EE75ED58FEC1CEBBFD5BFEA4" ID-GBIF-Occurrence="3897229314" accessionNumber="LC178621, LC178665, LC178647, LC178666" collectionCode="ICHUM" pageId="30" pageNumber="533" specimenCode="ICHUM 6325, ICHUM 6326" specimenCount="1">
From
<specimenCode id="DB85985FEE75ED58FE9ECEB8FDD1FF67" box="[375,550,197,219]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="30" pageNumber="533">ICHUM 6325</specimenCode>
:
<accessionNumber id="9470ADC7EE75ED58FDDECEB8FD4EFF67" box="[567,697,197,219]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178621" pageId="30" pageNumber="533">LC178621</accessionNumber>
, 28S (1136 bp);
<accessionNumber id="9470ADC7EE75ED58FEEDCE99FE8FFF46" box="[260,376,228,250]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178665" pageId="30" pageNumber="533">LC178665</accessionNumber>
, 16S (521 bp). From
<specimenCode id="DB85985FEE75ED58FDB2CE99FD02FF46" box="[603,757,228,250]" collectionCode="ICHUM" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="30" pageNumber="533">ICHUM 6326</specimenCode>
:
<accessionNumber id="9470ADC7EE75ED58FF79CF7FFEF2FEA4" box="[144,261,258,280]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178647" pageId="30" pageNumber="533">LC178647</accessionNumber>
,
<emphasis id="B957EC36EE75ED58FEF9CF7EFEC6FEA4" box="[272,305,259,280]" italics="true" pageId="30" pageNumber="533">H3</emphasis>
(331 bp);
<accessionNumber id="9470ADC7EE75ED58FE77CF7FFDE4FEA4" box="[414,531,258,280]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178666" pageId="30" pageNumber="533">LC178666</accessionNumber>
, 16S (521 bp)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9C3024EE75ED58FF78CF3FFD6FFE0E" blockId="30.[145,761,321,435]" pageId="30" pageNumber="533">
<emphasis id="B957EC36EE75ED58FF78CF3FFEDEFEEB" box="[145,297,322,343]" italics="true" pageId="30" pageNumber="533">Description:</emphasis>
Body up to
<quantity id="4CDB9DC1EE75ED58FE26CF3CFDEEFEEB" box="[463,537,321,343]" metricMagnitude="-1" metricUnit="m" metricValue="5.6" pageId="30" pageNumber="533" unit="cm" value="56.0">56 cm</quantity>
long,
<quantity id="4CDB9DC1EE75ED58FD85CF3CFD44FEEB" box="[620,691,321,343]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="533" unit="mm" value="3.0">3 mm</quantity>
wide (
<collectionCode id="ED32A8E1EE75ED58FF71CF1DFF0FFEC9" box="[152,248,352,373]" country="Japan" httpUri="http://grbio.org/cool/dk9x-54ca" name="Invertebrate Collection of the Hokkaido University Museum" pageId="30" pageNumber="533">ICHUM</collectionCode>
6325); background body colour pale beige, with pale-brown longitudinal stripes covering entire body (
<figureCitation id="13182CA1EE75ED58FF3CCFE0FEB2FE0F" box="[213,325,413,435]" captionStart="Figure 3" captionStartId="12.[142,222,1745,1767]" captionTargetBox="[222,1342,194,1706]" captionTargetId="figure-25@12.[222,1342,194,1706]" captionTargetPageId="12" captionText="Figure 3. A, Baseodiscus aff. marmoratus (Bürger, 1890) (voucher DNA, ICHUM 6322); B, Baseodiscus aff. maculosus (Bürger, 1895a) (voucher DNA, ICHUM 6324); CE, Baseodiscus delineatus (Delle Chiaje, 1822-1829), ICHUM 6326; F, Baseodiscus cf. curtus (Hubrecht, 1879), ICHUM 6328; G, H, Baseodiscus urgorrii sp. nov., holotype, MCZ IZ-135319, entire body (G) and protruding proboscis (indicated by an arrowhead, H); I, Baseodiscus punnetti (Coe, 1904), specimen different from any voucher specimens in this study; J, K, Baseodiscus kakuii sp. nov., holotype, ICHUM 6334, anterior body (J) and" figureDoi="http://doi.org/10.5281/zenodo.7037968" httpUri="https://zenodo.org/record/7037968/files/figure.png" pageId="30" pageNumber="533">Fig. 3C, D</figureCitation>
) except near mouth (
<figureCitation id="13182CA1EE75ED58FDD1CFE0FD7EFE0E" box="[568,649,413,435]" captionStart="Figure 3" captionStartId="12.[142,222,1745,1767]" captionTargetBox="[222,1342,194,1706]" captionTargetId="figure-25@12.[222,1342,194,1706]" captionTargetPageId="12" captionText="Figure 3. A, Baseodiscus aff. marmoratus (Bürger, 1890) (voucher DNA, ICHUM 6322); B, Baseodiscus aff. maculosus (Bürger, 1895a) (voucher DNA, ICHUM 6324); CE, Baseodiscus delineatus (Delle Chiaje, 1822-1829), ICHUM 6326; F, Baseodiscus cf. curtus (Hubrecht, 1879), ICHUM 6328; G, H, Baseodiscus urgorrii sp. nov., holotype, MCZ IZ-135319, entire body (G) and protruding proboscis (indicated by an arrowhead, H); I, Baseodiscus punnetti (Coe, 1904), specimen different from any voucher specimens in this study; J, K, Baseodiscus kakuii sp. nov., holotype, ICHUM 6334, anterior body (J) and" figureDoi="http://doi.org/10.5281/zenodo.7037968" httpUri="https://zenodo.org/record/7037968/files/figure.png" pageId="30" pageNumber="533">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8B9C3024EE75ED58FF78CFA1FDEBFD74" blockId="30.[145,762,476,712]" pageId="30" pageNumber="533">
<emphasis id="B957EC36EE75ED58FF78CFA1FED4FE4D" box="[145,291,476,497]" italics="true" pageId="30" pageNumber="533">Distribution:</emphasis>
Warm waters worldwide:
<collectingCountry id="F33470B4EE75ED58FDA3CFA1FD6EFE4D" box="[586,665,476,497]" name="France" pageId="30" pageNumber="533">France</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FD4FCFA1FF3FFDAC" author="Joubin L" pageId="30" pageNumber="533" pagination="461 - 602" refId="ref37886" refString="Joubin L. 1890. Recherches sur les turbellaries des cotes de France (Nemertes). Archives de Zoologie Experimentale et Generale, Deuxieme Serie 8: 461 - 602, pls XXV - XXXI." type="journal article" year="1890">Joubin, 1890</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FF3DCF86FEFCFDAC" author="Joubin L" box="[212,267,507,528]" pageId="30" pageNumber="533" refId="ref37923" refString="Joubin L. 1894. Faune Francaise, publiee par les soins de MM. Raphael Blanchard et Jules de Guerne; les Nemertiens. Paris: Societe d'Editions Scientifiques." type="book" year="1894">1894</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FEF6CF87FEA2FDAC" box="[287,341,506,528]" name="Italy" pageId="30" pageNumber="533">Italy</collectingCountry>
(Delle Chiaje, 182229;
<bibRefCitation id="EFB24DD5EE75ED58FD80CF87FD04FDAC" author="Grube AE" box="[617,755,506,528]" pageId="30" pageNumber="533" refId="ref37043" refString="Grube AE. 1840. Actinien, Echinodermen und Wurmer des Adriatischen- und Mittelmeers. Konigsberg [Kaliningrad]: J. H. Bon." type="book" year="1840">Grube, 1840</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FF78CC64FEBFFD92" author="Hubrecht AAW" box="[145,328,537,559]" pageId="30" pageNumber="533" pagination="193 - 232" refId="ref37309" refString="Hubrecht AAW. 1879. The genera of European nemerteans critically revised, with description of several new species. Notes from the Leyden Museum 1: 193 - 232." type="journal article" year="1879">Hubrecht, 1879</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FEBECC67FE09FD93" author="Burger O" box="[343,510,537,559]" pageId="30" pageNumber="533" refId="ref34395" refString="Burger O. 1895 a. Die Nemertinen des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel und der Angrenzenden Meeres-Abschnitte," type="book" year="1895">Bürger, 1895a</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FDE4CC64FD1EFD93" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" box="[525,745,537,559]" pageId="30" pageNumber="533" pagination="3785 - 3793" refId="ref41221" refString="Strand M, Hjelmgren A, Sundberg P. 2005. Genus Baseodiscus (Nemertea: Heteronemertea): molecular identification of a new species in a phylogenetic context. Journal of Natural History 39: 3785 - 3793." type="journal article" year="2005">
Strand
<emphasis id="B957EC36EE75ED58FD8ECC67FD54FD92" box="[615,675,537,559]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2005
</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FF78CC45FF0AFDF2" box="[145,253,568,590]" name="Australia" pageId="30" pageNumber="533">Australia</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FEE3CC45FE2FFDF2" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" box="[266,472,568,590]" pageId="30" pageNumber="533" pagination="3785 - 3793" refId="ref41221" refString="Strand M, Hjelmgren A, Sundberg P. 2005. Genus Baseodiscus (Nemertea: Heteronemertea): molecular identification of a new species in a phylogenetic context. Journal of Natural History 39: 3785 - 3793." type="journal article" year="2005">
Strand
<emphasis id="B957EC36EE75ED58FEB6CC44FE62FDF1" box="[351,405,568,590]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2005
</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FE0ACC45FD68FDF1" author="Kvist S &amp; Laumer CE &amp; Junoy J &amp; Giribet G" box="[483,671,568,590]" pageId="30" pageNumber="533" pagination="287 - 308" refId="ref39075" refString="Kvist S, Laumer CE, Junoy J, Giribet G. 2014. New insights into the phylogeny, systematics and DNA barcoding of Nemertea. Invertebrate Systematics 28: 287 - 308." type="journal article" year="2014">
Kvist
<emphasis id="B957EC36EE75ED58FDCFCC44FDABFDF1" box="[550,604,568,590]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2014
</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FD5ACC45FD0EFDF1" box="[691,761,568,589]" name="Japan" pageId="30" pageNumber="533">Japan</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FF70CC2BFEBDFDD0" author="Kajihara H" box="[153,330,598,620]" pageId="30" pageNumber="533" pagination="419 - 444" refId="ref38160" refString="Kajihara H. 2017. Species diversity of Japanese ribbon worms (Nemertea). In: Motokawa M, Kajihara H, eds. Species diversity of animals in Japan. Tokyo: Springer Japan, 419 - 444." type="book chapter" year="2017">Kajihara, 2017</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FEB0CC2BFDB3FDD0" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" box="[345,580,598,620]" pageId="30" pageNumber="533" pagination="348 - 353" refId="ref37407" refString="Ikenaga J, Hookabe N, Kohtsuka H, Yoshida M, Kajihara H. 2019. A population without females: males of Baseodiscus delineatus (Nemertea: Heteronemertea) reproduce asexually by fragmentation. Zoological Science 36: 348 - 353." type="journal article" year="2019">
Ikenaga
<emphasis id="B957EC36EE75ED58FE2BCC2AFE09FDD7" box="[450,510,598,620]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2019
</bibRefCitation>
),
<collectingRegion id="49E7FEC6EE75ED58FDB4CC2BFD42FDD0" box="[605,693,598,620]" country="United States of America" name="Florida" pageId="30" pageNumber="533">Florida</collectingRegion>
,
<collectingCountry id="F33470B4EE75ED58FD2BCC2AFD0FFDD0" box="[706,760,599,620]" name="United States of America" pageId="30" pageNumber="533">USA</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FF73CC08FEA5FD36" author="Schwartz ML" box="[154,338,629,651]" pageId="30" pageNumber="533" refId="ref40578" refString="Schwartz ML. 2009. Untying a Gordian knot of worms: systematics and taxonomy of the Pilidiophora (phylum Nemertea) from multiple data sets. Unpublished D. Phil. Thesis, The George Washington University." type="book" year="2009">Schwartz, 2009</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FE82CC08FE43FD37" box="[363,436,629,651]" name="Belize" pageId="30" pageNumber="533">Belize</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FE2FCC08FD88FD36" author="Schwartz ML" box="[454,639,629,651]" pageId="30" pageNumber="533" refId="ref40578" refString="Schwartz ML. 2009. Untying a Gordian knot of worms: systematics and taxonomy of the Pilidiophora (phylum Nemertea) from multiple data sets. Unpublished D. Phil. Thesis, The George Washington University." type="book" year="2009">Schwartz, 2009</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FD71CC0BFD0FFD37" box="[664,760,630,651]" name="Panama" pageId="30" pageNumber="533">Panama</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FF73CCE9FEA5FD15" author="Schwartz ML" box="[154,338,660,682]" pageId="30" pageNumber="533" refId="ref40578" refString="Schwartz ML. 2009. Untying a Gordian knot of worms: systematics and taxonomy of the Pilidiophora (phylum Nemertea) from multiple data sets. Unpublished D. Phil. Thesis, The George Washington University." type="book" year="2009">Schwartz, 2009</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FE85CCE9FE2AFD16" box="[364,477,660,682]" name="Colombia" pageId="30" pageNumber="533">Colombia</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FE06CCE9FF3FFD74" author="Gonzalez-Cueto J &amp; Quiroga S &amp; Norenburg J" pageId="30" pageNumber="533" pagination="83 - 108" refId="ref36953" refString="Gonzalez-Cueto J, Quiroga S, Norenburg J. 2014. A shore-based preliminary survey of marine ribbon worms (Nemertea) from the Caribbean coast of Colombia. ZooKeys 439: 83 - 108." type="journal article" year="2014">
Gonzalez-Cueto
<emphasis id="B957EC36EE75ED58FD5CCCE8FD04FD15" box="[693,755,660,682]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2014
</bibRefCitation>
),
<collectingCountry id="F33470B4EE75ED58FF34CCCFFED4FD74" box="[221,291,690,712]" name="Brazil" pageId="30" pageNumber="533">Brazil</collectingCountry>
(
<bibRefCitation id="EFB24DD5EE75ED58FEDBCCCFFDFAFD74" author="Mendes CB &amp; Matthews-Cascon H &amp; Norenburg JL" box="[306,525,690,712]" pageId="30" pageNumber="533" pagination="146 - 156" refId="ref39466" refString="Mendes CB, Matthews-Cascon H, Norenburg JL. 2016. New records of ribbon worms (Nemertea) from Ceara, northeast Brazil. Zootaxa 4061: 146 - 156." type="journal article" year="2016">
Mendes
<emphasis id="B957EC36EE75ED58FE78CCCEFE3EFD7B" box="[401,457,690,712]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2016
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9C3024EE75ED58FF78CC8CFD95FAD0" blockId="30.[144,762,753,1879]" pageId="30" pageNumber="533">
<emphasis id="B957EC36EE75ED58FF78CC8CFEFEFCBA" box="[145,265,753,774]" italics="true" pageId="30" pageNumber="533">Remarks:</emphasis>
In accordance with Article 12.2.7 of the
<emphasis id="B957EC36EE75ED58FF2BCD6DFF0AFC99" box="[194,253,784,805]" italics="true" pageId="30" pageNumber="533">Code</emphasis>
(
<bibRefCitation id="EFB24DD5EE75ED58FEF9CD6DFE6CFC99" author="ICZN" box="[272,411,784,805]" pageId="30" pageNumber="533" refId="ref37385" refString="ICZN. 1999. International code of zoological nomenclature, 4 th edn. London: International Trust for Zoological Nomenclature." type="book" year="1999">ICZN, 1999</bibRefCitation>
), the year of publication for
<taxonomicName id="4C234BA7EE75ED58FF78CD52FED8FCFF" authorityName=": Coe" authorityYear="1947" box="[145,303,814,836]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FF78CD52FED8FCFF" box="[145,303,814,836]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
should be 1822, rather than 1825 as previously regarded (e.g.
<bibRefCitation id="EFB24DD5EE75ED58FE5FCD30FF3FFC3D" author="Magarlamov TY &amp; Chernyshev AV" pageId="30" pageNumber="533" pagination="440 - 445" refId="ref39268" refString="Magarlamov TY, Chernyshev AV. 2011. The structure of the proboscis musculature in Baseodiscus delineatus (Delle Chiaje, 1825) (Heteronemertea) and the comparative analysis of the proboscis musculature in heteronemerteans. Russian Journal of Marine Biology 37: 440 - 445." type="journal article" year="2011">Magarlamov &amp; Chernyshev, 2011</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FF3CCD11FE40FC3D" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" box="[213,439,876,898]" pageId="30" pageNumber="533" pagination="348 - 353" refId="ref37407" refString="Ikenaga J, Hookabe N, Kohtsuka H, Yoshida M, Kajihara H. 2019. A population without females: males of Baseodiscus delineatus (Nemertea: Heteronemertea) reproduce asexually by fragmentation. Zoological Science 36: 348 - 353." type="journal article" year="2019">
Ikenaga
<emphasis id="B957EC36EE75ED58FED3CD10FE84FC3D" box="[314,371,876,898]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2019
</bibRefCitation>
). The species was originally established as
<taxonomicName id="4C234BA7EE75ED58FED7CDF7FE1AFC23" authorityName="Delle Chiaje" authorityYear="1825" box="[318,493,906,927]" class="Pilidiophora" family="Lineidae" genus="Polia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE75ED58FED7CDF7FE1AFC23" box="[318,493,906,927]" italics="true" pageId="30" pageNumber="533">Polia delineata</emphasis>
</taxonomicName>
in Delle Chiaje (1822 29), a work comprised of plates published in 1822 and four text volumes, with Volumes 1 and 2 issued in 1825 [or, Volume 2 may have been published in 1827 (
<bibRefCitation id="EFB24DD5EE75ED58FF70CA7BFE91FBA6" author="Neave SA" box="[153,358,1029,1051]" pageId="30" pageNumber="533" refId="ref39798" refString="Neave SA. 1940. Nomenclator zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus 1758 to the end of 1935, Vol. 3. M - P. London: Zoological Society of London." type="book" year="1940">Neave, 1940: 344</bibRefCitation>
)], Volume
<quantity id="4CDB9DC1EE75ED58FE08CA78FDF9FBA7" box="[481,526,1029,1051]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="30" pageNumber="533" unit="in" value="3.0">3 in</quantity>
1828 and Volume
<quantity id="4CDB9DC1EE75ED58FD02CA78FF5FFB86" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="30" pageNumber="533" unit="in" value="4.0">4 in</quantity>
1829. The description in words appeared in Volume 2 on p. 409 (as
<taxonomicName id="4C234BA7EE75ED58FEBACA3EFE32FBEB" box="[339,453,1090,1112]" class="Pilidiophora" family="Lineidae" genus="Polia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="lineata">
<emphasis id="B957EC36EE75ED58FEBACA3EFE32FBEB" box="[339,453,1090,1112]" italics="true" pageId="30" pageNumber="533">P. lineata</emphasis>
</taxonomicName>
) and pp. 427428 (
<bibRefCitation id="EFB24DD5EE75ED58FD50CA3FFED5FBCB" author="Delle Chiaje S" pageId="30" pageNumber="533" pagination="185 - 444" refId="ref35980" refString="Delle Chiaje S. 1822 - 1829. Memorie sulla storia e notomia degli animali senza vertebre del regno di Napoli. Naples: Societa' Tipografica [pls I - CIX published in 1822; Vol. 1, xii + 184 pp., in 1825; Vol. 2, 260 pp. (pp. 185 - 444), in 1825; Vol. 3, xx + 232 pp., in 1828; Vol. 4, viii + 214 pp., in 1829]." type="book chapter" year="1825">
Delle Chiaje,
<quantity id="4CDB9DC1EE75ED58FF03CA1CFED5FBCB" box="[234,290,1121,1143]" metricMagnitude="1" metricUnit="m" metricValue="4.6355" pageId="30" pageNumber="533" unit="in" value="1825.0">1825</quantity>
</bibRefCitation>
in 182229), although the species name
<taxonomicName id="4C234BA7EE75ED58FF78CAFDFEB5FB29" authorityName="Delle Chiaje" authorityYear="1825" box="[145,322,1152,1173]" class="Pilidiophora" family="Lineidae" genus="Polia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE75ED58FF78CAFDFEB5FB29" box="[145,322,1152,1173]" italics="true" pageId="30" pageNumber="533">Polia delineata</emphasis>
</taxonomicName>
had already appeared as the caption for an plate illustration published in 1822. Delle Chiajes 1822 plate numbers are discontinuous in several places: plate numbers XXX (30),
<collectionCode id="ED32A8E1EE75ED58FDBFCAA1FD61FB4D" box="[598,662,1244,1265]" pageId="30" pageNumber="533">LXIII</collectionCode>
(63) and CII (102) are missing; in addition, for each of the four numbers I (1), II (2),
<collectionCode id="ED32A8E1EE75ED58FE69CB67FE24FA93" box="[384,467,1306,1327]" pageId="30" pageNumber="533">XXVIII</collectionCode>
(28) and XXXIV (34), two different plates exist.
<taxonomicName id="4C234BA7EE75ED58FE6FCB45FDD9FAF1" authorityName="Delle Chiaje" authorityYear="1825" box="[390,558,1336,1357]" class="Pilidiophora" family="Lineidae" genus="Polia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE75ED58FE6FCB45FDD9FAF1" box="[390,558,1336,1357]" italics="true" pageId="30" pageNumber="533">Polia delineata</emphasis>
</taxonomicName>
is depicted on the second of the two plates labelled
<collectionCode id="ED32A8E1EE75ED58FDEECB2AFDAAFAD0" box="[519,605,1367,1388]" pageId="30" pageNumber="533">XXVIII</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C33963AFEE75ED58FF40CB08FBD5FC65" pageId="30" pageNumber="533" type="reference_group">
<paragraph id="8B9C3024EE75ED58FF40CB08FAE3FE2F" blockId="30.[144,762,753,1879]" lastBlockId="30.[809,1426,197,1231]" pageId="30" pageNumber="533">
<taxonomicName id="4C234BA7EE75ED58FF40CB08FE40FA36" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[169,439,1397,1418]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FF40CB08FE40FA36" box="[169,439,1397,1418]" italics="true" pageId="30" pageNumber="533">Baseodiscus delineatus</emphasis>
</taxonomicName>
has a pale background body colour with numerous brown longitudinal stripes. The broad range of variation in colour pattern has obscured the species identity of
<taxonomicName id="4C234BA7EE75ED58FDBCCBAFFD05FA5A" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[597,754,1489,1511]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FDBCCBAFFD05FA5A" box="[597,754,1489,1511]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
, especially in relation to a similar form,
<taxonomicName id="4C234BA7EE75ED58FD60CB8CFEADF998" ID-CoL="68D6P" authority="(Hubrecht, 1879)" baseAuthorityName="Hubrecht" baseAuthorityYear="1879" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="curtus">
<emphasis id="B957EC36EE75ED58FD60CB8CFD0FF9BA" box="[649,760,1521,1542]" italics="true" pageId="30" pageNumber="533">B. curtus</emphasis>
(
<bibRefCitation id="EFB24DD5EE75ED58FF70C873FEB9F998" author="Hubrecht AAW" box="[153,334,1550,1572]" pageId="30" pageNumber="533" pagination="193 - 232" refId="ref37309" refString="Hubrecht AAW. 1879. The genera of European nemerteans critically revised, with description of several new species. Notes from the Leyden Museum 1: 193 - 232." type="journal article" year="1879">Hubrecht, 1879</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C234BA7EE75ED58FE8DC873FDB4F998" ID-CoL="68D6P" baseAuthorityName="Hubrecht" baseAuthorityYear="1879" box="[356,579,1550,1572]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="curtus">
<emphasis id="B957EC36EE75ED58FE8DC873FDB4F998" box="[356,579,1550,1572]" italics="true" pageId="30" pageNumber="533">Baseodiscus curtus</emphasis>
</taxonomicName>
was previously regarded as a variety of
<taxonomicName id="4C234BA7EE75ED58FE2FC853FD91F9FE" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[454,614,1581,1603]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FE2FC853FD91F9FE" box="[454,614,1581,1603]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB24DD5EE75ED58FD93C850FD05F9FE" author="Coe WR" box="[634,754,1581,1602]" pageId="30" pageNumber="533" pagination="247 - 322" refId="ref35559" refString="Coe WR. 1940. Revision of the nemertean fauna of the Pacific coasts of North, Central, and northern South America. Allan Hancock Pacific Expeditions 2: 247 - 322." type="journal article" year="1940">Coe, 1940</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FF78C831FF38F9DD" author="Coe WR" box="[145,207,1612,1633]" pageId="30" pageNumber="533" pagination="27 - 32" refId="ref35593" refString="Coe WR. 1944. Geographic distribution of the nemerteans of the Pacific coast of North America, with descriptions of two new species. Journal of the Washington Academy of Sciences 34: 27 - 32." type="journal article" year="1944">1944</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FF0CC831FE63F9DD" author="Correa DD" box="[229,404,1612,1634]" pageId="30" pageNumber="533" pagination="441 - 455" refId="ref35723" refString="Correa DD. 1958. Nemertinos do litoral Brasileiro (VII). Anais da Academia Brasileira de Siencias 29: 441 - 455, pls 1 - 4." type="journal article" year="1958">Corrêa, 1958</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FE42C831FE1EF9DD" author="Correa DD" box="[427,489,1612,1634]" pageId="30" pageNumber="533" pagination="1 - 44" refId="ref35753" refString="Correa DD. 1961. Nemerteans from Florida and Virgin Islands. Bulletin of Marine Science of the Gulf and Caribbean 11: 1 - 44." type="journal article" year="1961">1961</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FE16C831FDC9F9DD" author="Correa DD" box="[511,574,1612,1634]" pageId="30" pageNumber="533" pagination="41 - 56" refId="ref35780" refString="Correa DD. 1963. Nemerteans from Curacao. Studies on the Fauna of Curacao and Other Caribbean Islands 17: 41 - 56." type="journal article" year="1963">1963</bibRefCitation>
) and thus as synonymous with it (
<bibRefCitation id="EFB24DD5EE75ED58FE65C817FDD1F93C" author="Gibson R" box="[396,550,1642,1664]" pageId="30" pageNumber="533" pagination="137 - 160" refId="ref36552" refString="Gibson R. 1979. Nemerteans of the Great Barrier Reef 2. Anopla Heteronemertea (Baseodiscidae). Zoological Journal of the Linnean Society 66: 137 - 160." type="journal article" year="1979">Gibson, 1979</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FDDCC817FD99F93C" author="Gibson R" box="[565,622,1642,1664]" pageId="30" pageNumber="533" pagination="271 - 561" refId="ref36771" refString="Gibson R. 1995. Nemertean genera and species of the world: an annotated checklist of original names and description citations, synonyms, current taxonomic status, habitats and recorded zoogeographic distribution. Journal of Natural History 29: 271 - 561." type="journal article" year="1995">1995</bibRefCitation>
). Moreover,
<bibRefCitation id="EFB24DD5EE75ED58FF78C8F7FEC2F922" author="Burger O" box="[145,309,1673,1695]" pageId="30" pageNumber="533" pagination="1 - 151" refId="ref34610" refString="Burger O. 1904. Nemertini. Das Tierreich 20: 1 - 151." type="journal article" year="1904">Bürger (1904)</bibRefCitation>
and
<bibRefCitation id="EFB24DD5EE75ED58FE86C8F4FDFEF922" author="Gibson R" box="[367,521,1673,1695]" pageId="30" pageNumber="533" pagination="137 - 160" refId="ref36552" refString="Gibson R. 1979. Nemerteans of the Great Barrier Reef 2. Anopla Heteronemertea (Baseodiscidae). Zoological Journal of the Linnean Society 66: 137 - 160." type="journal article" year="1979">Gibson (1979</bibRefCitation>
,
<bibRefCitation id="EFB24DD5EE75ED58FDFEC8F4FDB8F923" author="Gibson R" box="[535,591,1673,1695]" pageId="30" pageNumber="533" pagination="271 - 561" refId="ref36771" refString="Gibson R. 1995. Nemertean genera and species of the world: an annotated checklist of original names and description citations, synonyms, current taxonomic status, habitats and recorded zoogeographic distribution. Journal of Natural History 29: 271 - 561." type="journal article" year="1995">1995</bibRefCitation>
) synonymized the following six nominal species originally described from the Indo-Pacific with
<taxonomicName id="4C234BA7EE75ED58FE2FC8BAFDA9F967" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[454,606,1734,1756]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FE2FC8BAFDA9F967" box="[454,606,1734,1756]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
:
<taxonomicName id="4C234BA7EE75ED58FD83C8BBFD97F946" ID-CoL="84XNV" authority="Punnett &amp; Cooper, 1909" authorityName="Punnett &amp; Cooper" authorityYear="1909" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="insignis">
<emphasis id="B957EC36EE75ED58FD83C8BBFF0AF947" italics="true" pageId="30" pageNumber="533">Baseodiscus insignis</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FEE4C89BFD97F946" author="Punnett RC &amp; Cooper CF" box="[269,608,1765,1787]" pageId="30" pageNumber="533" pagination="1 - 15" refId="ref40244" refString="Punnett RC, Cooper CF. 1909. On some nemerteans from the eastern Indian Ocean. Transactions of the Linnean Society of London, 2 nd Series 13: 1 - 15." type="book chapter" year="1909">Punnett &amp; Cooper, 1909</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F33470B4EE75ED58FD93C89BFD05F947" box="[634,754,1765,1787]" name="Tanzania" pageId="30" pageNumber="533">Zanzibar</collectingCountry>
,
<collectingCountry id="F33470B4EE75ED58FF78C979FEF3F8A6" box="[145,260,1796,1818]" name="Tanzania" pageId="30" pageNumber="533">Tanzania</collectingCountry>
);
<taxonomicName id="4C234BA7EE75ED58FEF0C979FF1AF884" authority="Quoy &amp; Gaimard, 1833" authorityName="Quoy &amp; Gaimard" authorityYear="1833" class="Pilidiophora" family="Lineidae" genus="Borlasia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="striata">
<emphasis id="B957EC36EE75ED58FEF0C979FE2EF8A5" box="[281,473,1796,1817]" italics="true" pageId="30" pageNumber="533">Borlasia striata</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FE0BC979FD0EF8A5" author="Quoy JRC &amp; Gaimard JP" box="[482,761,1796,1818]" pageId="30" pageNumber="533" refId="ref40383" refString="Quoy JRC, Gaimard JP. 1833. Voyage de decouvertes de l'Astrolabe execute par ordre du Roi, pendant les annees 1826 - 1827 - 1828 - 1829, sou le commandement de M. J. Dumont d'Urville, Zoologie, tome quatrieme. Paris: Tastu." type="book" year="1833">Quoy &amp; Gaimard, 1833</bibRefCitation>
(
<collectingCountry id="F33470B4EE75ED58FF70C95EFF12F884" box="[153,229,1827,1848]" name="Guam" pageId="30" pageNumber="533">Guam</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="4C234BA7EE75ED58FF13C95FFD7AF884" ID-CoL="853QN" authority="Staub, 1900" authorityName="Staub" authorityYear="1900" box="[250,653,1826,1848]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="amboinensis">
<emphasis id="B957EC36EE75ED58FF13C95FFE01F88B" box="[250,502,1826,1847]" italics="true" pageId="30" pageNumber="533">Eupolia amboinensis</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FE17C95FFD7AF884" author="Staub J" box="[510,653,1826,1848]" pageId="30" pageNumber="533" pagination="591 - 614" refId="ref40883" refString="Staub J. 1900. Neue Nemertinen aus Amboina. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 8: 591 - 614 [75 - 98 in an alternative pagination]." type="journal article" year="1900">Staub, 1900</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F33470B4EE75ED58FD74C95FFD03F884" box="[669,756,1826,1848]" name="Indonesia" pageId="30" pageNumber="533">Ambon</collectingCountry>
,
<collectingCountry id="F33470B4EE75ED58FF78C93CFEFFF8EB" box="[145,264,1857,1879]" name="Indonesia" pageId="30" pageNumber="533">Indonesia</collectingCountry>
);
<taxonomicName id="4C234BA7EE75ED58FEF3C93CFD67F8EA" ID-CoL="853QQ" authority="Burger, 1890" authorityName="Burger" authorityYear="1890" box="[282,656,1857,1879]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="ascophora">
<emphasis id="B957EC36EE75ED58FEF3C93CFE03F8EA" box="[282,500,1857,1878]" italics="true" pageId="30" pageNumber="533">Eupolia ascophora</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FE12C93FFD67F8EA" author="Burger O" box="[507,656,1857,1879]" pageId="30" pageNumber="533" pagination="1 - 277" refId="ref34295" refString="Burger O. 1890. Untersuchungen uber die Anatomie und Histologie der Nemertinen nebst Beitragen zur Systematik. Zeitschrift fur Wissenschaftliche Zoologie 50: 1 - 277, pls I - X." type="journal article" year="1890">Bürger, 1890</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F33470B4EE75ED58FD77C93CFD03F8EB" box="[670,756,1857,1879]" name="Indonesia" pageId="30" pageNumber="533">Ambon</collectingCountry>
,
<collectingCountry id="F33470B4EE75ED58FCC0CEBBFC5CFF67" box="[809,939,197,219]" name="Indonesia" pageId="30" pageNumber="533">Indonesia</collectingCountry>
);
<taxonomicName id="4C234BA7EE75ED58FC25CEBBFA66FF66" authority="Burger, 1890" authorityName="Burger" authorityYear="1890" box="[972,1425,197,219]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="marmorata">
<emphasis id="B957EC36EE75ED58FC25CEBBFB22FF67" box="[972,1237,197,219]" italics="true" pageId="30" pageNumber="533">Eupolia marmorata</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FB0ACEBBFA66FF66" author="Burger O" box="[1251,1425,197,219]" pageId="30" pageNumber="533" pagination="1 - 277" refId="ref34295" refString="Burger O. 1890. Untersuchungen uber die Anatomie und Histologie der Nemertinen nebst Beitragen zur Systematik. Zeitschrift fur Wissenschaftliche Zoologie 50: 1 - 277, pls I - X." type="journal article" year="1890">Bürger, 1890</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F33470B4EE75ED58FCD9CE99FC7FFF46" box="[816,904,228,250]" name="Indonesia" pageId="30" pageNumber="533">Ambon</collectingCountry>
,
<collectingCountry id="F33470B4EE75ED58FC7FCE99FBE7FF46" box="[918,1040,228,250]" name="Indonesia" pageId="30" pageNumber="533">Indonesia</collectingCountry>
); and
<taxonomicName id="4C234BA7EE75ED58FBB5CE99FC95FEA4" authority="Staub, 1900" authorityName="Staub" authorityYear="1900" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="reticulata">
<emphasis id="B957EC36EE75ED58FBB5CE99FACDFF45" box="[1116,1338,228,249]" italics="true" pageId="30" pageNumber="533">Eupolia reticulata</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FAAACE99FC95FEA4" author="Staub J" pageId="30" pageNumber="533" pagination="591 - 614" refId="ref40883" refString="Staub J. 1900. Neue Nemertinen aus Amboina. Denkschriften der Medicinisch-Naturwissenschaftlichen Gesellschaft zu Jena 8: 591 - 614 [75 - 98 in an alternative pagination]." type="journal article" year="1900">Staub, 1900</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F33470B4EE75ED58FC98CF7FFC3FFEA4" box="[881,968,258,280]" name="Indonesia" pageId="30" pageNumber="533">Ambon</collectingCountry>
,
<collectingCountry id="F33470B4EE75ED58FC3CCF7FFBB9FEA4" box="[981,1102,258,280]" name="Indonesia" pageId="30" pageNumber="533">Indonesia</collectingCountry>
). This synonymization was probably an excessive lumping, as
<bibRefCitation id="EFB24DD5EE75ED58FB07CF5CFC81FEEA" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" pageId="30" pageNumber="533" pagination="3785 - 3793" refId="ref41221" refString="Strand M, Hjelmgren A, Sundberg P. 2005. Genus Baseodiscus (Nemertea: Heteronemertea): molecular identification of a new species in a phylogenetic context. Journal of Natural History 39: 3785 - 3793." type="journal article" year="2005">
Strand
<emphasis id="B957EC36EE75ED58FAB9CF5FFA66FE8A" box="[1360,1425,289,311]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
(2005)
</bibRefCitation>
discovered a new species,
<taxonomicName id="4C234BA7EE75ED58FB27CF3CFAB1FEE9" ID-CoL="5WCTT" box="[1230,1350,320,342]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="jonasii">
<emphasis id="B957EC36EE75ED58FB27CF3CFAB1FEE9" box="[1230,1350,320,342]" italics="true" pageId="30" pageNumber="533">B. jonasii</emphasis>
</taxonomicName>
, from
<collectingRegion id="49E7FEC6EE75ED58FCC0CF23FC36FEC8" box="[809,961,350,372]" country="Solomon Islands" name="Guadalcanal" pageId="30" pageNumber="533">Guadalcanal</collectingRegion>
(
<collectingCountry id="F33470B4EE75ED58FC39CF23FB60FEC8" box="[976,1175,350,372]" name="Solomon Islands" pageId="30" pageNumber="533">Solomon Islands</collectingCountry>
) that resembles, but is genetically distinct from,
<taxonomicName id="4C234BA7EE75ED58FB9DCF03FAFAFE2E" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[1140,1293,381,403]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FB9DCF03FAFAFE2E" box="[1140,1293,381,403]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C3024EE75ED58FCA8CFE1FBD5FC65" blockId="30.[809,1426,197,1231]" pageId="30" pageNumber="533">
Owing to the 16S gene sequence from
<collectingCountry id="F33470B4EE75ED58FABBCFE1FA79FE0E" box="[1362,1422,412,434]" name="Italy" pageId="30" pageNumber="533">Italy</collectingCountry>
(
<accessionNumber id="9470ADC7EE75ED58FCD9CFC7FC47FE6C" box="[816,944,442,464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY955227" pageId="30" pageNumber="533">AY955227</accessionNumber>
) provided by
<bibRefCitation id="EFB24DD5EE75ED58FB89CFC7FAA0FE6C" author="Strand M &amp; Hjelmgren A &amp; Sundberg P" box="[1120,1367,442,464]" pageId="30" pageNumber="533" pagination="3785 - 3793" refId="ref41221" refString="Strand M, Hjelmgren A, Sundberg P. 2005. Genus Baseodiscus (Nemertea: Heteronemertea): molecular identification of a new species in a phylogenetic context. Journal of Natural History 39: 3785 - 3793." type="journal article" year="2005">
Strand
<emphasis id="B957EC36EE75ED58FB29CFC6FB08FE73" box="[1216,1279,442,464]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
(2005)
</bibRefCitation>
, the species identity of
<taxonomicName id="4C234BA7EE75ED58FBEBCFA7FB60FE52" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[1026,1175,473,495]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FBEBCFA7FB60FE52" box="[1026,1175,473,495]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
has been established more firmly than before, enabling the application of reliable names to material from different localities. The six 16S sequences from four different countries (
<accessionNumber id="9470ADC7EE75ED58FCD9CC29FC59FDD6" box="[816,942,596,618]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY955227" pageId="30" pageNumber="533">AY955227</accessionNumber>
,
<collectingCountry id="F33470B4EE75ED58FC52CC29FC00FDD6" box="[955,1015,596,618]" name="Italy" pageId="30" pageNumber="533">Italy</collectingCountry>
;
<accessionNumber id="9470ADC7EE75ED58FBEDCC29FB77FDD6" box="[1028,1152,596,618]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY955232" pageId="30" pageNumber="533">AY955232</accessionNumber>
,
<accessionNumber id="9470ADC7EE75ED58FB64CC29FAFCFDD6" box="[1165,1291,596,618]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF935448" pageId="30" pageNumber="533">KF935448</accessionNumber>
,
<collectingCountry id="F33470B4EE75ED58FAF1CC29FA7BFDD6" box="[1304,1420,596,618]" name="Australia" pageId="30" pageNumber="533">Australia</collectingCountry>
;
<accessionNumber id="9470ADC7EE75ED58FCC0CC0FFC5DFD34" box="[809,938,626,648]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178665" pageId="30" pageNumber="533">LC178665</accessionNumber>
,
<accessionNumber id="9470ADC7EE75ED58FC53CC0FFBCCFD34" box="[954,1083,626,648]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC178666" pageId="30" pageNumber="533">LC178666</accessionNumber>
,
<collectingCountry id="F33470B4EE75ED58FBA2CC0EFB6EFD34" box="[1099,1177,627,648]" name="Japan" pageId="30" pageNumber="533">Japan</collectingCountry>
;
<accessionNumber id="9470ADC7EE75ED58FB40CC0FFADDFD34" box="[1193,1322,626,648]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EF124861" pageId="30" pageNumber="533">EF124861</accessionNumber>
,
<collectingCountry id="F33470B4EE75ED58FAD2CC0FFA7FFD34" box="[1339,1416,626,648]" name="Belize" pageId="30" pageNumber="533">Belize</collectingCountry>
) were identical (
<emphasis id="B957EC36EE75ED58FC13CCEFFBFFFD1B" box="[1018,1032,658,679]" italics="true" pageId="30" pageNumber="533">p</emphasis>
-distance = 0.00). In addition, recent technological progress in high-quality digital photography allows comparison of subtle differences in body coloration and markings between specimens from different localities (e.g.
<bibRefCitation id="EFB24DD5EE75ED58FB54CD71FA7EFC9D" author="Schwartz ML" box="[1213,1417,780,802]" pageId="30" pageNumber="533" refId="ref40578" refString="Schwartz ML. 2009. Untying a Gordian knot of worms: systematics and taxonomy of the Pilidiophora (phylum Nemertea) from multiple data sets. Unpublished D. Phil. Thesis, The George Washington University." type="book" year="2009">Schwartz, 2009</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FCC0CD57FB75FCFC" author="Gonzalez-Cueto J &amp; Quiroga S &amp; Norenburg J" box="[809,1154,810,832]" pageId="30" pageNumber="533" pagination="83 - 108" refId="ref36953" refString="Gonzalez-Cueto J, Quiroga S, Norenburg J. 2014. A shore-based preliminary survey of marine ribbon worms (Nemertea) from the Caribbean coast of Colombia. ZooKeys 439: 83 - 108." type="journal article" year="2014">
Gonzalez-Cueto
<emphasis id="B957EC36EE75ED58FC1FCD56FBC1FC83" box="[1014,1078,810,832]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2014
</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FB7DCD57FA7EFCFC" author="Mendes CB &amp; Matthews-Cascon H &amp; Norenburg JL" box="[1172,1417,810,832]" pageId="30" pageNumber="533" pagination="146 - 156" refId="ref39466" refString="Mendes CB, Matthews-Cascon H, Norenburg JL. 2016. New records of ribbon worms (Nemertea) from Ceara, northeast Brazil. Zootaxa 4061: 146 - 156." type="journal article" year="2016">
Mendes
<emphasis id="B957EC36EE75ED58FB17CD56FACAFC83" box="[1278,1341,810,832]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2016
</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FCC0CD34FC23FCE2" author="Kajihara H" box="[809,980,841,863]" pageId="30" pageNumber="533" pagination="419 - 444" refId="ref38160" refString="Kajihara H. 2017. Species diversity of Japanese ribbon worms (Nemertea). In: Motokawa M, Kajihara H, eds. Species diversity of animals in Japan. Tokyo: Springer Japan, 419 - 444." type="book chapter" year="2017">Kajihara, 2017</bibRefCitation>
;
<bibRefCitation id="EFB24DD5EE75ED58FC0BCD34FB32FCE2" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" box="[994,1221,841,863]" pageId="30" pageNumber="533" pagination="348 - 353" refId="ref37407" refString="Ikenaga J, Hookabe N, Kohtsuka H, Yoshida M, Kajihara H. 2019. A population without females: males of Baseodiscus delineatus (Nemertea: Heteronemertea) reproduce asexually by fragmentation. Zoological Science 36: 348 - 353." type="journal article" year="2019">
Ikenaga
<emphasis id="B957EC36EE75ED58FBAECD37FB76FCE2" box="[1095,1153,841,863]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2019
</bibRefCitation>
). These new data sources now indicate that
<taxonomicName id="4C234BA7EE75ED58FBB0CD14FB19FCC1" ID-CoL="5WCTV" authorityName=": Coe" authorityYear="1947" box="[1113,1262,872,894]" class="Pilidiophora" family="Valenciniidae" genus="Baseodiscus" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineatus">
<emphasis id="B957EC36EE75ED58FBB0CD14FB19FCC1" box="[1113,1262,872,894]" italics="true" pageId="30" pageNumber="533">B. delineatus</emphasis>
</taxonomicName>
is distributed circumglobally. The species can reproduce asexually by fragmentation followed by anterior regeneration (
<bibRefCitation id="EFB24DD5EE75ED58FCD8CDB9FBE5FC65" author="Ikenaga J &amp; Hookabe N &amp; Kohtsuka H &amp; Yoshida M &amp; Kajihara H" box="[817,1042,964,986]" pageId="30" pageNumber="533" pagination="348 - 353" refId="ref37407" refString="Ikenaga J, Hookabe N, Kohtsuka H, Yoshida M, Kajihara H. 2019. A population without females: males of Baseodiscus delineatus (Nemertea: Heteronemertea) reproduce asexually by fragmentation. Zoological Science 36: 348 - 353." type="journal article" year="2019">
Ikenaga
<emphasis id="B957EC36EE75ED58FC7FCDB8FC39FC65" box="[918,974,964,986]" italics="true" pageId="30" pageNumber="533">et al.</emphasis>
, 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33963AFEE75ED58FCA8CD9EFB9EFB72" pageId="30" pageNumber="533" type="description">
<paragraph id="8B9C3024EE75ED58FCA8CD9EFB9EFB72" blockId="30.[809,1426,197,1231]" pageId="30" pageNumber="533">
<bibRefCitation id="EFB24DD5EE75ED58FCA8CD9EFBA2FC44" author="Burger O" box="[833,1109,994,1016]" pageId="30" pageNumber="533" refId="ref34395" refString="Burger O. 1895 a. Die Nemertinen des Golfes von Neapel und der angrenzenden Meeres-Abschnitte. Fauna und Flora des Golfes von Neapel und der Angrenzenden Meeres-Abschnitte," type="book" year="1895">Bürger (1895a: 600)</bibRefCitation>
synonymized
<taxonomicName id="4C234BA7EE75ED58FAC9CD9EFB80FBAB" authority="Quatrefages, 1846" authorityName="Quatrefages" authorityYear="1846" class="Pilidiophora" family="Lineidae" genus="Borlasia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="carmellina">
<emphasis id="B957EC36EE75ED58FAC9CD9EFC53FBAA" italics="true" pageId="30" pageNumber="533">Borlasia carmellina</emphasis>
<bibRefCitation id="EFB24DD5EE75ED58FC40CA7CFB80FBAB" author="Quatrefages A de" box="[937,1143,1025,1047]" pageId="30" pageNumber="533" pagination="173 - 303" refId="ref40339" refString="Quatrefages A de. 1846. Etudes sur les types inferieurs de l'embranchement des anneles; memoire sur la famille des nemertiens (Nemertea). Annales des Sciences Naturelles, Troisieme Serie 6: 173 - 303, pls 8 - 14." type="journal article" year="1846">Quatrefages, 1846</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54988E86EE75ED58FB6CCA7CFB40FBAA" box="[1157,1207,1025,1046]" pageId="30" pageNumber="533">type</typeStatus>
locality, Favignana Island,
<collectingCountry id="F33470B4EE75ED58FC6DCA5DFC4AFB8A" box="[900,957,1056,1078]" name="Italy" pageId="30" pageNumber="533">Italy</collectingCountry>
) with
<taxonomicName id="4C234BA7EE75ED58FBE4CA5DFB15FB89" authorityName=": Burger" authorityYear="1893" box="[1037,1250,1056,1077]" class="Pilidiophora" family="Valenciniidae" genus="Eupolia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="delineata">
<emphasis id="B957EC36EE75ED58FBE4CA5DFB15FB89" box="[1037,1250,1056,1077]" italics="true" pageId="30" pageNumber="533">Eupolia delineata</emphasis>
</taxonomicName>
, but there are some differences between the descriptions of the two: in
<taxonomicName id="4C234BA7EE75ED58FCAECA20FBDEFBCE" authorityName="Quatrefages" authorityYear="1846" box="[839,1065,1117,1138]" class="Pilidiophora" family="Lineidae" genus="Borlasia" kingdom="Animalia" order="Heteronemertea" pageId="30" pageNumber="533" phylum="Nemertea" rank="species" species="carmellina">
<emphasis id="B957EC36EE75ED58FCAECA20FBDEFBCE" box="[839,1065,1117,1138]" italics="true" pageId="30" pageNumber="533">Borlasia carmellina</emphasis>
</taxonomicName>
, the body is reddish posteriorly and covered with small, elongate patches (petites taches allongée) rather than stripes, and eyes are lacking (
<bibRefCitation id="EFB24DD5EE75ED58FC62CAC4FBADFB73" author="Quatrefages A de" box="[907,1114,1209,1231]" pageId="30" pageNumber="533" pagination="173 - 303" refId="ref40339" refString="Quatrefages A de. 1846. Etudes sur les types inferieurs de l'embranchement des anneles; memoire sur la famille des nemertiens (Nemertea). Annales des Sciences Naturelles, Troisieme Serie 6: 173 - 303, pls 8 - 14." type="journal article" year="1846">Quatrefages, 1846</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>