treatments-xml/data/03/85/87/038587F4683AFFC14E8D659DFC6BFF7A.xml

<document id="B13BD1B881795DC3D9279AD0FBA301FB" ID-CLB-Dataset="7213" ID-DOI="10.11646/zootaxa.4958.1.31" ID-GBIF-Dataset="a1df68e3-0e4c-4010-9556-cb15d51a5499" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692369" ID-ZooBank="9E63F8E2-1CCA-4F58-9019-39AE7A3D0640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1618471284689" checkinUser="plazi" docAuthor="Salini, S. &amp; Kment, Petr" docDate="2021" docId="038587F4683AFFC14E8D659DFC6BFF7A" docLanguage="en" docName="zootaxa.4958.1.31.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agathocles joceliae Salini &amp; Kment 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="542" masterDocId="FFBCFF8C6823FFE14E1A672FFFFDFF80" masterDocTitle="The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species" masterLastPageNumber="559" masterPageNumber="510" pageNumber="535" updateTime="1698936917073" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="097A26F663E842E983FE74EF84578FC9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="893DA1E2A518F9DC0FCFF8F885432EAE">
<mods:title id="6832B23F33E8EEAEDE487DB1CAB505AB">The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="69C64B4FA45E7351124D7A9FA9CDFE7F" type="personal">
<mods:role id="519168B06B9DD6AAA991861076B7675A">
<mods:roleTerm id="FD56EC76445DDB3B5B5030423CAD8218">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A08EA2D2FFD4E86A2EE1CA08E38FBB4D">Salini, S.</mods:namePart>
<mods:affiliation id="54CBBDC6B08AE1BFA7F6CF97EBF7977E">National Bureau of Agricultural Insect Resources, Bangalore, 560024, India</mods:affiliation>
</mods:name>
<mods:name id="918A299EADA7E7AD79BF625750100144" type="personal">
<mods:role id="2524B140668C65ADED63F6DAE4A0874D">
<mods:roleTerm id="F80B07895FE1B9C98B5F1E5DCB094565">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A787139FC6671E422D11D7A2238A7FC">Kment, Petr</mods:namePart>
<mods:nameIdentifier id="9C2FF93AE528934FFCC8D3DE29B1F5E1" type="ORCID">0000-0002-7026-5691</mods:nameIdentifier>
<mods:affiliation id="D51DCF3AFDEFA181E50953357749901A">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
<mods:nameIdentifier id="14CBAEDED40520FEB4F165A9B52C2415" type="email">sigara@post.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E60BF40F5EA523CB385C2E8D9B41A09B">text</mods:typeOfResource>
<mods:relatedItem id="B3494F23B9BD04EF97C902257C1B9A3A" type="host">
<mods:titleInfo id="5F6CE1EA3CCCD8E6EE088BA2CAEFF020">
<mods:title id="5630ECD3F0FE410A533D5ABC754DEE8B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0E6119BAF2E5404A7210FC7BC89AE049">
<mods:date id="3767E01ABD5C67C0AF2CD532E078FCFB">2021</mods:date>
<mods:detail id="B300749FB47C31FC06910BA448A8E607" type="pubDate">
<mods:number id="0B28346B88ADAC08054CF7DB920D706C">2021-04-14</mods:number>
</mods:detail>
<mods:detail id="4B7D827D33439B8FB183B23917D94DF3" type="volume">
<mods:number id="04A99C085F2A299DC60D91B4171B4462">4958</mods:number>
</mods:detail>
<mods:detail id="C794FEBB0989B9E561EFA256881905AC" type="issue">
<mods:number id="95789A330772FE852F6B5F67DE4C684A">1</mods:number>
</mods:detail>
<mods:extent id="99152C26E6FC88B6119B46E4F0DFB335" unit="page">
<mods:start id="FC8BD4FE53570EDA9AAB2E26D2F4ED32">510</mods:start>
<mods:end id="D6B828EFBCC660DAE82B99D1502CA359">559</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FA06BFC186848BD1061FBD7C172C6716">journal article</mods:classification>
<mods:identifier id="E6168BD965FFFE59A4B331A7590F5811" type="CLB-Dataset">7213</mods:identifier>
<mods:identifier id="3CB00F37EFF79715BFCAE0EECA0F6AA6" type="DOI">10.11646/zootaxa.4958.1.31</mods:identifier>
<mods:identifier id="E1DDDB7F8969673DD2BD278A7CCCBFF1" type="GBIF-Dataset">a1df68e3-0e4c-4010-9556-cb15d51a5499</mods:identifier>
<mods:identifier id="5E22A837134C9B4F5609A29307005092" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3135B0CBB18E6C258FC7B69ABB895F9A" type="Zenodo-Dep">4692369</mods:identifier>
<mods:identifier id="82A8FA1AFEDE330A332267818B0AA761" type="ZooBank">9E63F8E2-1CCA-4F58-9019-39AE7A3D0640</mods:identifier>
</mods:mods>
<treatment id="038587F4683AFFC14E8D659DFC6BFF7A" ID-DOI="http://doi.org/10.5281/zenodo.4711371" ID-GBIF-Taxon="181531961" ID-Zenodo-Dep="4711371" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A" httpUri="http://treatment.plazi.org/id/038587F4683AFFC14E8D659DFC6BFF7A" lastPageId="32" lastPageNumber="542" pageId="25" pageNumber="535">
<subSubSection id="C3366569683AFFF84E8D659DFE1BFD4D" box="[151,486,690,717]" pageId="25" pageNumber="535" type="nomenclature">
<paragraph id="8B9336E2683AFFF84E8D659DFE1BFD4D" blockId="25.[151,701,690,753]" box="[151,486,690,717]" pageId="25" pageNumber="535">
<heading id="D0DB818E683AFFF84E8D659DFE1BFD4D" bold="true" box="[151,486,690,717]" fontSize="11" level="1" pageId="25" pageNumber="535" reason="1">
<emphasis id="B958EAF0683AFFF84E8D659DFE1BFD4D" bold="true" box="[151,486,690,717]" italics="true" pageId="25" pageNumber="535">
<taxonomicName id="4C2C4D61683AFFF84E8D659DFE7CFD4D" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[151,385,690,717]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="535" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">Agathocles joceliae</taxonomicName>
<taxonomicNameLabel id="A26B578B683AFFF84F92659CFE1BFD4D" box="[392,486,691,717]" pageId="25" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3366569683AFFF84E8D65F8FD40FD71" box="[151,701,726,753]" pageId="25" pageNumber="535" type="description">
<paragraph id="8B9336E2683AFFF84E8D65F8FD40FD71" blockId="25.[151,701,690,753]" box="[151,701,726,753]" pageId="25" pageNumber="535">
(
<figureCitation id="13172A67683AFFF84E8565F8FEDDFD71" box="[159,288,727,753]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="25" pageNumber="535">Figs 12–13</figureCitation>
, 
<figureCitation id="13172A67683AFFF84F3665F9FE62FD71" box="[300,415,726,753]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="25" pageNumber="535">15, 18, 20</figureCitation>
, 
<figureCitation id="13172A67683AFFF84FB065F8FE35FD71" box="[426,456,727,753]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="25" pageNumber="535">24</figureCitation>
, 
<figureCitation id="13172A67683AFFF84FCE65F8FE0FFD71" box="[468,498,727,753]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="25" pageNumber="535">32</figureCitation>
, 
<figureCitation id="13172A67683AFFF84FE465F8FDE1FD71" box="[510,540,727,753]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="25" pageNumber="535">36</figureCitation>
, 
<figureCitation id="13172A67683AFFF84C3265F9FD83FD71" box="[552,638,726,753]" captionStart-0="FIGURES 88–89" captionStart-1="FIGURES 90–96" captionStart-2="FIGURES 97–101" captionStart-3="FIGURES 102–107" captionStartId-0="25.[151,264,1912,1936]" captionStartId-1="26.[151,264,1564,1588]" captionStartId-2="27.[151,264,1687,1711]" captionStartId-3="28.[151,264,1763,1787]" captionTargetBox-0="[157,1428,892,1887]" captionTargetBox-1="[151,1436,181,1541]" captionTargetBox-2="[151,1436,453,1664]" captionTargetBox-3="[161,1424,189,1732]" captionTargetId-0="figure-264@25.[151,1436,885,1889]" captionTargetId-1="figure-19@26.[151,1436,181,1541]" captionTargetId-2="figure-147@27.[151,1436,453,1664]" captionTargetId-3="figure-19@28.[151,1436,181,1740]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="28" captionText-0="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." captionText-1="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." captionText-2="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." captionText-3="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi-0="http://doi.org/10.5281/zenodo.4692403" figureDoi-1="http://doi.org/10.5281/zenodo.4692405" figureDoi-2="http://doi.org/10.5281/zenodo.4692407" figureDoi-3="http://doi.org/10.5281/zenodo.4692409" httpUri-0="https://zenodo.org/record/4692403/files/figure.png" httpUri-1="https://zenodo.org/record/4692405/files/figure.png" httpUri-2="https://zenodo.org/record/4692407/files/figure.png" httpUri-3="https://zenodo.org/record/4692409/files/figure.png" pageId="25" pageNumber="535">88–105</figureCitation>
, 
<figureCitation id="13172A67683AFFF84C9065F8FD48FD71" box="[650,693,727,753]" captionStart="FIGURE 139" captionStartId="39.[151,250,1769,1793]" captionTargetBox="[151,1435,1007,1745]" captionTargetId="figure-340@39.[151,1435,1007,1745]" captionTargetPageId="39" captionText="FIGURE 139. Distribution of the Agathocles species. T—type locality;?—doubtful record. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692423" httpUri="https://zenodo.org/record/4692423/files/figure.png" pageId="25" pageNumber="535">139</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3366569683AFFFA4E8D6430FC71FE2D" lastPageId="27" lastPageNumber="537" pageId="25" pageNumber="535" type="materials_examined">
<paragraph id="8B9336E2683AFFF84E8D6430FD53FCDD" blockId="25.[151,1436,798,861]" pageId="25" pageNumber="535">
<emphasis id="B958EAF0683AFFF84E8D6430FECBFCB8" bold="true" box="[151,310,798,825]" pageId="25" pageNumber="535">Type locality.</emphasis>
<materialsCitation id="3B443CBF683AFFF84F276430FE3FFCDD" ID-GBIF-Occurrence="3083795319" country="Malaysia" elevation="1550" latitude="4.63" location="Kampung Raja" longLatPrecision="783" longitude="101.45" pageId="25" pageNumber="535" specimenCount="1" stateProvince="Kelantan">
<collectingCountry id="F33B7672683AFFF84F276430FE5BFCB9" box="[317,422,799,825]" name="Malaysia" pageId="25" pageNumber="535">Malaysia</collectingCountry>
, W 
<collectingRegion id="49E8F800683AFFF84FC96430FDC6FCB9" box="[467,571,799,825]" country="Malaysia" name="Kelantan" pageId="25" pageNumber="535">Kelantan</collectingRegion>
, road between 
<location id="8EF36039683AFFF84CFE6430FC70FCB9" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF36039683AFFF84CFE6430FC70FCB9" box="[740,909,799,825]" country="Malaysia" latitude="4.63" longLatPrecision="783" longitude="101.45" name="Kampung Raja" pageId="25" pageNumber="535" stateProvince="Kelantan">Kampung Raja</location>
and Gua Musang, 
<elevation id="0001D1D1683AFFF84A796430FAC6FCB9" box="[1123,1339,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="25" pageNumber="535" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">
<quantity id="4CD49B07683AFFF84A796430FB02FCB9" box="[1123,1279,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="25" pageNumber="535" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400‒1700 m</quantity>
a.s.l.
</elevation>
, Ladang Pandrak, 
<geoCoordinate id="EE185025683AFFF84F1B646CFEACFCDD" box="[257,337,835,861]" degrees="4.63" direction="north" orientation="latitude" pageId="25" pageNumber="535" precision="555" value="4.63">4.63°N</geoCoordinate>
<geoCoordinate id="EE185025683AFFF84F42646DFE3FFCDD" box="[344,450,834,861]" degrees="101.45" direction="east" orientation="longitude" pageId="25" pageNumber="535" precision="555" value="101.45">101.45°E</geoCoordinate>
</materialsCitation>
to 
<geoCoordinate id="EE185025683AFFF84FFF646CFDCBFCDD" box="[485,566,835,861]" degrees="4.88" direction="north" orientation="latitude" pageId="25" pageNumber="535" precision="555" value="4.88">4.88°N</geoCoordinate>
<geoCoordinate id="EE185025683AFFF84C27646DFD54FCDD" box="[573,681,834,861]" degrees="101.95" direction="east" orientation="longitude" pageId="25" pageNumber="535" precision="555" value="101.95">101.95°E</geoCoordinate>
.
</paragraph>
<caption id="DF53666A683AFFF84E8D6057FE37F834" ID-DOI="http://doi.org/10.5281/zenodo.4692403" ID-Zenodo-Dep="4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="25" pageNumber="535" startId="25.[151,264,1912,1936]" targetBox="[157,1428,892,1887]" targetPageId="25">
<paragraph id="8B9336E2683AFFF84E8D6057FE37F834" blockId="25.[151,1437,1912,1972]" pageId="25" pageNumber="535">
<emphasis id="B958EAF0683AFFF84E8D6057FEAAF810" bold="true" box="[151,343,1912,1936]" pageId="25" pageNumber="535">FIGURES 88–89.</emphasis>
<taxonomicName id="4C2C4D61683AFFF84F7A6057FDD7F810" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[352,554,1912,1936]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="535" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0683AFFF84F7A6057FDD7F810" box="[352,554,1912,1936]" italics="true" pageId="25" pageNumber="535">Agathocles joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0683AFFF84C296056FD7AF811" bold="true" box="[563,647,1913,1937]" pageId="25" pageNumber="535">
<taxonomicNameLabel id="A26B578B683AFFF84C296056FD7AF811" box="[563,647,1913,1937]" pageId="25" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment.
</paragraph>
</caption>
<caption id="DF53666A6839FFFB4E8D6133FBE3F920" ID-DOI="http://doi.org/10.5281/zenodo.4692405" ID-Zenodo-Dep="4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="26" pageNumber="536" startId="26.[151,264,1564,1588]" targetBox="[151,1436,181,1541]" targetPageId="26">
<paragraph id="8B9336E26839FFFB4E8D6133FBE3F920" blockId="26.[151,1436,1564,1697]" pageId="26" pageNumber="536">
<emphasis id="B958EAF06839FFFB4E8D6133FEABF9B5" bold="true" box="[151,342,1564,1589]" pageId="26" pageNumber="536">FIGURES 90–96.</emphasis>
Male genital capsule of 
<taxonomicName id="4C2C4D616839FFFB4C4C6132FCE2F9B5" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[598,799,1565,1589]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="536" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF06839FFFB4C4C6132FCE2F9B5" box="[598,799,1565,1589]" italics="true" pageId="26" pageNumber="536">Agathocles joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF06839FFFB4D3D6132FC84F9B5" bold="true" box="[807,889,1565,1589]" pageId="26" pageNumber="536">
<taxonomicNameLabel id="A26B578B6839FFFB4D3D6132FC84F9B5" box="[807,889,1565,1589]" pageId="26" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment.
</paragraph>
</caption>
<paragraph id="8B9336E26839FFFA4EDD61DCFEFFFF55" blockId="26.[151,1436,1779,2021]" lastBlockId="27.[151,1437,151,431]" lastPageId="27" lastPageNumber="537" pageId="26" pageNumber="536">
<emphasis id="B958EAF06839FFFB4EDD61DCFE8BF88D" bold="true" box="[199,374,1779,1805]" pageId="26" pageNumber="536">Type material.</emphasis>
<materialsCitation id="3B443CBF6839FFFB4F6461DCFBE9F81D" ID-GBIF-Occurrence="3083795327" collectingDate="2006-04-01" collectingDateMax="2006-04-28" collectingDateMin="2006-04-01" collectionCode="NHMW, MUSEUM, WIEN" country="Malaysia" county="Road" elevation="1550" location="Gua Musang" longitude="101.75" municipality="Kampung Raja" pageId="26" pageNumber="536" specimenCount="2" specimenCount-male="2" stateProvince="Kelantan" typeStatus="holotype">
<typeStatus id="549788406839FFFB4F6461DCFE16F88D" box="[382,491,1779,1805]" pageId="26" pageNumber="536" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D2AFD6B6839FFFB4FEC61DCFDF6F88D" box="[502,523,1779,1805]" pageId="26" pageNumber="536" type="male">♂</specimenCount>
(
<collectionCode id="ED3DAE276839FFFB4C0161DCFD80F88D" box="[539,637,1779,1805]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="536" type="Museum">NHMW</collectionCode>
), ‘ 
<collectingCountry id="F33B76726839FFFB4C8F61DCFCD3F88D" box="[661,814,1779,1805]" name="Malaysia" pageId="26" pageNumber="536">MALAYSIA</collectingCountry>
, W 
<collectingRegion id="49E8F8006839FFFB4D4661DCFC0FF88D" box="[860,1010,1779,1805]" country="Malaysia" name="Kelantan" pageId="26" pageNumber="536">KELANTAN</collectingRegion>
/ 
<collectingCounty id="62F24E6E6839FFFB4A1061DCFBB8F88D" box="[1034,1093,1779,1805]" pageId="26" pageNumber="536">Road</collectingCounty>
between 
<collectingMunicipality id="6BF7AC986839FFFB4AA861DCFAA1F88D" box="[1202,1372,1779,1805]" pageId="26" pageNumber="536">Kampung Raja</collectingMunicipality>
/ and 
<location id="8EF360396839FFFB4E8D6038FED1F8B1" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396839FFFB4E8D6038FED1F8B1" box="[151,300,1815,1841]" country="Malaysia" county="Road" longitude="101.75" municipality="Kampung Raja" name="Gua Musang" pageId="26" pageNumber="536" stateProvince="Kelantan">Gua Musang</location>
, 
<quantity id="4CD49B076839FFFB4F2A6038FE3DF8B1" box="[304,448,1815,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="26" pageNumber="536" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">
<elevation id="0001D1D16839FFFB4F2A6038FE3DF8B1" box="[304,448,1815,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="26" pageNumber="536" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400-1700m</elevation>
</quantity>
, / (Ladang Pandrak), 
<date id="FF9210226839FFFB4CAE6038FCB5F8B1" box="[692,840,1815,1841]" pageId="26" pageNumber="536" value="2006-04-01" valueMax="2006-04-28" valueMin="2006-04-01">
<collectingDate id="EFD6E9CA6839FFFB4CAE6038FCB5F8B1" box="[692,840,1815,1841]" pageId="26" pageNumber="536" value="2006-04-01" valueMax="2006-04-28" valueMin="2006-04-01">1.-28.iv.2006</collectingDate>
</date>
/ 4°63´N, 
<geoCoordinate id="EE1850256839FFFB4DDA6038FBD3F8B2" box="[960,1070,1815,1842]" degrees="101" direction="east" minutes="45" orientation="longitude" pageId="26" pageNumber="536" precision="925" value="101.75">101°45´E</geoCoordinate>
, 4°88´N-/ 101°95´E, 
<location id="8EF360396839FFFB4B3A6037FF39F8D5" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396839FFFB4B3A6037FF39F8D5" country="Malaysia" county="Road" longitude="101.75" municipality="Kampung Raja" name="Cechovsky Petr" pageId="26" pageNumber="536" stateProvince="Kelantan">Čechovský Petr</location>
lgt. [p]’ // ‘ 
<collectingCountry id="F33B76726839FFFB4F5D6014FE1DF8D5" box="[327,480,1851,1877]" name="Malaysia" pageId="26" pageNumber="536">MALAYSIA</collectingCountry>
, W 
<collectingRegion id="49E8F8006839FFFB4C156014FD58F8D5" box="[527,677,1851,1877]" country="Malaysia" name="Kelantan" pageId="26" pageNumber="536">KELANTAN</collectingRegion>
/ 
<location id="8EF360396839FFFB4CA56014FD07F8D5" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396839FFFB4CA56014FD07F8D5" box="[703,762,1851,1877]" country="Malaysia" county="Road" longitude="101.75" municipality="Kampung Raja" name="Road" pageId="26" pageNumber="536" stateProvince="Kelantan">Road</location>
between 
<location id="8EF360396839FFFB4D726014FBEEF8D5" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396839FFFB4D726014FBEEF8D5" box="[872,1043,1851,1877]" country="Malaysia" county="Road" longitude="101.75" municipality="Kampung Raja" name="Kampong Raja" pageId="26" pageNumber="536" stateProvince="Kelantan">Kampong Raja</location>
/ and 
<location id="8EF360396839FFFB4A476014FB0EF8D5" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396839FFFB4A476014FB0EF8D5" box="[1117,1267,1851,1877]" country="Malaysia" county="Road" longitude="101.75" municipality="Kampung Raja" name="Gua Musang" pageId="26" pageNumber="536" stateProvince="Kelantan">Gua Musang</location>
, 
<quantity id="4CD49B076839FFFB4AED6014FA7AF8D5" box="[1271,1415,1851,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="26" pageNumber="536" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">
<elevation id="0001D1D16839FFFB4AED6014FA7AF8D5" box="[1271,1415,1851,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.7" metricValueMin="1.4" pageId="26" pageNumber="536" unit="m" value="1550.0" valueMax="1700.0" valueMin="1400.0">1400-1700m</elevation>
</quantity>
, / (LadangPandrak), 
<date id="FF9210226839FFFB4F726070FE00F8F9" box="[360,509,1887,1913]" pageId="26" pageNumber="536" value="2006-04-01" valueMax="2006-04-28" valueMin="2006-04-01">
<collectingDate id="EFD6E9CA6839FFFB4F726070FE00F8F9" box="[360,509,1887,1913]" pageId="26" pageNumber="536" value="2006-04-01" valueMax="2006-04-28" valueMin="2006-04-01">1.-28.iv.2006</collectingDate>
</date>
/ 4°63´N, 
<geoCoordinate id="EE1850256839FFFB4C6F6070FD1FF8FA" box="[629,738,1887,1914]" degrees="101" direction="east" minutes="45" orientation="longitude" pageId="26" pageNumber="536" precision="925" value="101.75">101°45´E</geoCoordinate>
, 4°88´N- / 101°95´E, Cechovsky Petr lgt. [p, partly illegible]’ // COLLECTION / NATURHISTORISCHE / 
<collectionCode id="ED3DAE276839FFFB4CBF60ACFC8FF81D" box="[677,882,1923,1949]" pageId="26" pageNumber="536">MUSEUM WIEN</collectionCode>
[p] // 
<specimenCount id="9D2AFD6B6839FFFB4DA760ACFC2FF81D" box="[957,978,1923,1949]" pageId="26" pageNumber="536" type="male">♂</specimenCount>
[p] //
</materialsCitation>
<materialsCitation id="3B443CBF6839FFFB4A0760ACFA61F865" ID-GBIF-Occurrence="3083795356" collectionCode="II, III" collectorName="Salini &amp; Kment" country="Malaysia" pageId="26" pageNumber="536" specimenCount="2" stateProvince="Kelantan" typeStatus="holotype">
<typeStatus id="549788406839FFFB4A0760ACFB3EF81D" box="[1053,1219,1923,1949]" pageId="26" pageNumber="536" type="holotype">HOLOTYPUS</typeStatus>
/ 
<taxonomicName id="4C2C4D616839FFFB4AC760ACFA71F81D" box="[1245,1420,1923,1949]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06839FFFB4AC760ACFA71F81D" box="[1245,1420,1923,1949]" italics="true" pageId="26" pageNumber="536">AGATHOCLES</emphasis>
</taxonomicName>
/ 
<emphasis id="B958EAF06839FFFB4E8D6088FEEAF841" box="[151,279,1959,1985]" italics="true" pageId="26" pageNumber="536">JOCELIAE</emphasis>
/ sp. nov. / des. 
<collectorName id="26D953346839FFFB4FCC6088FDEAF841" box="[470,535,1959,1985]" pageId="26" pageNumber="536">Salini</collectorName>
&amp; 
<collectorName id="26D953346839FFFB4C256088FD76F841" box="[575,651,1959,1985]" pageId="26" pageNumber="536">Kment</collectorName>
2020 [p, red label]. 
<collectorName id="26D953346839FFFB4D6D6088FC5EF841" box="[887,931,1959,1985]" pageId="26" pageNumber="536">The</collectorName>
<typeStatus id="549788406839FFFB4DB16088FBF0F841" box="[939,1037,1959,1985]" pageId="26" pageNumber="536" type="holotype">holotype</typeStatus>
is card-mounted, complete, except detached genital capsule and right metatarsomeres 
<collectionCode id="ED3DAE276839FFFB4CC560E4FD0FF865" box="[735,754,1995,2021]" pageId="26" pageNumber="536">II</collectionCode>
+ 
<collectionCode id="ED3DAE276839FFFB4D0E60E4FCCDF865" box="[788,816,1995,2021]" pageId="26" pageNumber="536">III</collectionCode>
glued to the same piece of card. The original locality
</materialsCitation>
label is quite illegible; the data were reconstructed with help of 
<bibRefCitation id="EFBD4B136838FFFA4D6067B8FBE6FF31" author="Novak, V." box="[890,1051,151,177]" pageId="27" pageNumber="537" pagination="451 - 467" refId="ref32426" refString="Novak, V. (2018) New genera of Alleculinae (Coleoptera: Tenebrionidae: Alleculinae: Alleculini) from the Oriental Region. Part IX - Dorota gen. nov. Studies and Reports, Taxonomical Series, 14 (1), 451 - 467." type="journal article" year="2018">Novák (2018)</bibRefCitation>
, and replacement locality label is provided.
</paragraph>
<paragraph id="8B9336E26838FFFA4EDD67F0FC71FE2D" blockId="27.[151,1437,151,431]" pageId="27" pageNumber="537">
<materialsCitation id="3B443CBF6838FFFA4EDD67F0FD09FF79" ID-GBIF-Occurrence="3083795335" box="[199,756,222,249]" country="Malaysia" location="Kelantan" pageId="27" pageNumber="537" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Kelantan" typeStatus="paratype">
<typeStatus id="549788406838FFFA4EDD67F0FEC9FF79" box="[199,308,223,249]" pageId="27" pageNumber="537" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D2AFD6B6838FFFA4F5367F0FE87FF79" box="[329,378,223,249]" pageId="27" pageNumber="537" type="male">4 ♂</specimenCount>
<specimenCount id="9D2AFD6B6838FFFA4F9267F0FE4AFF79" box="[392,439,223,249]" pageId="27" pageNumber="537" type="female">6 ♀</specimenCount>
). 
<emphasis id="B958EAF06838FFFA4FCF67F1FD09FF79" bold="true" box="[469,756,222,249]" pageId="27" pageNumber="537">
<collectingCountry id="F33B76726838FFFA4FCF67F1FD92FF78" box="[469,623,222,248]" name="Malaysia" pageId="27" pageNumber="537">MALAYSIA</collectingCountry>
: 
<collectingRegion id="49E8F8006838FFFA4C9B67F0FD09FF79" box="[641,756,223,249]" country="Malaysia" name="Kelantan" pageId="27" pageNumber="537">Kelantan</collectingRegion>
</emphasis>
</materialsCitation>
<emphasis id="B958EAF06838FFFA4CEE67F0FD04FF79" bold="true" box="[756,761,223,249]" pageId="27" pageNumber="537">:</emphasis>
<materialsCitation id="3B443CBF6838FFFA4D1C67F0FA65FE9D" ID-GBIF-Occurrence="3083795347" collectingDate="2011-05-27" collectingDateMax="2011-06-19" collectingDateMin="2011-05-27" collectionCode="NHMW" collectorName="Kampong Sungai Om &amp; Petr Cechovsky" country="Malaysia" elevation="900" latitude="4.9027777" location="Ulu Lalat Mt." longLatPrecision="21" longitude="101.8275" pageId="27" pageNumber="537" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kelantan" typeStatus="paratype">
<locationDeviation id="9A212FC06838FFFA4D1C67F0FBA5FF79" box="[774,1112,223,249]" location="Gua Musang" pageId="27" pageNumber="537">
<quantity id="4CD49B076838FFFA4D1C67F0FCAEFF79" box="[774,851,223,249]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="537" unit="km" value="30.0">30 km</quantity>
NW of Gua Musang
</locationDeviation>
, 
<location id="8EF360396838FFFA4A7367F0FAF2FF79" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396838FFFA4A7367F0FAF2FF79" box="[1129,1295,223,249]" country="Malaysia" latitude="4.9027777" longLatPrecision="21" longitude="101.8275" name="Ulu Lalat Mt." pageId="27" pageNumber="537" stateProvince="Kelantan">Ulu Lalat Mt.</location>
[
<geoCoordinate id="EE1850256838FFFA4B3867CFFA61FF7B" box="[1314,1436,224,251]" degrees="4" direction="north" minutes="54" orientation="latitude" pageId="27" pageNumber="537" precision="15" seconds="10" value="4.9027777">4°54′10″N</geoCoordinate>
<geoCoordinate id="EE1850256838FFFA4E8D662BFED1FE9F" box="[151,300,260,287]" degrees="101" direction="east" minutes="49" orientation="longitude" pageId="27" pageNumber="537" precision="15" seconds="39" value="101.8275">101°49′39″E</geoCoordinate>
], 
<elevation id="0001D1D16838FFFA4F24662CFDFAFE9D" box="[318,519,259,285]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="27" pageNumber="537" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<quantity id="4CD49B076838FFFA4F24662CFE36FE9D" box="[318,459,259,285]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="27" pageNumber="537" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</quantity>
a.s.l.
</elevation>
, 
<collectorName id="26D953346838FFFA4C09662CFCF4FE9D" box="[531,777,259,285]" pageId="27" pageNumber="537">Kampong Sungai Om</collectorName>
, 
<date id="FF9210226838FFFA4D0E662CFC2CFE9D" box="[788,977,259,285]" pageId="27" pageNumber="537" value="2011-05-27" valueMax="2011-06-19" valueMin="2011-05-27">
<collectingDate id="EFD6E9CA6838FFFA4D0E662CFC2CFE9D" box="[788,977,259,285]" pageId="27" pageNumber="537" value="2011-05-27" valueMax="2011-06-19" valueMin="2011-05-27">27.v.–19.vi.2011</collectingDate>
</date>
, 
<specimenCount id="9D2AFD6B6838FFFA4DC6662CFBFBFE9D" box="[988,1030,259,285]" pageId="27" pageNumber="537" type="male">1 ♂</specimenCount>
<specimenCount id="9D2AFD6B6838FFFA4A17662CFBCBFE9D" box="[1037,1078,259,285]" pageId="27" pageNumber="537" type="female">1 ♀</specimenCount>
, 
<collectorName id="26D953346838FFFA4A5E662CFB09FE9F" box="[1092,1268,259,287]" pageId="27" pageNumber="537">Petr Čechovský</collectorName>
lgt. (
<collectionCode id="ED3DAE276838FFFA4B34662CFA6DFE9D" box="[1326,1424,259,285]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="537" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B443CBF6838FFFA4E8D6608FA65FEC1" ID-GBIF-Occurrence="3083795338" box="[151,1432,295,323]" collectingDate="2010-06-21" collectingDateMax="2010-07-14" collectingDateMin="2010-06-21" collectionCode="NHMW, NMPC" collectorName="Petr Cechovsky" country="Malaysia" elevation="900" latitude="4.9027777" location="Ulu Lalat Mt." longLatPrecision="21" longitude="101.8275" pageId="27" pageNumber="537" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="Kelantan" typeStatus="paratype">
same locality, 
<date id="FF9210226838FFFA4F216608FDF6FEC1" box="[315,523,295,321]" pageId="27" pageNumber="537" value="2010-06-21" valueMax="2010-07-14" valueMin="2010-06-21">
<collectingDate id="EFD6E9CA6838FFFA4F216608FDF6FEC1" box="[315,523,295,321]" pageId="27" pageNumber="537" value="2010-06-21" valueMax="2010-07-14" valueMin="2010-06-21">21.vi.–14.vii.2010</collectingDate>
</date>
, 
<specimenCount id="9D2AFD6B6838FFFA4C026608FDB9FEC1" box="[536,580,295,321]" pageId="27" pageNumber="537" type="male">2 ♂</specimenCount>
<specimenCount id="9D2AFD6B6838FFFA4C566608FD85FEC1" box="[588,632,295,321]" pageId="27" pageNumber="537" type="female">4 ♀</specimenCount>
, 
<collectorName id="26D953346838FFFA4C926608FCC7FEC3" box="[648,826,295,323]" pageId="27" pageNumber="537">Petr Čechovský</collectorName>
lgt. (
<specimenCount id="9D2AFD6B6838FFFA4D616608FC58FEC1" box="[891,933,295,321]" pageId="27" pageNumber="537" type="male">1 ♂</specimenCount>
<specimenCount id="9D2AFD6B6838FFFA4DB76608FC24FEC1" box="[941,985,295,321]" pageId="27" pageNumber="537" type="female">3 ♀</specimenCount>
<collectionCode id="ED3DAE276838FFFA4DF86608FBC3FEC1" box="[994,1086,295,321]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="537" type="Museum">NHMW</collectionCode>
, 
<specimenCount id="9D2AFD6B6838FFFA4A516608FB8BFEC1" box="[1099,1142,295,321]" pageId="27" pageNumber="537" type="male">1 ♂</specimenCount>
<specimenCount id="9D2AFD6B6838FFFA4A656608FB56FEC1" box="[1151,1195,295,321]" pageId="27" pageNumber="537" type="female">1 ♀</specimenCount>
<collectionCode id="ED3DAE276838FFFA4AA96608FAF3FEC1" box="[1203,1294,295,321]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="537" type="Museum">NHMW</collectionCode>
→ 
<collectionCode id="ED3DAE276838FFFA4B266608FA6CFEC1" box="[1340,1425,295,321]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="27" pageNumber="537">NMPC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B443CBF6838FFFA4E8D6664FE4DFE09" ID-GBIF-Occurrence="3083795330" collectingDate="2018-04-10" collectingDateMax="2018-05-06" collectingDateMin="2018-04-10" collectionCode="NMPC" collectorName="Petr Cechovsky" country="Malaysia" latitude="5.423611" location="Kampong Timor" longLatPrecision="21" longitude="101.84667" municipality="Gunung Jual" pageId="27" pageNumber="537" specimenCount="1" specimenCount-female="1" stateProvince="Kelantan" typeStatus="paratype">
<locationDeviation id="9A212FC06838FFFA4E8D6664FEB8FEE5" box="[151,325,331,357]" location="Jeli" pageId="27" pageNumber="537">
<quantity id="4CD49B076838FFFA4E8D6664FF22FEE5" box="[151,223,331,357]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="537" unit="km" value="30.0">30 km</quantity>
S of Jeli
</locationDeviation>
[
<geoCoordinate id="EE1850256838FFFA4F4E6663FE33FEE7" box="[340,462,332,359]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="27" pageNumber="537" precision="15" seconds="25" value="5.423611">5°25′25″N</geoCoordinate>
<geoCoordinate id="EE1850256838FFFA4FCC6663FD96FEE7" box="[470,619,332,359]" degrees="101" direction="east" minutes="50" orientation="longitude" pageId="27" pageNumber="537" precision="15" seconds="48" value="101.84667">101°50′48″E</geoCoordinate>
], 
<collectingMunicipality id="6BF7AC986838FFFA4C656664FCECFEE5" box="[639,785,331,357]" pageId="27" pageNumber="537">Gunung Jual</collectingMunicipality>
, 
<location id="8EF360396838FFFA4D076664FC27FEE5" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396838FFFA4D076664FC27FEE5" box="[797,986,331,357]" country="Malaysia" latitude="5.423611" longLatPrecision="21" longitude="101.84667" municipality="Gunung Jual" name="Kampong Timor" pageId="27" pageNumber="537" stateProvince="Kelantan">Kampong Timor</location>
, 
<date id="FF9210226838FFFA4DFC6664FB68FEE5" box="[998,1173,331,357]" pageId="27" pageNumber="537" value="2018-04-10" valueMax="2018-05-06" valueMin="2018-04-10">
<collectingDate id="EFD6E9CA6838FFFA4DFC6664FB68FEE5" box="[998,1173,331,357]" pageId="27" pageNumber="537" value="2018-04-10" valueMax="2018-05-06" valueMin="2018-04-10">
10.iv.–6.v. 
<quantity id="4CD49B076838FFFA4A4C6664FB68FEE5" box="[1110,1173,331,357]" metricMagnitude="6" metricUnit="m" metricValue="2.0187999999999997" pageId="27" pageNumber="537" unit="m" value="2018800.0">2018</quantity>
</collectingDate>
</date>
, 800 m a.s.l., 
<specimenCount id="9D2AFD6B6838FFFA4B2F6664FA9DFEE5" box="[1333,1376,331,357]" pageId="27" pageNumber="537" type="female">1 ♀</specimenCount>
, 
<collectorName id="26D953346838FFFA4B756664FEEEFE0B" pageId="27" pageNumber="537">Petr Čechovský</collectorName>
lgt. (
<collectionCode id="ED3DAE276838FFFA4F486640FE55FE09" box="[338,424,367,393]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="27" pageNumber="537">NMPC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B443CBF6838FFFA4FA66640FC75FE2D" ID-GBIF-Occurrence="3083795323" collectingDate="2004-03-29" collectingDateMax="2004-04-15" collectingDateMin="2004-03-29" collectionCode="NHMW" collectorName="Petr Cechovsky" country="Malaysia" elevation="900" latitude="4.405139" location="Ringlet" longLatPrecision="1" longitude="101.38675" municipality="Banjaran Titiwangsa" pageId="27" pageNumber="537" specimenCount="1" specimenCount-male="1" stateProvince="Perak" typeStatus="paratype">
<emphasis id="B958EAF06838FFFA4FA66640FDF0FE09" bold="true" box="[444,525,367,393]" pageId="27" pageNumber="537">
<collectingRegion id="49E8F8006838FFFA4FA66640FDF5FE09" box="[444,520,367,393]" country="Malaysia" name="Perak" pageId="27" pageNumber="537">Perak</collectingRegion>
:
</emphasis>
<locationDeviation id="9A212FC06838FFFA4C0C6640FD1AFE09" box="[534,743,367,393]" location="Ipoh" pageId="27" pageNumber="537">
<quantity id="4CD49B076838FFFA4C0C6640FDA2FE09" box="[534,607,367,393]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="27" pageNumber="537" unit="km" value="40.0">40 km</quantity>
SE of Ipoh
</locationDeviation>
, 
<collectingMunicipality id="6BF7AC986838FFFA4CEE6640FC1DFE09" box="[756,992,367,393]" pageId="27" pageNumber="537">Banjaran Titiwangsa</collectingMunicipality>
, 
<location id="8EF360396838FFFA4DF76640FBC2FE09" LSID="urn:lsid:plazi:treatment:038587F4683AFFC14E8D659DFC6BFF7A:8EF360396838FFFA4DF76640FBC2FE09" box="[1005,1087,367,393]" country="Malaysia" latitude="4.405139" longLatPrecision="1" longitude="101.38675" municipality="Banjaran Titiwangsa" name="Ringlet" pageId="27" pageNumber="537" stateProvince="Perak">Ringlet</location>
[
<geoCoordinate id="EE1850256838FFFA4A55665FFB23FE0B" box="[1103,1246,368,395]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="27" pageNumber="537" precision="1" seconds="18.5" value="4.405139">4°24′18.5″N</geoCoordinate>
<geoCoordinate id="EE1850256838FFFA4AFD665FFA6CFE0B" box="[1255,1425,368,395]" degrees="101" direction="east" minutes="23" orientation="longitude" pageId="27" pageNumber="537" precision="1" seconds="12.3" value="101.38675">101°23′12.3″E</geoCoordinate>
], 
<elevation id="0001D1D16838FFFA4E8D66BCFEE6FE2D" box="[151,283,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="537" unit="m" value="900.0">
<quantity id="4CD49B076838FFFA4E8D66BCFF23FE2D" box="[151,222,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="27" pageNumber="537" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
, 
<date id="FF9210226838FFFA4F3D66BDFE12FE2D" box="[295,495,402,429]" pageId="27" pageNumber="537" value="2004-03-29" valueMax="2004-04-15" valueMin="2004-03-29">
<collectingDate id="EFD6E9CA6838FFFA4F3D66BDFE12FE2D" box="[295,495,402,429]" pageId="27" pageNumber="537" value="2004-03-29" valueMax="2004-04-15" valueMin="2004-03-29">29.iii.–15.iv.2004</collectingDate>
</date>
, 
<specimenCount id="9D2AFD6B6838FFFA4FE066BCFDDBFE2D" box="[506,550,403,429]" pageId="27" pageNumber="537" type="male">1 ♂</specimenCount>
, 
<collectorName id="26D953346838FFFA4C2866BCFD1FFE2F" box="[562,738,403,431]" pageId="27" pageNumber="537">Petr Čechovský</collectorName>
lgt. (
<collectionCode id="ED3DAE276838FFFA4D0766BCFC82FE2D" box="[797,895,403,429]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="537" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF53666A6838FFFA4E8D61B8FC4BF952" ID-DOI="http://doi.org/10.5281/zenodo.4692407" ID-Zenodo-Dep="4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="27" pageNumber="537" startId="27.[151,264,1687,1711]" targetBox="[151,1436,453,1664]" targetPageId="27">
<paragraph id="8B9336E26838FFFA4E8D61B8FC4BF952" blockId="27.[151,1436,1687,1747]" pageId="27" pageNumber="537">
<emphasis id="B958EAF06838FFFA4E8D61B8FE9EF92F" bold="true" box="[151,355,1687,1711]" pageId="27" pageNumber="537">FIGURES 97–101.</emphasis>
Paramere of 
<taxonomicName id="4C2C4D616838FFFA4FE861B8FD41F92F" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[498,700,1687,1711]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="537" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF06838FFFA4FE861B8FD41F92F" box="[498,700,1687,1711]" italics="true" pageId="27" pageNumber="537">Agathocles joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF06838FFFA4CDF61B8FCEAF92F" bold="true" box="[709,791,1687,1711]" pageId="27" pageNumber="537">
<taxonomicNameLabel id="A26B578B6838FFFA4CDF61B8FCEAF92F" box="[709,791,1687,1711]" pageId="27" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment.
</paragraph>
</caption>
<subSubSection id="C33665696838FFFC4EDD602EFB21F8CE" lastPageId="29" lastPageNumber="539" pageId="27" pageNumber="537" type="description">
<paragraph id="8B9336E26838FFFA4EDD602EFD5BF873" blockId="27.[151,1436,1793,2035]" pageId="27" pageNumber="537">
<emphasis id="B958EAF06838FFFA4EDD602EFE12F89C" bold="true" box="[199,495,1793,1820]" pageId="27" pageNumber="537">
Description. 
<emphasis id="B958EAF06838FFFA4F79602EFE12F89C" bold="true" box="[355,495,1793,1820]" italics="true" pageId="27" pageNumber="537">Colouration</emphasis>
</emphasis>
(
<figureCitation id="13172A676838FFFA4C18602EFD74F89B" box="[514,649,1793,1819]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="27" pageNumber="537">Figs 88–89</figureCitation>
). Body above and below, including posterior portion of pygophore, black, except clavus and corium dark brown; connexival segments ventrally with narrow brown stripe sublaterally; acetabula and peritremal surface brownish; antennomeres I–III pale brown, distiflagellum (IV) pale ochraceous; legs brown, femora apically darker; labium and base of pygophore pale ochraceous. Apex of scutellum with large yellowish spot, smaller yellowish callosities on apices of humeral angles, at the end of median furrow on corium, and anterolaterally on metepimeron; dorsal surface of body, especially on corium, with very small yellowish ochraceous callosities. Membrane dark brown, translucent.
</paragraph>
<caption id="DF53666A683FFFFD4E8D61CCFD22F8E7" ID-DOI="http://doi.org/10.5281/zenodo.4692409" ID-Zenodo-Dep="4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="28" pageNumber="538" startId="28.[151,264,1763,1787]" targetBox="[161,1424,189,1732]" targetPageId="28">
<paragraph id="8B9336E2683FFFFD4E8D61CCFD22F8E7" blockId="28.[151,1437,1763,1896]" pageId="28" pageNumber="538">
<emphasis id="B958EAF0683FFFFD4E8D61CCFE8FF97C" bold="true" box="[151,370,1763,1788]" pageId="28" pageNumber="538">FIGURES 102–107.</emphasis>
Penis of 
<taxonomicName id="4C2C4D61683FFFFD4FC661CBFDB0F97C" box="[476,589,1764,1788]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0683FFFFD4FC661CBFDB0F97C" box="[476,589,1764,1788]" italics="true" pageId="28" pageNumber="538">Agathocles</emphasis>
</taxonomicName>
species. 102–105— 
<taxonomicName id="4C2C4D61683FFFFD4D3961CBFC6EF97C" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[803,915,1764,1788]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="538" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0683FFFFD4D3961CBFC6EF97C" box="[803,915,1764,1788]" italics="true" pageId="28" pageNumber="538">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0683FFFFD4D8461CBFC0FF97C" bold="true" box="[926,1010,1764,1788]" pageId="28" pageNumber="538">
<taxonomicNameLabel id="A26B578B683FFFFD4D8461CBFC0FF97C" box="[926,1010,1764,1788]" pageId="28" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107— 
<taxonomicName id="4C2C4D61683FFFFD4A046027FAC1F8A0" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[1054,1340,1800,1824]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="538" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0683FFFFD4A046027FB6BF8A0" box="[1054,1174,1800,1824]" italics="true" pageId="28" pageNumber="538">A. normalis</emphasis>
(Distant, 1901)
</taxonomicName>
, inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment.
</paragraph>
</caption>
<paragraph id="8B9336E2683EFFFC4EDD67B8FE2CFEC1" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD67B8FE09FF31" bold="true" box="[199,500,150,177]" italics="true" pageId="29" pageNumber="539">Integument and vestiture.</emphasis>
Body shining, clavus, corium and evaporatorium matt. Body above covered with dense, coarse, black punctures, except impunctate apex of scutellum; endocorium with punctures sparser but equally large as on exocorium; disc of pronotum (posteromedially) and scutellum (anteriorly) with prominent transverse wrinkles (
<figureCitation id="13172A67683EFFFC4F1D662CFEA7FE9D" box="[263,346,259,285]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="29" pageNumber="539">Fig. 88</figureCitation>
); head underside and thoracic sterna with dense, coarse punctures; surface of abdominal sternites smooth with fine punctures.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD6664FB9AFDE1" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD6664FEC6FEE5" bold="true" box="[199,315,331,357]" italics="true" pageId="29" pageNumber="539">Structure.</emphasis>
Head (
<figureCitation id="13172A67683EFFFC4F946664FE1FFEE5" box="[398,482,331,357]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="29" pageNumber="539">Fig. 12</figureCitation>
). Head slightly shorter than wide (length: width ratio 0.79–0.88). Mandibular plates with lateral margins distinctly sinuate in front of eyes above the large antenniferous tubercle, parallel-sided about their midlength, and narrowly rounded anteriorly; mandibular plates not meeting anteriorly, apex of clypeus completely free, apical incisure in front of clypeus wide, trapeziform.Antennomeres from shortest to longest: I &lt;IIa &lt;IV &lt;IIb = III, but ratios of antennal segments highly variable. Scape (I) surpassing apex of head of about its halflength, basipedicellite (IIa) distinctly (1: 1.41‒1.93×) longer than scape and usually shorter than distipedicellite (IIb) (length of IIa: IIb ratio: 1: 1.00‒1.70). Labiomeres from shortest to longest (n = 1): IV (
<quantity id="4CD49B07683EFFFC4AD8650CFAD4FDBD" box="[1218,1321,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="29" pageNumber="539" unit="mm" value="1.23">1.23 mm</quantity>
) &lt;I (
<quantity id="4CD49B07683EFFFC4B70650DFF39FDE1" metricMagnitude="-3" metricUnit="m" metricValue="1.57" pageId="29" pageNumber="539" unit="mm" value="1.57">1.57 mm</quantity>
) &lt;III (
<quantity id="4CD49B07683EFFFC4F0F6568FE86FDE1" box="[277,379,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="29" pageNumber="539" unit="mm" value="1.86">1.86 mm</quantity>
) &lt;II (
<quantity id="4CD49B07683EFFFC4FD86568FDD1FDE1" box="[450,556,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" pageId="29" pageNumber="539" unit="mm" value="2.06">2.06 mm</quantity>
). Labium reaching posterior margin of mesocoxae.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD6544FE56FD4D" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD6544FECBFD04" box="[199,310,619,644]" italics="true" pageId="29" pageNumber="539">Pronotum</emphasis>
(
<figureCitation id="13172A67683EFFFC4F5F6544FE65FD05" box="[325,408,619,645]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="29" pageNumber="539">Fig. 88</figureCitation>
). Lateral tooth on anterolateral angles prominent, directed laterad (
<figureCitation id="13172A67683EFFFC4A986544FB28FD05" box="[1154,1237,619,645]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="29" pageNumber="539">Fig. 12</figureCitation>
). Lateral pronotal margin anteriorly slightly serrate. Humeri rectangular, slightly surpassing laterad over base of corium. Posterior margin slightly concave.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD65F8FF06FC95" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD65F8FEC8FD71" box="[199,309,727,753]" italics="true" pageId="29" pageNumber="539">Scutellum</emphasis>
(
<figureCitation id="13172A67683EFFFC4F5D65F8FE61FD71" box="[327,412,727,753]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="29" pageNumber="539">Fig. 88</figureCitation>
). Disc of scutellum convex in prefrenal portion, its postfrenal portion flat; apex narrowly rounded.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD6430FBCFFCB9" blockId="29.[151,1437,150,1977]" box="[199,1074,799,825]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD6430FEB8FCB9" box="[199,325,799,825]" italics="true" pageId="29" pageNumber="539">Hemelytra.</emphasis>
Costal margin of corium only slightly insinuated basally (
<figureCitation id="13172A67683EFFFC4DCB6430FBD9FCB9" box="[977,1060,799,825]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="29" pageNumber="539">Fig. 88</figureCitation>
).
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD646CFC96FC01" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD646CFEDDFCDD" box="[199,288,835,861]" italics="true" pageId="29" pageNumber="539">Thorax.</emphasis>
Ostiole large, oval, opening posteriad; median furrow short and wide, not apparent in apical half of peritreme length (
<figureCitation id="13172A67683EFFFC4F456448FE4FFC01" box="[351,434,871,897]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="29" pageNumber="539">Fig. 18</figureCitation>
). Metasternum medium wide (
<figureCitation id="13172A67683EFFFC4D106448FCA3FC01" box="[778,862,870,897]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="29" pageNumber="539">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD64A4FE58FB6A" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD64A4FE8BFC25" box="[199,374,907,933]" italics="true" pageId="29" pageNumber="539">Male genitalia.</emphasis>
Genital capsule (
<figureCitation id="13172A67683EFFFC4C5964A4FD3AFC25" box="[579,711,907,933]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Figs 90–94</figureCitation>
) nearly quadrangular in dorsal/ventral view, longer than wide. Ventral rim deeply incised medially, sharp (
<figureCitation id="13172A67683EFFFC4C886480FD15FC49" box="[658,744,943,969]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Fig. 93</figureCitation>
); posterolateral projections prominent (
<figureCitation id="13172A67683EFFFC4AB46480FACAFC49" box="[1198,1335,943,969]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Figs 93, 94</figureCitation>
); ventral rim infolding not visible (
<figureCitation id="13172A67683EFFFC4FDB64FCFDEAFC6D" box="[449,535,979,1005]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Fig. 91</figureCitation>
). Superior processes of dorsal rim elongate, crescent-shaped (
<figureCitation id="13172A67683EFFFC4AC464FCFA6DFC6E" box="[1246,1424,979,1006]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Figs 90–91, 96</figureCitation>
). Proctiger trapezoid, short, without any constriction (
<figureCitation id="13172A67683EFFFC4CFC64D8FCC6FB91" box="[742,827,1014,1041]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Fig. 95</figureCitation>
). Parameral socket open (
<figureCitation id="13172A67683EFFFC4A4564D8FB49FB91" box="[1119,1204,1015,1041]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="29" pageNumber="539">Fig. 91</figureCitation>
: ps). Paramere (
<figureCitation id="13172A67683EFFFC4B7064D8FF0DFBB5" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="29" pageNumber="539">Figs 97–101</figureCitation>
): basal process (bp) with a lamellate, horizontally oriented semicircular lobe, apically with group of few long setae (
<figureCitation id="13172A67683EFFFC4F066310FE4FFBD9" box="[284,434,1087,1113]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="29" pageNumber="539">Figs 97, 101</figureCitation>
); apical process (ap) hook-shaped, apically more pointed (
<figureCitation id="13172A67683EFFFC4A4D6310FB21FBD9" box="[1111,1244,1087,1113]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="29" pageNumber="539">Figs 97–99</figureCitation>
); mesal angle of stem without any additional projection (
<figureCitation id="13172A67683EFFFC4C78634CFCDFFBFD" box="[610,802,1123,1149]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="29" pageNumber="539">Figs 97, 99, 101</figureCitation>
). Phallus (
<figureCitation id="13172A67683EFFFC4D87634CFBC2FBFD" box="[925,1087,1122,1149]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="29" pageNumber="539">Figs 102–105</figureCitation>
): A pair of dorsal conjunctival processes (dlc), transversely placed and ending in triangular sclerotised processes (
<figureCitation id="13172A67683EFFFC4A2763A8FB5DFB21" box="[1085,1184,1158,1185]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="29" pageNumber="539">Fig. 105</figureCitation>
); median penial plates (mpp) or processes of aedeagus fused basally, outer margins broadly and thickly sclerotized, encircling short, curved aedeagus (
<figureCitation id="13172A67683EFFFC4F1763E0FE92FB69" box="[269,367,1230,1257]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="29" pageNumber="539">Fig. 105</figureCitation>
: ad).
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD63DCFD6EF9AD" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD63DCFE74FA8D" box="[199,393,1267,1293]" italics="true" pageId="29" pageNumber="539">Female genitalia</emphasis>
(
<figureCitation id="13172A67683EFFFC4F8363DCFE0EFA8D" box="[409,499,1267,1293]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="29" pageNumber="539">Figs 32</figureCitation>
, 
<figureCitation id="13172A67683EFFFC4C1B63DCFDDCFA8E" box="[513,545,1267,1294]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">36</figureCitation>
). Laterotergite VIII (lt8) subtriangular. Laterotergites IX (lt9) widely rounded apically, not touching each other medially, segment X visible (
<figureCitation id="13172A67683EFFFC4D436238FC52FAB1" box="[857,943,1303,1329]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="29" pageNumber="539">Fig. 32</figureCitation>
). Valvifers VIII (vf8) quadrant with median margins juxtaposed and dorsal margins slightly concave (
<figureCitation id="13172A67683EFFFC4D0C6214FC97FAD5" box="[790,874,1339,1365]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="29" pageNumber="539">Fig. 32</figureCitation>
). Valvifers IX (vf9) fused medially, with indistinct median fusion line, flat. Ring sclerites roughly circular, not apparently sclerotized in the specimen examined (
<figureCitation id="13172A67683EFFFC4B756270FF4AFA1E" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">Fig. 36</figureCitation>
). Spermatheca: dilation in the examined specimen swollen, whitish and not transparent, constricted subbasaly with sclerotized ring (
<figureCitation id="13172A67683EFFFC4F946288FE1CFA41" box="[398,481,1447,1473]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">Fig. 33</figureCitation>
), distal invagination (‘sclerotized rod’; 
<figureCitation id="13172A67683EFFFC4D846288FC12FA41" box="[926,1007,1447,1473]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">Fig. 33</figureCitation>
) visible only basally, strongly curved; distal flange (df) tightly attached to apical receptacle (ar) (
<figureCitation id="13172A67683EFFFC4D2D62E4FC73FA66" box="[823,910,1483,1510]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">Fig. 36</figureCitation>
); apical receptacle subhemispherical (
<figureCitation id="13172A67683EFFFC4B5A62E4FA69FA66" box="[1344,1428,1483,1510]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="29" pageNumber="539">Fig. 36</figureCitation>
) with one long projection directed proximad and nearly reaching proximal flange (pf), and one short processes directed proximad and reaching distal flange.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD6118FE38F93D" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD6118FE94F9D0" box="[199,361,1591,1616]" italics="true" pageId="29" pageNumber="539">Measurements</emphasis>
(mm). See 
<tableCitation id="C6AE0359683EFFFC4FF66118FDBCF9D1" box="[492,577,1591,1617]" captionStart="TABLE 2" captionStartId="34.[151,239,507,533]" captionText="TABLE 2. Measurements of the Agathocles species presented as median, with minimum and maximum values given in parentheses." pageId="29" pageNumber="539">Table 2</tableCitation>
. 
<typeStatus id="54978840683EFFFC4C566118FD49F9D1" box="[588,692,1591,1617]" pageId="29" pageNumber="539" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D2AFD6B683EFFFC4CDF6118FD20F9D1" box="[709,733,1591,1617]" pageId="29" pageNumber="539" type="male">♂</specimenCount>
): Body length 19.27; head: length 3.43, width (including eyes) 3.92, interocular width 2.45; lengths of antennomeres: I—1.47, IIa—2.35, IIb—2.94, III—2.75, IV—2.45; pronotum: length 4.51, width (across humeri) 10.39; scutellum: length 6.96, width (across basal angles) 5.88; abdomen width (across segment III) 10.69.
</paragraph>
<paragraph id="8B9336E2683EFFFC4EDD61E9FB21F8CE" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD61E9FEC2F960" bold="true" box="[199,319,1734,1760]" pageId="29" pageNumber="539">Variation.</emphasis>
<materialsCitation id="3B443CBF683EFFFC4F5F61E8FEE5F8AA" ID-GBIF-Occurrence="3083795358" collectionCode="IV, III" pageId="29" pageNumber="539" specimenCount="3">
Distribution of small yellowish callosities on dorsal surface of body varies among specimens. Ratios of length of antennomeres IIa to 
<collectionCode id="ED3DAE27683EFFFC4C1561C4FDD1F885" box="[527,556,1771,1797]" pageId="29" pageNumber="539">IV</collectionCode>
highly variable, arranged from shortest to longest: I &lt;IIa &lt;
<collectionCode id="ED3DAE27683EFFFC4AFB61C4FB03F885" box="[1249,1278,1771,1797]" pageId="29" pageNumber="539">IV</collectionCode>
&lt;
<collectionCode id="ED3DAE27683EFFFC4B0561C4FAC6F885" box="[1311,1339,1771,1797]" pageId="29" pageNumber="539">III</collectionCode>
&lt;IIb (
<specimenCount id="9D2AFD6B683EFFFC4B9461C4FEEDF8AA" pageId="29" pageNumber="539" type="generic">3 specimens</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="3B443CBF683EFFFC4F396020FCF4F8AA" ID-GBIF-Occurrence="3083795329" box="[291,777,1807,1834]" collectionCode="III, IV" pageId="29" pageNumber="539" specimenCount="1">
I &lt;IIa = IIb &lt;
<collectionCode id="ED3DAE27683EFFFC4FDD6020FE1EF8A9" box="[455,483,1807,1833]" pageId="29" pageNumber="539">III</collectionCode>
(
<collectionCode id="ED3DAE27683EFFFC4FEB6020FDEDF8A9" box="[497,528,1807,1833]" pageId="29" pageNumber="539">IV</collectionCode>
missing; 
<specimenCount id="9D2AFD6B683EFFFC4C676020FCFDF8AA" box="[637,768,1807,1834]" pageId="29" pageNumber="539" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="3B443CBF683EFFFC4D0E6020FB04F8AA" ID-GBIF-Occurrence="3083795352" box="[788,1273,1807,1834]" collectionCode="III, IV" pageId="29" pageNumber="539" specimenCount="1">
I &lt;IIa &lt;IIb &lt;
<collectionCode id="ED3DAE27683EFFFC4DAD6020FC2EF8A9" box="[951,979,1807,1833]" pageId="29" pageNumber="539">III</collectionCode>
(
<collectionCode id="ED3DAE27683EFFFC4DF86020FBFCF8A9" box="[994,1025,1807,1833]" pageId="29" pageNumber="539">IV</collectionCode>
missing; 
<specimenCount id="9D2AFD6B683EFFFC4A776020FB0CF8AA" box="[1133,1265,1807,1834]" pageId="29" pageNumber="539" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="3B443CBF683EFFFC4B1E6020FE65F8CE" ID-GBIF-Occurrence="3083795315" collectionCode="IV, III" pageId="29" pageNumber="539" specimenCount="2">
I &lt;IIa &lt;
<collectionCode id="ED3DAE27683EFFFC4B726020FA78F8A9" box="[1384,1413,1807,1833]" pageId="29" pageNumber="539">IV</collectionCode>
&lt;IIb &lt;
<collectionCode id="ED3DAE27683EFFFC4ECC601CFF0FF8CD" box="[214,242,1843,1869]" pageId="29" pageNumber="539">III</collectionCode>
(
<specimenCount id="9D2AFD6B683EFFFC4F1B601CFE6DF8CE" box="[257,400,1843,1870]" pageId="29" pageNumber="539" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="3B443CBF683EFFFC4FB9601CFCC5F8CE" ID-GBIF-Occurrence="3083795353" box="[419,824,1843,1870]" collectionCode="IV, III" pageId="29" pageNumber="539" specimenCount="2">
I &lt;IIa &lt;
<collectionCode id="ED3DAE27683EFFFC4C1D601CFDD9F8CD" box="[519,548,1843,1869]" pageId="29" pageNumber="539">IV</collectionCode>
&lt;IIb = 
<collectionCode id="ED3DAE27683EFFFC4C9A601CFD61F8CD" box="[640,668,1843,1869]" pageId="29" pageNumber="539">III</collectionCode>
(
<specimenCount id="9D2AFD6B683EFFFC4CB6601CFCD2F8CE" box="[684,815,1843,1870]" pageId="29" pageNumber="539" type="generic">2 specimen</specimenCount>
)
</materialsCitation>
, 
<materialsCitation id="3B443CBF683EFFFC4D58601CFB25F8CE" ID-GBIF-Occurrence="3083795345" box="[834,1240,1843,1870]" collectionCode="IV, III" pageId="29" pageNumber="539" specimenCount="1">
I &lt;
<collectionCode id="ED3DAE27683EFFFC4D70601CFC7AF8CD" box="[874,903,1843,1869]" pageId="29" pageNumber="539">IV</collectionCode>
&lt;IIa &lt;IIb = 
<collectionCode id="ED3DAE27683EFFFC4A3A601CFBC1F8CD" box="[1056,1084,1843,1869]" pageId="29" pageNumber="539">III</collectionCode>
(
<specimenCount id="9D2AFD6B683EFFFC4A54601CFB2DF8CE" box="[1102,1232,1843,1870]" pageId="29" pageNumber="539" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3366569683EFFFC4EDD6079FE9BF839" pageId="29" pageNumber="539" type="diagnosis">
<paragraph id="8B9336E2683EFFFC4EDD6079FE9BF839" blockId="29.[151,1437,150,1977]" pageId="29" pageNumber="539">
<emphasis id="B958EAF0683EFFFC4EDD6079FE33F8F0" bold="true" box="[199,462,1878,1904]" pageId="29" pageNumber="539">Differential diagnosis.</emphasis>
See key and 
<tableCitation id="C6AE0359683EFFFC4C7F6078FD47F8F1" box="[613,698,1879,1905]" captionStart="TABLE 3" captionStartId="40.[151,239,479,505]" captionText="TABLE 3. Summary of main morphological differences of Agathocles species." pageId="29" pageNumber="539">Table 3</tableCitation>
. This species is easy to recognize due to its apically free clypeus, scape surpassing apex of head, and blackish-brown colouration with conspicuous whitish unpunctured spot at the apex of scutellum.
</paragraph>
</subSubSection>
<caption id="DF53666A683DFFFF4E8D615BFC21F955" ID-DOI="http://doi.org/10.5281/zenodo.4692411" ID-Zenodo-Dep="4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="30" pageNumber="540" startId="30.[151,264,1652,1676]" targetBox="[153,1428,193,1628]" targetPageId="30">
<paragraph id="8B9336E2683DFFFF4E8D615BFC21F955" blockId="30.[151,1436,1652,1749]" pageId="30" pageNumber="540">
<emphasis id="B958EAF0683DFFFF4E8D615BFE90F90C" bold="true" box="[151,365,1652,1676]" pageId="30" pageNumber="540">FIGURES 108–112.</emphasis>
<taxonomicName id="4C2C4D61683DFFFF4F6E615BFD17F90D" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[372,746,1652,1677]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="540" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0683DFFFF4F6E615BFDBBF90C" box="[372,582,1652,1676]" italics="true" pageId="30" pageNumber="540">Agathocles normalis</emphasis>
(Distant, 1901)
</taxonomicName>
, habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of 
<taxonomicName id="4C2C4D61683DFFFF4D5261B6FB75F931" authority="Distant, 1901" authorityName="Distant. We" authorityYear="1901" box="[840,1160,1688,1713]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="30" pageNumber="540" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0683DFFFF4D5261B6FC05F930" box="[840,1016,1688,1712]" italics="true" pageId="30" pageNumber="540">Surenus normalis</emphasis>
Distant, 1901
</taxonomicName>
, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112).
</paragraph>
</caption>
<caption id="DF53666A683CFFFE4E8D64AFFE84FB84" ID-DOI="http://doi.org/10.5281/zenodo.4692413" ID-Zenodo-Dep="4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="31" pageNumber="541" startId="31.[151,264,896,920]" targetBox="[306,1282,181,873]" targetPageId="31">
<paragraph id="8B9336E2683CFFFE4E8D64AFFE84FB84" blockId="31.[151,1437,896,1029]" pageId="31" pageNumber="541">
<emphasis id="B958EAF0683CFFFE4E8D64AFFE94FC19" bold="true" box="[151,361,896,921]" pageId="31" pageNumber="541">FIGURES 113–117.</emphasis>
Male genital capsule of 
<taxonomicName id="4C2C4D61683CFFFE4C7B64AEFC29FC19" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[609,980,897,921]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="541" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0683CFFFE4C7B64AEFCCCFC19" box="[609,817,897,921]" italics="true" pageId="31" pageNumber="541">Agathocles normalis</emphasis>
(Distant, 1901)
</taxonomicName>
: 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment.
</paragraph>
</caption>
<caption id="DF53666A683CFFFE4E8D60ABFDADF864" ID-DOI="http://doi.org/10.5281/zenodo.4692415" ID-Zenodo-Dep="4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="31" pageNumber="541" startId="31.[151,264,1924,1948]" targetBox="[311,1277,1053,1901]" targetPageId="31">
<paragraph id="8B9336E2683CFFFE4E8D60ABFDADF864" blockId="31.[151,1436,1924,2020]" pageId="31" pageNumber="541">
<emphasis id="B958EAF0683CFFFE4E8D60ABFE92F81C" bold="true" box="[151,367,1924,1948]" pageId="31" pageNumber="541">FIGURES 118–124.</emphasis>
Paramere of 
<taxonomicName id="4C2C4D61683CFFFE4C1A60ABFC80F81C" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[512,893,1924,1948]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="541" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0683CFFFE4C1A60ABFD28F81C" box="[512,725,1924,1948]" italics="true" pageId="31" pageNumber="541">Agathocles normalis</emphasis>
(Distant, 1901)
</taxonomicName>
, five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment.
</paragraph>
</caption>
<subSubSection id="C33665696803FFC14EDD67B8FC6BFF7A" pageId="32" pageNumber="542" type="etymology">
<paragraph id="8B9336E26803FFC14EDD67B8FC6BFF7A" blockId="32.[151,1436,151,250]" pageId="32" pageNumber="542">
<emphasis id="B958EAF06803FFC14EDD67B8FEB3FF31" bold="true" box="[199,334,151,177]" pageId="32" pageNumber="542">Etymology.</emphasis>
We are pleased to dedicate this elegant new species to our dear colleague, Professor Jocelia Grazia, with our thanks for her outstanding contributions to pentatomoid systematics, morphology and phylogeny. 
<emphasis id="B958EAF06803FFC14EDD67F1FEA4FF78" bold="true" box="[199,345,222,248]" pageId="32" pageNumber="542">Distribution</emphasis>
(
<figureCitation id="13172A676803FFC14F7367F0FE36FF79" box="[361,459,223,249]" captionStart="FIGURE 139" captionStartId="39.[151,250,1769,1793]" captionTargetBox="[151,1435,1007,1745]" captionTargetId="figure-340@39.[151,1435,1007,1745]" captionTargetPageId="39" captionText="FIGURE 139. Distribution of the Agathocles species. T—type locality;?—doubtful record. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692423" httpUri="https://zenodo.org/record/4692423/files/figure.png" pageId="32" pageNumber="542">Fig. 139</figureCitation>
). Peninsular 
<collectingCountry id="F33B76726803FFC14C4667F0FD3EFF79" box="[604,707,223,249]" name="Malaysia" pageId="32" pageNumber="542">Malaysia</collectingCountry>
(
<collectingRegion id="49E8F8006803FFC14CCB67F0FCC6FF79" box="[721,827,223,249]" country="Malaysia" name="Kelantan" pageId="32" pageNumber="542">Kelantan</collectingRegion>
, 
<collectingRegion id="49E8F8006803FFC14D5C67F0FC76FF7A" box="[838,907,223,250]" country="Malaysia" name="Perak" pageId="32" pageNumber="542">Perak</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>