416 lines
62 KiB
XML
416 lines
62 KiB
XML
<document id="2EA2895C6ABC1756BAFDB00EB7E3D9F3" ID-CLB-Dataset="267614" ID-DOI="10.1163/18759866-BJA10027" ID-GBIF-Dataset="2fd2ff98-fcd5-484e-a7b8-6a60f12e7084" ID-ISSN="1875-9866" ID-Zenodo-Dep="8348969" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1694665046491" checkinUser="felipe" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung & Panha, Somsak" docDate="2022" docId="038587FD036AFFCC745BFEC695C4FA46" docLanguage="en" docName="ContribZool.91.2.97-132.pdf" docOrigin="Contributions to Zoology 91 (2)" docSource="http://dx.doi.org/10.1163/18759866-bja10027" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Sarika costabilis Pholyotha & Sutcharit & Lin & Panha 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="126" masterDocId="FFBCFF850373FFD17439FF98972DFFE8" masterDocTitle="Multigene phylogeny reveals the ribbed shell morphotypes in the land snail genus Sarika (Eupulmonata: Ariophantidae), with description of two new species from Thailand and Myanmar" masterLastPageNumber="132" masterPageNumber="97" pageNumber="122" updateTime="1694783892711" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="32D8544C8EE8CC68B4893E00F708A94D">Multigene phylogeny reveals the ribbed shell morphotypes in the land snail genus Sarika (Eupulmonata: Ariophantidae), with description of two new species from Thailand and Myanmar</mods:title>
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<mods:roleTerm id="1E0C71B9568EB18D55F7549FF9317156">Author</mods:roleTerm>
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<mods:namePart id="CCDB89B5A93DFAD1D57CC254D64D8E9E">Pholyotha, Arthit</mods:namePart>
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<mods:namePart id="C60DB2CF91AA67AB2C99D00919FEE88E">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="59F89A7617411304891F1E79638B3350">Lin, Aung</mods:namePart>
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<mods:namePart id="1696FC06C4E24E79602CE8F696EE229D">Panha, Somsak</mods:namePart>
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<mods:title id="3FB5E01E786E097FC43E1D1F72FB2896">Contributions to Zoology</mods:title>
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<mods:part id="9A183BC92BF25EDCC5581512079E4465">
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<mods:date id="D6165097DE750B04F6229FE53BDC5215">2022</mods:date>
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<mods:number id="8B59DE527AFD0AB045B8BE3FBAB1B9B3">2022-03-21</mods:number>
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<treatment id="038587FD036AFFCC745BFEC695C4FA46" ID-DOI="http://doi.org/10.5281/zenodo.8348981" ID-Zenodo-Dep="8348981" LSID="urn:lsid:plazi:treatment:038587FD036AFFCC745BFEC695C4FA46" httpUri="http://treatment.plazi.org/id/038587FD036AFFCC745BFEC695C4FA46" lastPageId="29" lastPageNumber="126" pageId="25" pageNumber="122">
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<subSubSection id="C3366560036AFFC8745BFEC696D4FE91" box="[98,505,350,377]" pageId="25" pageNumber="122" type="nomenclature">
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<heading id="D0DB8187036AFFC8745BFEC696D4FE91" box="[98,505,350,377]" fontSize="11" level="2" pageId="25" pageNumber="122" reason="2">
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<emphasis id="B958EAF9036AFFC8745BFEC696D4FE91" bold="true" box="[98,505,350,377]" pageId="25" pageNumber="122">
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<taxonomicName id="4C2C4D68036AFFC8745BFEC696B7FE91" ID-CoL="8YNYM" authority="Pholyotha" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[98,410,350,377]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="122" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
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<emphasis id="B958EAF9036AFFC8745BFEC69630FE91" bold="true" box="[98,285,350,377]" italics="true" pageId="25" pageNumber="122">Sarika costabilis</emphasis>
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Pholyotha
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,
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<taxonomicNameLabel id="A26B5782036AFFC8759CFEC796D4FE91" box="[421,505,351,377]" pageId="25" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3366560036AFFCC745BFE1B95C4FA46" lastPageId="29" lastPageNumber="126" pageId="25" pageNumber="122" type="description">
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<paragraph id="8B9336EB036AFFC8745BFE1B96B3FE75" blockId="25.[98,610,387,557]" box="[98,414,387,413]" pageId="25" pageNumber="122">
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(
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<figureCitation id="13172A6E036AFFC87450FE1B978FFE75" box="[105,162,387,413]" captionStart="FIGURE 1" captionStartId="3.[99,166,977,998]" captionTargetBox="[193,1062,164,958]" captionTargetId="figure-233@3.[193,1062,164,958]" captionTargetPageId="3" captionText="FIGURE 1 Geographic distribution of ribbed shell Sarika spp. based on the specimens examined herein. A) Distribution of S. theodori, S.costabilis sp.nov., and S. costata sp. nov. B) Distribution of S. siamensis. Black symbols indicate the DNA extracted specimens used in the molecular analysis; white symbols indicate shells or other preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.8348977" httpUri="https://zenodo.org/record/8348977/files/figure.png" pageId="25" pageNumber="122">figs 1</figureCitation>
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,
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<figureCitation id="13172A6E036AFFC87495FE1B9797FE75" box="[172,186,387,413]" captionStart="FIGURE 2" captionStartId="11.[100,167,1258,1279]" captionTargetBox="[193,1061,164,1238]" captionTargetId="figure-122@11.[193,1062,164,1239]" captionTargetPageId="11" captionText="FIGURE 2 Bayesian inference tree based on a 1495 bp concatenated alignment dataset of COI + 16S + 28S genes. Numbers by the nodes are the Bayesian posterior probabilities (left) and ML bootstrap values (right); shown only for the nodes supported by BI ≥ 0.95 or ML ≥ 70%. Clade A consists of the S. resplendens-S. hainesi species groups.Clade B consists of the S.siamensis species group.Images of living snails are not to scale." figureDoi="http://doi.org/10.5281/zenodo.8348979" httpUri="https://zenodo.org/record/8348979/files/figure.png" pageId="25" pageNumber="122">2</figureCitation>
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,
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<figureCitation id="13172A6E036AFFC874FDFE1B962DFE75" box="[196,256,387,413]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="25" pageNumber="122">8A, B</figureCitation>
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,
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<figureCitation id="13172A6E036AFFC87533FE1B966BFE75" box="[266,326,387,413]" captionStart="FIGURE 9" captionStartId="27.[98,165,1350,1371]" captionTargetBox="[193,1062,164,1331]" captionTargetId="figure-97@27.[193,1062,164,1331]" captionTargetPageId="27" captionText="FIGURE 9 Genitalia of Sarika spp.A, B) S. costabilis sp. nov.paratype CUMZ 7945.A) General view of the genital system.B) Internal structure of penis.C, D) S.costata sp. nov. paratype CUMZ 7947.C) General view of the genital system.D) Internal structure of penis. White arrows indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.8349001" httpUri="https://zenodo.org/record/8349001/files/figure.png" pageId="25" pageNumber="122">9A, B</figureCitation>
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,
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||
<figureCitation id="13172A6E036AFFC87569FE1B9644FE75" box="[336,361,387,413]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="25" pageNumber="122">10</figureCitation>
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,
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<figureCitation id="13172A6E036AFFC8754AFE1B96BAFE75" box="[371,407,387,413]" captionStart="FIGURE 11" captionStartId="29.[98,165,1042,1063]" captionTargetBox="[193,1062,164,1023]" captionTargetId="figure-192@29.[193,1062,164,1023]" captionTargetPageId="29" captionText="FIGURE 11 Radula of Sarika spp. A) S.theodori specimen CUMZ 7942.B) S.siamensis specimen CUMZ 7951.C) S.costabilis sp. nov. paratype CUMZ 7945. D) S. costata sp. nov. paratype CUMZ 7947.Central tooth indicated by ‘C’ and blue colour; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.8349005" httpUri="https://zenodo.org/record/8349005/files/figure.png" pageId="25" pageNumber="122">11C</figureCitation>
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)
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</paragraph>
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<paragraph id="8B9336EB036AFFC8745BFE30954CFDC5" blockId="25.[98,610,387,557]" pageId="25" pageNumber="122">
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<emphasis id="B958EAF9036AFFC8745BFE3096A4FE29" box="[98,393,423,449]" italics="true" pageId="25" pageNumber="122">Type material examined.</emphasis>
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<materialsCitation id="3B443CB6036AFFC875A5FE3F9573FE0D" collectionCode="CUMZ" pageId="25" pageNumber="122" specimenCode="CUMZ 7943" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54978849036AFFC875A5FE3F952AFE29" box="[412,519,423,449]" pageId="25" pageNumber="122">Holotype</typeStatus>
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.
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<specimenCode id="DB8A9E90036AFFC87624FE3F97B8FE0D" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="25" pageNumber="122">CUMZ 7943</specimenCode>
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(
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<figureCitation id="13172A6E036AFFC8749CFE5397C6FE0D" box="[165,235,459,485]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="25" pageNumber="122">fig. 8A</figureCitation>
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; width
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, height
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<quantity id="4CD49B0E036AFFC875CEFE53957BFE0D" box="[503,598,459,485]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="25" pageNumber="122" unit="mm" value="16.7">16.7 mm</quantity>
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)
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.
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<materialsCitation id="3B443CB6036AFFC8745BFE779570FDC5" collectionCode="CUMZ" pageId="25" pageNumber="122" specimenCode="CUMZ 7944" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54978849036AFFC8745BFE7797FCFDE1" box="[98,209,495,521]" pageId="25" pageNumber="122">Paratypes</typeStatus>
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. Same locality as holotype:
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<specimenCode id="DB8A9E90036AFFC87624FE7797BBFDC5" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="25" pageNumber="122">CUMZ 7944</specimenCode>
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(
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<figureCitation id="13172A6E036AFFC8749CFD8A97C0FDC5" box="[165,237,530,557]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="25" pageNumber="122">fig. 8B</figureCitation>
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, width
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, height
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<quantity id="4CD49B0E036AFFC875C2FD8A9578FDC5" box="[507,597,530,557]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="25" pageNumber="122" unit="mm" value="16.1">16.1 mm</quantity>
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)
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,
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<paragraph id="8B9336EB036AFFC876BCFF349426FEBE" blockId="25.[645,1157,171,557]" pageId="25" pageNumber="122">
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NHMUK 0000 (two shells), and
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<materialsCitation id="3B443CB6036AFFC87036FF3493ACFED9" collectionCode="ZRC" country="Thailand" county="Thong Pha Phum District" latitude="14.941973" location="Limestone" longLatPrecision="1" longitude="98.669556" municipality="Cha Lae" pageId="25" pageNumber="122" specimenCount="1" stateProvince="Kanchanaburi">
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<collectionCode id="ED3DAE2E036AFFC87036FF349316FF2E" box="[1039,1083,172,198]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="25" pageNumber="122">ZRC</collectionCode>
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0000 (two shells).
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<location id="8EF36030036AFFC87725FF5794BFFF01" LSID="urn:lsid:plazi:treatment:038587FD036AFFCC745BFEC695C4FA46:8EF36030036AFFC87725FF5794BFFF01" box="[796,914,207,233]" country="Thailand" county="Thong Pha Phum District" latitude="14.941973" longLatPrecision="1" longitude="98.669556" municipality="Cha Lae" name="Limestone" pageId="25" pageNumber="122" stateProvince="Kanchanaburi">Limestone</location>
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outcrop in
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<collectingMunicipality id="6BF7AC91036AFFC87019FF5793ACFF02" box="[1056,1153,207,234]" pageId="25" pageNumber="122">Cha Lae</collectingMunicipality>
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,
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<collectingCounty id="62F24E67036AFFC876BCFF6B94E0FEE5" box="[645,973,243,269]" pageId="25" pageNumber="122">Thong Pha Phum District</collectingCounty>
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,
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<collectingRegion id="49E8F809036AFFC877DDFF6B95C4FED9" country="Thailand" name="Kanchanaburi" pageId="25" pageNumber="122">Kanchanaburi Province</collectingRegion>
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,
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<collectingCountry id="F33B767B036AFFC876CEFE8F9470FED9" box="[759,861,279,305]" name="Thailand" pageId="25" pageNumber="122">Thailand</collectingCountry>
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,
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,
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<materialsCitation id="3B443CB6036AFFC876BCFEA3942AFEBE" box="[645,775,315,342]" collectionCode="CUMZ" country="Thailand" pageId="25" pageNumber="122" specimenCode="CUMZ 7945" specimenCount="1" stateProvince="Kanchanaburi">
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<specimenCode id="DB8A9E90036AFFC876BCFEA3942AFEBE" box="[645,775,315,342]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="25" pageNumber="122">CUMZ 7945</specimenCode>
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</materialsCitation>
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.
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</paragraph>
|
||
<paragraph id="8B9336EB036AFFC8769DFEF89355FE2A" blockId="25.[645,1157,171,557]" pageId="25" pageNumber="122">
|
||
<materialsCitation id="3B443CB6036AFFC8769DFEF89359FE2A" country="Thailand" county="Sangkhla Buri District" latitude="15.036861" location="Wat Pa Tham Sukho Prang" longLatPrecision="1" longitude="98.58283" municipality="Pa Tham Sukho Prang Phe" pageId="25" pageNumber="122" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
|
||
<emphasis id="B958EAF9036AFFC8769DFEF89415FE91" box="[676,824,351,377]" italics="true" pageId="25" pageNumber="122">
|
||
<typeStatus id="54978849036AFFC8769DFEF895F5FE91" box="[676,728,352,377]" pageId="25" pageNumber="122">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EF36030036AFFC8777BFEC793A8FE91" LSID="urn:lsid:plazi:treatment:038587FD036AFFCC745BFEC695C4FA46:8EF36030036AFFC8777BFEC793A8FE91" box="[834,1157,351,377]" country="Thailand" county="Sangkhla Buri District" latitude="15.036861" longLatPrecision="1" longitude="98.58283" municipality="Pa Tham Sukho Prang Phe" name="Wat Pa Tham Sukho Prang" pageId="25" pageNumber="122" stateProvince="Kanchanaburi">
|
||
Wat
|
||
<collectingMunicipality id="6BF7AC91036AFFC87745FEC793A8FE91" box="[892,1157,351,377]" pageId="25" pageNumber="122">Pa Tham Sukho Prang</collectingMunicipality>
|
||
</location>
|
||
Phe,
|
||
<collectingCounty id="62F24E67036AFFC876FFFE1B94FCFE75" box="[710,977,387,413]" pageId="25" pageNumber="122">Sangkhla Buri District</collectingCounty>
|
||
,
|
||
<collectingRegion id="49E8F809036AFFC877DDFE1B95C4FE29" country="Thailand" name="Kanchanaburi" pageId="25" pageNumber="122">Kanchanaburi Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33B767B036AFFC876CAFE3F947AFE29" box="[755,855,423,449]" name="Thailand" pageId="25" pageNumber="122">Thailand</collectingCountry>
|
||
<geoCoordinate id="EE18502C036AFFC87764FE3F94CCFE2A" box="[861,993,423,450]" degrees="15" direction="north" minutes="02" orientation="latitude" pageId="25" pageNumber="122" precision="1" seconds="12.7" value="15.036861">15°02’12.7”N</geoCoordinate>
|
||
<geoCoordinate id="EE18502C036AFFC877DEFE3F9359FE2A" box="[999,1140,423,450]" degrees="98" direction="east" minutes="34" orientation="longitude" pageId="25" pageNumber="122" precision="1" seconds="58.2" value="98.58283">98°34’58.2”E</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036AFFCB769DFE539639FF02" blockId="25.[645,1157,171,557]" lastBlockId="26.[129,640,171,1742]" lastPageId="26" lastPageNumber="123" pageId="25" pageNumber="122">
|
||
<emphasis id="B958EAF9036AFFC8769DFE539431FE0D" box="[676,796,459,485]" italics="true" pageId="25" pageNumber="122">Etymology.</emphasis>
|
||
The specific name “
|
||
<taxonomicName id="4C2C4D68036AFFC877C7FE539349FE0D" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[1022,1124,459,485]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="122" phylum="Mollusca" rank="species" species="costabilis">
|
||
<emphasis id="B958EAF9036AFFC877C7FE539349FE0D" box="[1022,1124,459,485]" italics="true" pageId="25" pageNumber="122">costabilis</emphasis>
|
||
</taxonomicName>
|
||
” is from the Latin word meaning “riblike”, which refers to shell surface with transversely dashed ribs that characterises this species within the genus
|
||
<taxonomicName id="4C2C4D680369FFCB74F1FF579623FF01" authorityName="Godwin-Austen" authorityYear="1907" box="[200,270,207,233]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="123" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B958EAF90369FFCB74F1FF579623FF01" box="[200,270,207,233]" italics="true" pageId="26" pageNumber="123">Sarika</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF536663036AFFC87458F9039403F926" ID-DOI="http://doi.org/10.5281/zenodo.8348999" ID-Zenodo-Dep="8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="25" pageNumber="122" startId="25.[97,164,1691,1712]" targetBox="[193,1062,591,1673]" targetPageId="25" targetType="figure">
|
||
<paragraph id="8B9336EB036AFFC87458F9039403F926" blockId="25.[97,1157,1691,1743]" pageId="25" pageNumber="122">
|
||
FIGURE 8 Shells of
|
||
<taxonomicName id="4C2C4D68036AFFC87514F9039688F958" box="[301,421,1691,1713]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="122" phylum="Mollusca" rank="species" species="undefined-A">
|
||
<emphasis id="B958EAF9036AFFC87514F903964BF958" box="[301,358,1691,1712]" italics="true" pageId="25" pageNumber="122">Sarika</emphasis>
|
||
spp. A
|
||
</taxonomicName>
|
||
, B)
|
||
<taxonomicName id="4C2C4D68036AFFC875FEF9049503F958" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[455,558,1691,1713]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="122" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036AFFC875FEF9049503F958" box="[455,558,1691,1713]" italics="true" pageId="25" pageNumber="122">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036AFFC8760AF904955AF959" box="[563,631,1692,1713]" pageId="25" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A) Holotype CUMZ 7943. B) Paratype CUMZ 7944. C, D)
|
||
<taxonomicName id="4C2C4D68036AFFC8704DF9049631F926" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="122" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036AFFC8704DF9049631F926" italics="true" pageId="25" pageNumber="122">S. costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036AFFC87518F9219648F926" box="[289,357,1721,1742]" pageId="25" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
|
||
C) Holotype CUMZ 7946. D) Paratype CUMZ 7947.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9336EB0369FFCB74A6FF6C9502FDB9" blockId="26.[129,640,171,1742]" pageId="26" pageNumber="123">
|
||
<emphasis id="B958EAF90369FFCB74A6FF6C96FDFEE5" box="[159,464,243,269]" italics="true" pageId="26" pageNumber="123">Diagnostic characteristics.</emphasis>
|
||
Shell large, depressed; body whorl obtusely angulated; shell ribbed with nodules arranged on radial ribs. Animal dark creamy-grey body and four mantle edges. Genitalia with short and straight epiphallic caecum and cuboidal penial pilasters. Spermatophore: head filament with smooth longitudinal folds; tail filament bearing three spines near sperm sac and with branching spines on middle and terminal part.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0369FFCB74A6FDC3966BFB3F" blockId="26.[129,640,171,1742]" pageId="26" pageNumber="123">
|
||
<emphasis id="B958EAF90369FFCB74A6FDC39674FD9C" box="[159,345,602,628]" italics="true" pageId="26" pageNumber="123">Description.Shell</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E0369FFCB755BFDC296EFFD9D" box="[354,450,602,629]" captionStart="FIGURE 8" captionStartId="25.[97,164,1691,1712]" captionTargetBox="[193,1062,591,1673]" captionTargetId="figure-133@25.[193,1062,591,1673]" captionTargetPageId="25" captionText="FIGURE 8 Shells of Sarika spp.A, B) S. costabilis sp. nov.A) Holotype CUMZ 7943.B) Paratype CUMZ 7944.C, D) S. costata sp.nov.C) Holotype CUMZ 7946.D) Paratype CUMZ 7947." figureDoi="http://doi.org/10.5281/zenodo.8348999" httpUri="https://zenodo.org/record/8348999/files/figure.png" pageId="26" pageNumber="123">fig.8A,B</figureCitation>
|
||
).Shelldepressed, large size (width up to
|
||
<quantity id="4CD49B0E0369FFCB75B1FDE696C5FD71" box="[392,488,638,665]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="26" pageNumber="123" unit="mm" value="29.3">29.3 mm</quantity>
|
||
, height up to
|
||
<quantity id="4CD49B0E0369FFCB74B8FD3A97F3FD55" box="[129,222,674,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="26" pageNumber="123" unit="mm" value="16.7">16.7 mm</quantity>
|
||
), and rather solid; shell colour yellowish-brown. Whorls 6–6½, increasing regularly; near aperture opening slightly sloping downwards. Spire dome-shaped; apex slightly protruding; suture impressed. Embryonic shell nearly smooth with very weak radial wrinkles. Body whorl large and obtusely angulated. Upper shell surface sculptured with distinct radial ribs, crossed by spiral lines, and with series of nodules arranged on radial ribs, and then radial ribs gradually disappearing below periphery towards umbilicus. Aperture crescent-shaped, little descending, and lip little thickened. Columellar margin simple and slightly reflected near umbilicus. Umbilicus narrowly opened.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0369FFCB74A6FB789486FF01" blockId="26.[129,640,171,1742]" lastBlockId="26.[676,1188,171,1742]" pageId="26" pageNumber="123">
|
||
<emphasis id="B958EAF90369FFCB74A6FB789665FB12" box="[159,328,1248,1274]" italics="true" pageId="26" pageNumber="123">Genital organs</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E0369FFCB755BFB7896F7FB13" box="[354,474,1248,1275]" captionStart="FIGURE 9" captionStartId="27.[98,165,1350,1371]" captionTargetBox="[193,1062,164,1331]" captionTargetId="figure-97@27.[193,1062,164,1331]" captionTargetPageId="27" captionText="FIGURE 9 Genitalia of Sarika spp.A, B) S. costabilis sp. nov.paratype CUMZ 7945.A) General view of the genital system.B) Internal structure of penis.C, D) S.costata sp. nov. paratype CUMZ 7947.C) General view of the genital system.D) Internal structure of penis. White arrows indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.8349001" httpUri="https://zenodo.org/record/8349001/files/figure.png" pageId="26" pageNumber="123">fig. 9A, B</figureCitation>
|
||
). Atrium (at) enlarged and very short. Penis (p) elongate, cylindrical, with thin penial sheath covering near atrium. Inner surface sculptured, very finely folded to nearly smooth from atrium to one-third of penis, and then gradually transformed to cuboidal penial pilasters (pp). Epiphallus(e) cylindrical, slightly shorter than penis. Epiphallic caecum (ec) short, straight, diameter nearly equal to proximal epiphallus, and located at middle of epiphallus. Penial retractor muscle (prm) thin and attached at tip of epiphallic caecum. Flagellum (fl) very long approximately twice of epiphallus length, slender and enlarged at the tip. Vas deferens (vd) thin tube.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0369FFCB76FBFF6B935BFDB8" blockId="26.[676,1188,171,1742]" pageId="26" pageNumber="123">Vagina (v) cylindrical and slightly shorter than half of penis length. Dart apparatus (da) enlarged cylindrical and joined to atrium at vagina and penis junction. Gametolytic sac (gs) elongate bulbous (containing one spermatophore inside); gametolytic duct (gd) cylindrical and enlarged at based. Free oviduct (fo) enlarged cylindrical, approximately two times of vagina length. Oviduct large lobules; prostate gland running alongside oviduct.</paragraph>
|
||
<paragraph id="8B9336EB0369FFCB76FBFDC29442FB5B" blockId="26.[676,1188,171,1742]" pageId="26" pageNumber="123">
|
||
<emphasis id="B958EAF90369FFCB76FBFDC29446FD9C" box="[706,875,602,628]" italics="true" pageId="26" pageNumber="123">Spermatophore</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E0369FFCB77BCFDC294FBFD9C" box="[901,982,602,628]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10</figureCitation>
|
||
). Spermatophore long and needle-shaped (
|
||
<figureCitation id="13172A6E0369FFCB77FAFDE69331FD70" box="[963,1052,638,664]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10A</figureCitation>
|
||
). Sperm sac (ss) enlarged and elongate oval. Head filament (hf) elongate with smooth longitudinal folds (
|
||
<figureCitation id="13172A6E0369FFCB7692FD729428FCEC" box="[683,773,746,772]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10B</figureCitation>
|
||
). Tail filament (tf), region close to sperm sac bearing three spines: spine I very small and simple; spine II longer than spine I with bifurcating into spinules; spine III longest with complicated branching into small spinules (
|
||
<figureCitation id="13172A6E0369FFCB7729FC05944AFC5F" box="[784,871,925,951]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10C</figureCitation>
|
||
). Region furthest away with series of ten branching spines arranged in row located in middle (red dashed line in
|
||
<figureCitation id="13172A6E0369FFCB7068FC7D93B2FC17" box="[1105,1183,997,1023]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig 10A</figureCitation>
|
||
;
|
||
<figureCitation id="13172A6E0369FFCB769DFB9195D7FBCB" box="[676,762,1033,1059]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10D</figureCitation>
|
||
); terminal part about one-tenth of its length, composed of series of short to long delicate branching spines arranged in multiple rows, and covering terminal part of tail filament (
|
||
<figureCitation id="13172A6E0369FFCB7729FB01944EFB5B" box="[784,867,1177,1203]" captionStart="FIGURE 10" captionStartId="28.[129,196,1324,1345]" captionTargetBox="[223,1092,164,1305]" captionTargetId="figure-100@28.[223,1092,164,1305]" captionTargetPageId="28" captionText="FIGURE 10 Spermatophore of Sarika costabilis sp.nov.paratype CUMZ 7945.A) General view of spermatophore that was broken into two parts.B) Head filament.C–E) Tail filament.C) Three spines located close to the sperm sac.D) Region with and without branching spines.E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length." figureDoi="http://doi.org/10.5281/zenodo.8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="26" pageNumber="123">fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0369FFCB76FBFB24947FF9D5" blockId="26.[676,1188,171,1742]" pageId="26" pageNumber="123">
|
||
<taxonomicName id="4C2C4D680369FFCB76FBFB24943CFB3E" box="[706,785,1212,1238]" class="Bivalvia" family="Limidae" genus="Radula" kingdom="Animalia" order="Limida" pageId="26" pageNumber="123" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B958EAF90369FFCB76FBFB24943CFB3E" box="[706,785,1212,1238]" italics="true" pageId="26" pageNumber="123">Radula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13172A6E0369FFCB771AFB249455FB3F" box="[803,888,1212,1239]" captionStart="FIGURE 11" captionStartId="29.[98,165,1042,1063]" captionTargetBox="[193,1062,164,1023]" captionTargetId="figure-192@29.[193,1062,164,1023]" captionTargetPageId="29" captionText="FIGURE 11 Radula of Sarika spp. A) S.theodori specimen CUMZ 7942.B) S.siamensis specimen CUMZ 7951.C) S.costabilis sp. nov. paratype CUMZ 7945. D) S. costata sp. nov. paratype CUMZ 7947.Central tooth indicated by ‘C’ and blue colour; yellow colour indicates lateral teeth in the transition to marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.8349005" httpUri="https://zenodo.org/record/8349005/files/figure.png" pageId="26" pageNumber="123">fig. 11C</figureCitation>
|
||
). Teeth arranged in wide, straight, and with half row formula: 1–(19– 20)–70. Central tooth symmetrical tricuspid with large mesocone and small ectocones. Lateral teeth asymmetrical tricuspid with large mesocone, and small endocone and ectocone. Marginal teeth started around tooth number 19 or 20; inner teeth elongate bicuspid with lanceolate endocone and minute ectocone; and outermost teeth gradually reduced in size.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0369FFCA76FBF9DF9657F96E" blockId="26.[676,1188,171,1742]" lastBlockId="27.[98,610,1500,1742]" lastPageId="27" lastPageNumber="124" pageId="26" pageNumber="123">
|
||
<emphasis id="B958EAF90369FFCB76FBF9DF94A7F98A" box="[706,906,1607,1634]" italics="true" pageId="26" pageNumber="123">External features</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E0369FFCB779DF9DF94C7F98A" box="[932,1002,1607,1634]" captionStart="FIGURE 2" captionStartId="11.[100,167,1258,1279]" captionTargetBox="[193,1061,164,1238]" captionTargetId="figure-122@11.[193,1062,164,1239]" captionTargetPageId="11" captionText="FIGURE 2 Bayesian inference tree based on a 1495 bp concatenated alignment dataset of COI + 16S + 28S genes. Numbers by the nodes are the Bayesian posterior probabilities (left) and ML bootstrap values (right); shown only for the nodes supported by BI ≥ 0.95 or ML ≥ 70%. Clade A consists of the S. resplendens-S. hainesi species groups.Clade B consists of the S.siamensis species group.Images of living snails are not to scale." figureDoi="http://doi.org/10.5281/zenodo.8348979" httpUri="https://zenodo.org/record/8348979/files/figure.png" pageId="26" pageNumber="123">fig. 2</figureCitation>
|
||
). Living snails with reticulated skin, dark grey to black eye stalks, and pale to dark creamy-grey body. Four well-developed mantle lobes with same colour as body. Right shell lobe short and left shell lobe absent. Dorsal lobes large and broad; anterior and posterior left dorsal lobes smaller than right dorsal lobe. Caudal fossa and caudal horn present.
|
||
</paragraph>
|
||
<caption id="DF5366630368FFCA745BFADE9325FA7F" ID-DOI="http://doi.org/10.5281/zenodo.8349001" ID-Zenodo-Dep="8349001" httpUri="https://zenodo.org/record/8349001/files/figure.png" pageId="27" pageNumber="124" startId="27.[98,165,1350,1371]" targetBox="[193,1062,164,1331]" targetPageId="27" targetType="figure">
|
||
<paragraph id="8B9336EB0368FFCA745BFADE9325FA7F" blockId="27.[98,1149,1350,1431]" pageId="27" pageNumber="124">
|
||
FIGURE 9 Genitalia of
|
||
<taxonomicName id="4C2C4D680368FFCA7575FADE96E9FAB3" box="[332,452,1350,1372]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="124" phylum="Mollusca" rank="species" species="undefined-A">
|
||
<emphasis id="B958EAF90368FFCA7575FADE96A8FAB3" box="[332,389,1350,1371]" italics="true" pageId="27" pageNumber="124">Sarika</emphasis>
|
||
spp. A
|
||
</taxonomicName>
|
||
, B)
|
||
<taxonomicName id="4C2C4D680368FFCA75DFFADF9560FAB3" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[486,589,1350,1372]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="124" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF90368FFCA75DFFADF9560FAB3" box="[486,589,1350,1372]" italics="true" pageId="27" pageNumber="124">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57820368FFCA7668FADF95BBFAB4" box="[593,662,1351,1372]" pageId="27" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype CUMZ 7945. A) General view of the genital system. B) Internal structure of penis. C, D)
|
||
<taxonomicName id="4C2C4D680368FFCA7652FAFC9592FA91" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[619,703,1380,1401]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="124" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF90368FFCA7652FAFC9592FA91" box="[619,703,1380,1401]" italics="true" pageId="27" pageNumber="124">S. costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57820368FFCA76FDFAFC9425FA91" box="[708,776,1380,1401]" pageId="27" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype CUMZ 7947. C) General view of the genital system. D) Internal structure of penis. White arrows indicate the end of penis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9336EB0368FFCA74B8F917947EF96D" blockId="27.[98,610,1500,1742]" lastBlockId="27.[645,1157,1500,1741]" pageId="27" pageNumber="124">
|
||
<emphasis id="B958EAF90368FFCA74B8F9179628F941" box="[129,261,1679,1705]" italics="true" pageId="27" pageNumber="124">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="13172A6E0368FFCA752BF9179668F942" box="[274,325,1679,1706]" captionStart="FIGURE 1" captionStartId="3.[99,166,977,998]" captionTargetBox="[193,1062,164,958]" captionTargetId="figure-233@3.[193,1062,164,958]" captionTargetPageId="3" captionText="FIGURE 1 Geographic distribution of ribbed shell Sarika spp. based on the specimens examined herein. A) Distribution of S. theodori, S.costabilis sp.nov., and S. costata sp. nov. B) Distribution of S. siamensis. Black symbols indicate the DNA extracted specimens used in the molecular analysis; white symbols indicate shells or other preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.8348977" httpUri="https://zenodo.org/record/8348977/files/figure.png" pageId="27" pageNumber="124">fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="4C2C4D680368FFCA7560F9179527F941" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[345,522,1679,1705]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="124" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF90368FFCA7560F9179527F941" box="[345,522,1679,1705]" italics="true" pageId="27" pageNumber="124">Sarika costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57820368FFCA7629F908954FF942" box="[528,610,1680,1706]" pageId="27" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is endemic to
|
||
<collectingCountry id="F33B767B0368FFCA753BF92B964BF925" box="[258,358,1715,1741]" name="Thailand" pageId="27" pageNumber="124">Thailand</collectingCountry>
|
||
and, currently known from a few limestone localities in mountainous areas in
|
||
<collectingRegion id="49E8F8090368FFCA7727F9989301F9F2" box="[798,1068,1536,1562]" country="Thailand" name="Kanchanaburi" pageId="27" pageNumber="124">Kanchanaburi Province</collectingRegion>
|
||
(
|
||
<figureCitation id="13172A6E0368FFCA7007F998935BF9F2" box="[1086,1142,1536,1562]" captionStart="FIGURE 1" captionStartId="3.[99,166,977,998]" captionTargetBox="[193,1062,164,958]" captionTargetId="figure-233@3.[193,1062,164,958]" captionTargetPageId="3" captionText="FIGURE 1 Geographic distribution of ribbed shell Sarika spp. based on the specimens examined herein. A) Distribution of S. theodori, S.costabilis sp.nov., and S. costata sp. nov. B) Distribution of S. siamensis. Black symbols indicate the DNA extracted specimens used in the molecular analysis; white symbols indicate shells or other preserved specimens." figureDoi="http://doi.org/10.5281/zenodo.8348977" httpUri="https://zenodo.org/record/8348977/files/figure.png" pageId="27" pageNumber="124">fig. 1</figureCitation>
|
||
). We extended our survey in other karstic habitats in
|
||
<collectingCountry id="F33B767B0368FFCA76E6F9D09468F98A" box="[735,837,1608,1634]" name="Thailand" pageId="27" pageNumber="124">Thailand</collectingCountry>
|
||
, but we could not find this species elsewhere.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB0368FFCC769DF91797D2FB13" blockId="27.[645,1157,1500,1741]" lastBlockId="29.[98,609,1213,1742]" lastPageId="29" lastPageNumber="126" pageId="27" pageNumber="124">
|
||
<emphasis id="B958EAF90368FFCA769DF9179427F941" box="[676,778,1679,1705]" italics="true" pageId="27" pageNumber="124">Remarks.</emphasis>
|
||
Among the
|
||
<taxonomicName id="4C2C4D680368FFCA77AEF917931BF941" box="[919,1078,1679,1705]" pageId="27" pageNumber="124">
|
||
<emphasis id="B958EAF90368FFCA77AEF91794F0F941" box="[919,989,1679,1705]" italics="true" pageId="27" pageNumber="124">Sarika</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with a ribbed shell, the distinguishing character of
|
||
<taxonomicName id="4C2C4D68036FFFCD74B8FA44962CFA1E" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[129,257,1500,1526]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="125" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036FFFCD74B8FA44962CFA1E" box="[129,257,1500,1526]" italics="true" pageId="28" pageNumber="125">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036FFFCD7531FA459671FA1F" box="[264,348,1501,1527]" pageId="28" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the obtusely angulated last whorl with descending aperture, genitalia with short and large epiphallic caecum and long flagellum, and spermatophore with branching spines at the middle and terminal part of tail filament. In contrast,
|
||
<taxonomicName id="4C2C4D68036FFFCD75C3F90895ADF941" box="[506,640,1680,1706]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="125" phylum="Mollusca" rank="species" species="siamensis">
|
||
<emphasis id="B958EAF9036FFFCD75C3F90895ADF941" box="[506,640,1680,1706]" italics="true" pageId="28" pageNumber="125">S. siamensis</emphasis>
|
||
</taxonomicName>
|
||
has a well-rounded last whorl, aperture not descending, genitalia with long epiphallic caecum and short flagellum, and spermatophore having spines nearly the entire tail filament length. Shell morphology of
|
||
<taxonomicName id="4C2C4D68036FFFCD701DF9D0938EF98A" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[1060,1187,1608,1634]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="125" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036FFFCD701DF9D0938EF98A" box="[1060,1187,1608,1634]" italics="true" pageId="28" pageNumber="125">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036FFFCD769DF9F495D1F96E" box="[676,764,1644,1670]" pageId="28" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is somewhat similar to
|
||
<taxonomicName id="4C2C4D68036FFFCD701AF9F493B0F96D" authorityName=": Godwin-Austen" authorityYear="1907" box="[1059,1181,1643,1670]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="125" phylum="Mollusca" rank="species" species="theodori">
|
||
<emphasis id="B958EAF9036FFFCD701AF9F493B0F96D" box="[1059,1181,1643,1670]" italics="true" pageId="28" pageNumber="125">S. theodori</emphasis>
|
||
</taxonomicName>
|
||
, but the latter species have long epiphallic caecum, short flagellum, and spermatophore having spines about two-thirds of the tail filament length.
|
||
</paragraph>
|
||
<caption id="DF536663036FFFCD74B8FAB494CEFA72" ID-DOI="http://doi.org/10.5281/zenodo.8349003" ID-Zenodo-Dep="8349003" httpUri="https://zenodo.org/record/8349003/files/figure.png" pageId="28" pageNumber="125" startId="28.[129,196,1324,1345]" targetBox="[223,1092,164,1305]" targetPageId="28" targetType="figure">
|
||
<paragraph id="8B9336EB036FFFCD74B8FAB494CEFA72" blockId="28.[129,1168,1324,1434]" pageId="28" pageNumber="125">
|
||
FIGURE 10 Spermatophore of
|
||
<taxonomicName id="4C2C4D68036FFFCD759CFAB4951BFAA9" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[421,566,1324,1345]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="125" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036FFFCD759CFAB4951BFAA9" box="[421,566,1324,1345]" italics="true" pageId="28" pageNumber="125">Sarika costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036FFFCD7602FAB59552FAAA" box="[571,639,1325,1346]" pageId="28" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype CUMZ 7945. A) General view of spermatophore that was broken into two parts. B) Head filament. C–E) Tail filament. C) Three spines located close to the sperm sac. D) Region with and without branching spines. E) Branching spines on the tip region. Red dashed line indicates a series of ten branching spines near the middle length.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF536663036EFFCC745BFB8A963CFB68" ID-DOI="http://doi.org/10.5281/zenodo.8349005" ID-Zenodo-Dep="8349005" httpUri="https://zenodo.org/record/8349005/files/figure.png" pageId="29" pageNumber="126" startId="29.[98,165,1042,1063]" targetBox="[193,1062,164,1023]" targetPageId="29" targetType="figure">
|
||
<paragraph id="8B9336EB036EFFCC745BFB8A963CFB68" blockId="29.[98,1111,1042,1152]" pageId="29" pageNumber="126">
|
||
FIGURE 11 Radula of
|
||
<taxonomicName id="4C2C4D68036EFFCC7501FB8A969DFBC0" box="[312,432,1042,1064]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="undefined-A">
|
||
<emphasis id="B958EAF9036EFFCC7501FB8A965CFBCF" box="[312,369,1042,1063]" italics="true" pageId="29" pageNumber="126">Sarika</emphasis>
|
||
spp. A
|
||
</taxonomicName>
|
||
)
|
||
<taxonomicName id="4C2C4D68036EFFCC7582FB8A9537FBCF" authorityName=": Godwin-Austen" authorityYear="1907" box="[443,538,1042,1063]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="theodori">
|
||
<emphasis id="B958EAF9036EFFCC7582FB8A9537FBCF" box="[443,538,1042,1063]" italics="true" pageId="29" pageNumber="126">S. theodori</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 7942. B)
|
||
<taxonomicName id="4C2C4D68036EFFCC773EFB8A945EFBC0" box="[775,883,1042,1064]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="siamensis">
|
||
<emphasis id="B958EAF9036EFFCC773EFB8A945EFBC0" box="[775,883,1042,1064]" italics="true" pageId="29" pageNumber="126">S. siamensis</emphasis>
|
||
</taxonomicName>
|
||
specimen CUMZ 7951. C)
|
||
<taxonomicName id="4C2C4D68036EFFCC74E2FBA8966FFBAC" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[219,322,1071,1093]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036EFFCC74E2FBA8966FFBAC" box="[219,322,1071,1093]" italics="true" pageId="29" pageNumber="126">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036EFFCC757EFBA896A6FBAD" box="[327,395,1072,1093]" pageId="29" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype CUMZ 7945. D)
|
||
<taxonomicName id="4C2C4D68036EFFCC764BFBA895EAFBAD" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[626,711,1072,1093]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="costata" status="sp. nov.">
|
||
<emphasis id="B958EAF9036EFFCC764BFBA895EAFBAD" box="[626,711,1072,1093]" italics="true" pageId="29" pageNumber="126">S. costata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036EFFCC76F2FBA89422FBAD" box="[715,783,1072,1093]" pageId="29" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype CUMZ 7947. Central tooth indicated by ‘C’ and blue colour; yellow colour indicates lateral teeth in the transition to marginal teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9336EB036EFFCC74B8FA9C954CF926" blockId="29.[98,609,1213,1742]" pageId="29" pageNumber="126">
|
||
Compared with other ariophantids with ribbed shells,
|
||
<taxonomicName id="4C2C4D68036EFFCC7538FAB196ACFAAA" authority="Pholyotha & Sutcharit & Lin & Panha, 2022" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[257,385,1320,1347]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="costabilis" status="sp. nov.">
|
||
<emphasis id="B958EAF9036EFFCC7538FAB196ACFAAA" box="[257,385,1320,1347]" italics="true" pageId="29" pageNumber="126">S. costabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B5782036EFFCC75B0FAB196F0FAAB" box="[393,477,1321,1347]" pageId="29" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="4C2C4D68036EFFCC7524FAD497B1FA63" authority="(Benson, 1856)" baseAuthorityName="Benson" baseAuthorityYear="1856" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="textrina">
|
||
<emphasis id="B958EAF9036EFFCC7524FAD496D5FA8E" box="[285,504,1356,1382]" italics="true" pageId="29" pageNumber="126">Hemiplecta textrina</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD4B1A036EFFCC763FFAD497B8FA63" author="Benson, W. H." pageId="29" pageNumber="126" pagination="249 - 254" refId="ref14962" refString="Benson, W. H. (1856) Descriptions of one Indian and nine new Burmese Helices; and notes on two Burmese Cyclostomacea. Ann. Mag. Nat. Hist., 18, 249 - 254." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by a coarse shell sculpture and descending aperture. In contrast, the shell sculpture of
|
||
<taxonomicName id="4C2C4D68036EFFCC745BFA2197F7FA3A" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[98,218,1465,1490]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="textrina">
|
||
<emphasis id="B958EAF9036EFFCC745BFA2197F7FA3A" box="[98,218,1465,1490]" italics="true" pageId="29" pageNumber="126">H. textrina</emphasis>
|
||
</taxonomicName>
|
||
is slightly coarse and its aperture is simple (
|
||
<bibRefCitation id="EFBD4B1A036EFFCC74E5FA449573FA1E" author="Blanford, W. T. & Godwin-Austen, H. H." box="[220,606,1500,1526]" pageId="29" pageNumber="126" refId="ref15040" refString="Blanford, W. T. & Godwin-Austen, H. H. (1908) The fauna of British India. Mollusca. Testacellidae and Zonitidae. In: C. T. Bingham (Ed) The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. https: // www. biodiversitylibrary. org / item / 47068 # page / 5 / mode / 1 up." type="url" year="1908">Blanford & Godwin-Austen, 1908</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD4B1A036EFFCC745BF9989641F9F2" author="Sutcharit, C. & Panha, S." box="[98,364,1536,1562]" pageId="29" pageNumber="126" pagination="101 - 154" refId="ref18564" refString="Sutcharit, C. & Panha, S. (2021) Systematic review of the dextral Hemiplecta Albers, 1850 (Eupulmonata, Ariophantidae) from Thailand with description of a new species and list of all the Indochinese species. ZooKeys, 1047, 101 - 154. https: // doi. org / 10.3897 / zookeys. 1047.65735." type="journal article" year="2021">Sutcharit & Panha, 2021</bibRefCitation>
|
||
).This new species can be separated from
|
||
<taxonomicName id="4C2C4D68036EFFCC757DF9BD97D1F98A" authority="(Tapparone-Canefri, 1889)" baseAuthorityName="Tapparone-Canefri" baseAuthorityYear="1889" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="auriettae">
|
||
<emphasis id="B958EAF9036EFFCC757DF9BD96E6F9D6" box="[324,459,1573,1598]" italics="true" pageId="29" pageNumber="126">H. auriettae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBD4B1A036EFFCC75E7F9BC97D8F98A" author="Tapparone-Canefri, C." pageId="29" pageNumber="126" pagination="295 - 359" refId="ref18806" refString="Tapparone-Canefri, C. (1889) Viaggio de Leonardo Fea in Birmania e regioni vicine. XVIII. Molluschi terrestri e dacqua dolce. Annali del Museo Civico di Storia Naturale di Genova. 27 [= series 2 volume 7]: 295 - 359. https: // www. biodiversitylibrary. org / page / 35995965." type="journal article" year="1889">Tapparone-Canefri, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by its small and narrow umbilicus. In contrast, the umbilicus of
|
||
<taxonomicName id="4C2C4D68036EFFCC75E6F9F4954CF96D" baseAuthorityName="Tapparone-Canefri" baseAuthorityYear="1889" box="[479,609,1644,1669]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="auriettae">
|
||
<emphasis id="B958EAF9036EFFCC75E6F9F4954CF96D" box="[479,609,1644,1669]" italics="true" pageId="29" pageNumber="126">H. auriettae</emphasis>
|
||
</taxonomicName>
|
||
is slightly widely opened and shows all whorls (
|
||
<bibRefCitation id="EFBD4B1A036EFFCC7453F92B96F7F926" author="Blanford, W. T. & Godwin-Austen, H. H." box="[106,474,1715,1742]" pageId="29" pageNumber="126" refId="ref15040" refString="Blanford, W. T. & Godwin-Austen, H. H. (1908) The fauna of British India. Mollusca. Testacellidae and Zonitidae. In: C. T. Bingham (Ed) The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. https: // www. biodiversitylibrary. org / item / 47068 # page / 5 / mode / 1 up." type="url" year="1908">Blanford & Godwin-Austen, 1908</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBD4B1A036EFFCC75DBF92B954CF926" author="Sutcharit, C. & Panha, S." box="[482,609,1715,1742]" pageId="29" pageNumber="126" pagination="101 - 154" refId="ref18564" refString="Sutcharit, C. & Panha, S. (2021) Systematic review of the dextral Hemiplecta Albers, 1850 (Eupulmonata, Ariophantidae) from Thailand with description of a new species and list of all the Indochinese species. ZooKeys, 1047, 101 - 154. https: // doi. org / 10.3897 / zookeys. 1047.65735." type="journal article" year="2021">Sutcharit &</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036EFFCC76BCFB25944BFAF6" blockId="29.[645,1157,1213,1454]" pageId="29" pageNumber="126">
|
||
Panha, 2021). However, the anatomical data of
|
||
<taxonomicName id="4C2C4D68036EFFCC76BCFB7995D7FB12" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[645,762,1249,1274]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="textrina">
|
||
<emphasis id="B958EAF9036EFFCC76BCFB7995D7FB12" box="[645,762,1249,1274]" italics="true" pageId="29" pageNumber="126">H. textrina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2C4D68036EFFCC7709FB79949DFB12" baseAuthorityName="Tapparone-Canefri" baseAuthorityYear="1889" box="[816,944,1249,1274]" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="auriettae">
|
||
<emphasis id="B958EAF9036EFFCC7709FB79949DFB12" box="[816,944,1249,1274]" italics="true" pageId="29" pageNumber="126">H. auriettae</emphasis>
|
||
</taxonomicName>
|
||
are unavailable for further comparison.
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036EFFCC769DFAB093A8FAAA" blockId="29.[645,1157,1213,1454]" box="[676,1157,1320,1347]" pageId="29" pageNumber="126">
|
||
<emphasis id="B958EAF9036EFFCC769DFAB094DEFAAB" bold="true" box="[676,1011,1320,1347]" pageId="29" pageNumber="126">
|
||
<taxonomicName id="4C2C4D68036EFFCC769DFAB0946AFAAB" authorityName="Pholyotha & Sutcharit & Lin & Panha" authorityYear="2022" box="[676,839,1320,1347]" class="Gastropoda" family="Ariophantidae" genus="Sarika" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="126" phylum="Mollusca" rank="species" species="costata">
|
||
<emphasis id="B958EAF9036EFFCC769DFAB0946AFAAB" bold="true" box="[676,839,1320,1347]" italics="true" pageId="29" pageNumber="126">Sarika costata</emphasis>
|
||
</taxonomicName>
|
||
species group:
|
||
</emphasis>
|
||
species with
|
||
</paragraph>
|
||
<paragraph id="8B9336EB036EFFCC76BCFAD595C4FA46" blockId="29.[645,1157,1213,1454]" pageId="29" pageNumber="126">a shell surface having fine radial ribs without fine impressed spiral lines and with five mantle lobes.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |