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<mods:title id="A6D24B2C08715AF7EF13340F9433D057">The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species</mods:title>
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<mods:namePart id="426CE0A962E0D98ED6C3ABA74E32EBD0">Salini, S.</mods:namePart>
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<mods:namePart id="801E7B2CEF0E8585DAC1F43B7DFF87FF">Kment, Petr</mods:namePart>
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<mods:affiliation id="70D6B90A1B54B70988CD5A55D858C399">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<taxonomicName id="4C2C4D616826FFE44E8D6311FE62FBD8" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[151,415,1085,1113]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EAF06826FFE44E8D6311FEE0FBD9" bold="true" box="[151,285,1086,1113]" italics="true" pageId="5" pageNumber="515">Agathocles</emphasis>
<bibRefCitation id="EFBD4B136826FFE44F3E6312FE62FBD8" author="Stal, C." box="[292,415,1085,1112]" pageId="5" pageNumber="515" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
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<taxonomicName id="4C2C4D616826FFE44E8D63ACFE60FB1B" authority="Stal, 1876: 69" authorityName="Stal" authorityPageNumber="69" authorityYear="1876" box="[151,413,1155,1179]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06826FFE44E8D63ACFEF5FB1B" box="[151,264,1155,1179]" italics="true" pageId="5" pageNumber="515">Agathocles</emphasis>
<bibRefCitation id="EFBD4B136826FFE44F1563ACFE60FB1B" author="Stal, C." box="[271,413,1155,1179]" pageId="5" pageNumber="515" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876: 69</bibRefCitation>
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(key to genera), 106 (description of 
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species). 
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species: 
<taxonomicName id="4C2C4D616826FFE44A2263ACFA85FB1B" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[1080,1400,1155,1179]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF06826FFE44A2263ACFAFAFB1B" box="[1080,1287,1155,1179]" italics="true" pageId="5" pageNumber="515">Agathocles limbatus</emphasis>
<bibRefCitation id="EFBD4B136826FFE44B1463ACFA85FB1B" author="Stal, C." box="[1294,1400,1155,1179]" pageId="5" pageNumber="515" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
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, by monotypy.
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<taxonomicName id="4C2C4D616826FFE44E8D63EFFE46FB59" authority=": Atkinson (1888)" authorityName=": Atkinson" authorityYear="1888" box="[151,443,1216,1241]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06826FFE44E8D63EFFEF5FB58" box="[151,264,1216,1240]" italics="true" pageId="5" pageNumber="515">Agathocles</emphasis>
: 
<bibRefCitation id="EFBD4B136826FFE44F0863EFFE46FB59" author="Atkinson, E. T." box="[274,443,1216,1241]" pageId="5" pageNumber="515" pagination="118 - 184" refId="ref28739" refString="Atkinson, E. T. (1888) Notes on Indian Rhynchota, Heteroptera. No. 3 &amp; 4. Journal of the Asiatic Society of Bengal, 57, 118 - 184." type="journal article" year="1888">Atkinson (1888)</bibRefCitation>
</taxonomicName>
: 152
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(redescription); 
<treatmentCitation id="0A8D10F36826FFE44C9563EFFC2EFB59" author="Lethierry, L. &amp; Severin, G." box="[655,979,1216,1241]" page="183" pageId="5" pageNumber="515" year="1893">
<bibRefCitation id="EFBD4B136826FFE44C9563EFFC58FB59" author="Lethierry, L. &amp; Severin, G." box="[655,933,1216,1241]" pageId="5" pageNumber="515" refId="ref31678" refString="Lethierry, L. &amp; Severin, G. (1893) Catalogue general des Hemipteres. Vol. 1. Pentatomidae. Bruxelles, x + 286 pp. https: // doi. org / 10.5962 / bhl. title. 15830" type="book" year="1893">Lethierry &amp; Severin (1893)</bibRefCitation>
: 183
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(catalogue); 
<treatmentCitation id="0A8D10F36826FFE44A4F63EFFAE4FB59" author="Distant, W. L." box="[1109,1305,1216,1241]" page="173" pageId="5" pageNumber="515" year="1900">
<bibRefCitation id="EFBD4B136826FFE44A4F63EFFB16FB59" author="Distant, W. L." box="[1109,1259,1216,1241]" pageId="5" pageNumber="515" pagination="163 - 174" refId="ref29399" refString="Distant, W. L. (1900) Undescribed genera and species belonging to the rhynchotal family Pentatomidae. Transactions of the Entomological Society of London, 1900 (1), 163 - 174, pl. II. https: // doi. org / 10.1111 / j. 1365 - 2311.1900. tb 00998. x" type="journal article" year="1900">Distant (1900)</bibRefCitation>
: 173
</treatmentCitation>
(comparison with 
<emphasis id="B958EAF06826FFE44EE163F0FEA1FB77" box="[251,348,1247,1271]" italics="true" pageId="5" pageNumber="515">Fernelius</emphasis>
); 
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<bibRefCitation id="EFBD4B136826FFE44F6963F0FDF7FB77" author="Distant, W. L." box="[371,522,1247,1271]" pageId="5" pageNumber="515" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant (1902)</bibRefCitation>
: 196
</treatmentCitation>
(key to genera), 198 (redescription); 
<treatmentCitation id="0A8D10F36826FFE44DA363F0FB6FFB77" author="Kirkaldy, G. W." box="[953,1170,1247,1271]" page="147" pageId="5" pageNumber="515" year="1909">
<bibRefCitation id="EFBD4B136826FFE44DA363F0FB9FFB77" author="Kirkaldy, G. W." box="[953,1122,1247,1271]" pageId="5" pageNumber="515" refId="ref31156" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 147
</treatmentCitation>
(catalogue); Ahmad 
<emphasis id="B958EAF06826FFE44B7363CFFA60FB77" box="[1385,1437,1247,1271]" italics="true" pageId="5" pageNumber="515">et al.</emphasis>
(1974): 88, 89 (figure legend); 
<treatmentCitation id="0A8D10F36826FFE44C1263D1FD3BFA96" author="Ahmad, I." box="[520,710,1278,1302]" page="16" pageId="5" pageNumber="515" year="1981">
<bibRefCitation id="EFBD4B136826FFE44C1263D1FD5FFA96" author="Ahmad, I." box="[520,674,1278,1302]" pageId="5" pageNumber="515" pagination="1 - 113" refId="ref27623" refString="Ahmad, I. (1981) A revision of the superfamilies Coreoidea and Pentatomoidea (Heteroptera: Pentatomomorpha) from Pakistan, Azad Kashmir and Bangladesh. Part 1: Additions and corrections of coreid and pentatomid fauna with phylogenetic considerations. Supplement of the Entomological Society of Karachi, 4 (1), 1 - 113. [1979]" type="journal article" year="1981">Ahmad (1981)</bibRefCitation>
: 16
</treatmentCitation>
(key to genera), 43 (list); 
<treatmentCitation id="0A8D10F36826FFE44DC263D1FB6DFA96" author="Rider, D. A." box="[984,1168,1278,1302]" page="402" pageId="5" pageNumber="515" year="1997">
<bibRefCitation id="EFBD4B136826FFE44DC263D1FBA2FA96" author="Rider, D. A." box="[984,1119,1278,1302]" pageId="5" pageNumber="515" pagination="401 - 403" refId="ref32620" refString="Rider, D. A. (1997) Rolstoniellini, replacement name proposed for Compastini Distant, 1902. A tribal name based on a generic junior homonym (Heteroptera: Pentatomidae: Pentatominae). Journal of New York Entomological Society, 103 (4), 401 - 403. [1995]" type="journal article" year="1997">Rider (1997)</bibRefCitation>
: 402
</treatmentCitation>
(checklist); 
<treatmentCitation id="0A8D10F36826FFE44B0863D1FF10FAB5" author="Rider, D. A." page="342" pageId="5" pageNumber="515" year="2006">
<bibRefCitation id="EFBD4B136826FFE44B0863D1FA64FA96" author="Rider, D. A." box="[1298,1433,1278,1302]" pageId="5" pageNumber="515" pagination="1815" refId="ref32676" refString="Rider, D. A. (2006) Family Pentatomidae Leach, 1815. In: Aukema, B. &amp; Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. Pentatomomorpha II. The Netherlands Entomological Society, Amsterdam, pp. 233 - 402." type="book chapter" year="2006">Rider (2006)</bibRefCitation>
: 342
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0A8D10F36826FFE44F6D6232FDECFAB4" author="Fan, Z. - H." box="[375,529,1308,1333]" page="15" pageId="5" pageNumber="515" year="2011">
<bibRefCitation id="EFBD4B136826FFE44F6D6232FE11FAB5" author="Fan, Z. - H." box="[375,492,1309,1333]" pageId="5" pageNumber="515" refId="ref29913" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Unpublished Ph. D., Nankai University, Tianjin, 614 pp." type="book" year="2011">Fan (2011)</bibRefCitation>
: 15
</treatmentCitation>
(key to genera), 43 (diagnosis, key to Chinese species); 
<treatmentCitation id="0A8D10F36826FFE44A7E6233FAACFAB5" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1124,1361,1308,1333]" page="80" pageId="5" pageNumber="515" year="2018">
<bibRefCitation id="EFBD4B136826FFE44A7E6233FAD4FAB5" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[1124,1321,1308,1333]" pageId="5" pageNumber="515" pagination="25 - 201" refId="ref32736" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): Biology, higher systematics, semiochemistry, and management. CRC Press, Taylor &amp; Francis Group, Boca Raton, London and New York, pp. 25 - 201. https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider 
<emphasis id="B958EAF06826FFE44ABE6232FB27FAB4" box="[1188,1242,1308,1332]" italics="true" pageId="5" pageNumber="515">et al.</emphasis>
(2018)
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: 80
</treatmentCitation>
, 96–97 (comments on tribal placement).
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</paragraph>
<paragraph id="8B9336E26826FFE44E8D6275FC89FAF2" blockId="5.[151,1437,1155,1609]" box="[151,884,1370,1394]" pageId="5" pageNumber="515">
<emphasis id="B958EAF06826FFE44E8D6275FEF4FAF2" box="[151,265,1370,1394]" italics="true" pageId="5" pageNumber="515">Agathooles</emphasis>
(incorrect subsequent spelling): 
<treatmentCitation id="0A8D10F36826FFE44C4F6275FD06FAF2" box="[597,763,1370,1394]" pageId="5" pageNumber="515">
<bibRefCitation id="EFBD4B136826FFE44C4F6275FD25FAF2" author="Jiang, Z. - L." box="[597,728,1370,1394]" pageId="5" pageNumber="515" pagination="54 - 67" refId="ref31067" refString="Jiang, Z. - L. (1985) [The Hemiptera of Yunnan - A list of the family Pentatomidae]. Yunnan Forestry Science and Technology, 1985 (2), 54 - 67. [in Chinese]" type="journal article" year="1985">Jiang (1985)</bibRefCitation>
: 67
</treatmentCitation>
(checklist).
</paragraph>
<paragraph id="8B9336E26826FFE44E8D6257FC09FA10" blockId="5.[151,1437,1155,1609]" box="[151,1012,1400,1425]" pageId="5" pageNumber="515">
<emphasis id="B958EAF06826FFE44E8D6257FEF5FA10" box="[151,264,1400,1424]" italics="true" pageId="5" pageNumber="515">Agothocles</emphasis>
(incorrect subsequent spelling): Ahmad 
<emphasis id="B958EAF06826FFE44CB96256FD2AFA10" box="[675,727,1400,1424]" italics="true" pageId="5" pageNumber="515">et al.</emphasis>
(1974): 43 (list, host plant).
</paragraph>
<paragraph id="8B9336E26826FFE44E8D62B8FD01FA4E" blockId="5.[151,1437,1155,1609]" pageId="5" pageNumber="515">
<treatmentCitationGroup id="AB3C11CC6826FFE44E8D62B8FD01FA4E" pageId="5" pageNumber="515">
<treatmentCitation id="0A8D10F36826FFE44E8D62B8FE48FA2F" author="Distant, W. L." box="[151,437,1431,1455]" page="101" pageId="5" pageNumber="515" year="1901">
<taxonomicName id="4C2C4D616826FFE44E8D62B8FE48FA2F" authority="Distant, 1901 a: 101" authorityName="Distant" authorityPageNumber="101" authorityYear="1901" box="[151,437,1431,1455]" class="Insecta" family="Pentatomidae" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06826FFE44E8D62B8FF15FA2E" box="[151,232,1431,1454]" italics="true" pageId="5" pageNumber="515">Surenus</emphasis>
<bibRefCitation id="EFBD4B136826FFE44EF462B8FE48FA2F" author="Distant, W. L." box="[238,437,1431,1455]" pageId="5" pageNumber="515" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901a: 101</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(list), 106–107 (original description, systematic position). 
<typeStatus id="549788406826FFE44A1D62B7FBC4FA30" box="[1031,1081,1432,1456]" pageId="5" pageNumber="515">Type</typeStatus>
species: 
<taxonomicName id="4C2C4D616826FFE44A8C62B8FEFAFA4E" authority="Distant, 1901 a" authorityName="Distant. We" authorityYear="1901" class="Insecta" family="Pentatomidae" genus="Surenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF06826FFE44A8C62B8FABAFA2F" box="[1174,1351,1431,1455]" italics="true" pageId="5" pageNumber="515">Surenus normalis</emphasis>
<bibRefCitation id="EFBD4B136826FFE44B5762B8FEFAFA4E" author="Distant, W. L." pageId="5" pageNumber="515" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901a</bibRefCitation>
</taxonomicName>
, by monotypy. 
<emphasis id="B958EAF06826FFE44FB96299FD01FA4E" bold="true" box="[419,764,1462,1486]" pageId="5" pageNumber="515">New junior subjective synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9336E26826FFE44E8D62FAFAFFF9C8" blockId="5.[151,1437,1155,1609]" pageId="5" pageNumber="515">
<treatmentCitationGroup id="AB3C11CC6826FFE44E8D62FAFAFFF9C8" pageId="5" pageNumber="515">
<treatmentCitation id="0A8D10F36826FFE44E8D62FAFE3BFA6D" author="Distant, W. L." box="[151,454,1492,1517]" page="110" pageId="5" pageNumber="515" year="1902">
<taxonomicName id="4C2C4D616826FFE44E8D62FAFE69FA6D" authority=": Distant (1902)" authorityName=": Distant" authorityYear="1902" box="[151,404,1492,1517]" class="Insecta" family="Pentatomidae" genus="Surenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06826FFE44E8D62FAFF15FA6C" box="[151,232,1493,1516]" italics="true" pageId="5" pageNumber="515">Surenus</emphasis>
: 
<bibRefCitation id="EFBD4B136826FFE44EE362FBFE69FA6D" author="Distant, W. L." box="[249,404,1492,1517]" pageId="5" pageNumber="515" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant (1902)</bibRefCitation>
</taxonomicName>
: 110
</treatmentCitation>
(key to genera), 116–117 (description repeated); 
<treatmentCitation id="0A8D10F36826FFE44DCC62FBFB48FA6C" author="Bergroth, E." box="[982,1205,1492,1517]" page="154" pageId="5" pageNumber="515" year="1908">
<bibRefCitation id="EFBD4B136826FFE44DCC62FBFB7CFA6D" author="Bergroth, E." box="[982,1153,1492,1517]" pageId="5" pageNumber="515" pagination="131 - 200" refId="ref28911" refString="Bergroth, E. (1908) Enumeratio Pentatomidarum post Catalogum bruxellensem descriptarum. Memoires de la Societe Entomologique de Belgique, 15, 131 - 200." type="journal article" year="1908">Bergroth (1908)</bibRefCitation>
: 154
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0A8D10F36826FFE44B5862FBFEBDF98B" author="Kirkaldy, G. W." page="195" pageId="5" pageNumber="515" year="1909">
<bibRefCitation id="EFBD4B136826FFE44B5862FBFEF2F98B" author="Kirkaldy, G. W." pageId="5" pageNumber="515" refId="ref31156" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Prefaced by a discussion on Nomenclature and an analytical table of families. Vol. I. Cimicidae. Felix L. Dames, Berlin, xl + 392 pp. https: // doi. org / 10.5962 / bhl. title. 15205" type="book" year="1909">Kirkaldy (1909)</bibRefCitation>
: 195
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0A8D10F36826FFE44FD362DBFD1CF98B" author="Memon, N. &amp; Gilbert, F. &amp; Ahmad, I." box="[457,737,1523,1547]" page="1151" pageId="5" pageNumber="515" year="2011">
<bibRefCitation id="EFBD4B136826FFE44FD362DBFD59F98B" author="Memon, N. &amp; Gilbert, F. &amp; Ahmad, I." box="[457,676,1523,1547]" pageId="5" pageNumber="515" pagination="1149 - 1169" refId="ref32198" refString="Memon, N., Gilbert, F. &amp; Ahmad, I. (2011) Phylogeny of the South Asian Halyine stink bugs (Hemiptera: Pentatomidae: Halyini) based on morphological characters. Annals of Entomological Society of America, 104 (6), 1149 - 1169. https: // doi. org / 10.1603 / AN 10109" type="journal article" year="2011">
Memon 
<emphasis id="B958EAF06826FFE44C3A62DBFDA8F98B" box="[544,597,1523,1547]" italics="true" pageId="5" pageNumber="515">et al.</emphasis>
(2011)
</bibRefCitation>
: 1151
</treatmentCitation>
–1155, 1157, 1162–1163 (phylogenetic analysis, key to the genera); 
<treatmentCitation id="0A8D10F36826FFE44EDD613DFDF3F9AA" author="Salini, S. &amp; Viraktamath, C. A." box="[199,526,1554,1578]" page="10" pageId="5" pageNumber="515" year="2015">
<bibRefCitation id="EFBD4B136826FFE44EDD613DFE16F9AA" author="Salini, S. &amp; Viraktamath, C. A." box="[199,491,1554,1578]" pageId="5" pageNumber="515" pagination="1 - 76" refId="ref33142" refString="Salini, S. &amp; Viraktamath, C. A. (2015) Genera of Pentatomidae (Hemiptera: Pentatomoidea) from south India - an illustrated key to genera and checklist of species. Zootaxa, 3924 (1), 1 - 76. https: // doi. org / 10.11646 / zootaxa. 3924.1.1" type="journal article" year="2015">Salini &amp; Viraktamath (2015)</bibRefCitation>
: 10
</treatmentCitation>
, 22, 51, 59 (key to tribes and genera, checklist for south Indian 
<taxonomicName id="4C2C4D616826FFE44A93613DFAEBF9AA" ID-CoL="62662" authorityName="Leach" authorityYear="1815" box="[1161,1302,1554,1578]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="515" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
, illustration); 
<treatmentCitation id="0A8D10F36826FFE44EDD611FFE1BF9C9" author="Arnold, K. &amp; Arnold, M. - L." box="[199,486,1584,1609]" page="13" pageId="5" pageNumber="515" year="2017">
<bibRefCitation id="EFBD4B136826FFE44EDD611FFE39F9C9" author="Arnold, K. &amp; Arnold, M. - L." box="[199,452,1584,1609]" pageId="5" pageNumber="515" pagination="9 - 21" refId="ref28681" refString="Arnold, K. &amp; Arnold, M. - L. (2017) 7. Beitrag zur Heteropterenfauna der Inselwelt Indonesiens, speziell der Insel Sumatra so wie weiterer ostlicher bzw. sudostlicher Bereiche unter Einbeziehung der sudostlichen Orientalischen Region (Insecta: Hemiptera: Heteroptera). Edessana, 7, 9 - 21." type="journal article" year="2017">Arnold &amp; Arnold (2017)</bibRefCitation>
: 13
</treatmentCitation>
(faunistic record); 
<treatmentCitation id="0A8D10F36826FFE44CB3611FFC73F9C9" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[681,910,1584,1609]" page="80" pageId="5" pageNumber="515" year="2018">
<bibRefCitation id="EFBD4B136826FFE44CB3611FFC94F9C9" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[681,873,1584,1609]" pageId="5" pageNumber="515" pagination="25 - 201" refId="ref32736" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): Biology, higher systematics, semiochemistry, and management. CRC Press, Taylor &amp; Francis Group, Boca Raton, London and New York, pp. 25 - 201. https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider 
<emphasis id="B958EAF06826FFE44CF2611EFCE6F9C8" box="[744,795,1584,1608]" italics="true" pageId="5" pageNumber="515">et al.</emphasis>
(2018)
</bibRefCitation>
: 80
</treatmentCitation>
, 86 (comments on tribal placement).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33665696826FFED4E8D6159FED5FE51" lastPageId="12" lastPageNumber="522" pageId="5" pageNumber="515" type="description">
<paragraph id="8B9336E26826FFE44E8D6159FDC0F97D" blockId="5.[151,1437,1654,2041]" pageId="5" pageNumber="515">
<emphasis id="B958EAF06826FFE44E8D6159FE1EF911" bold="true" box="[151,483,1654,1681]" pageId="5" pageNumber="515">
Redescription. 
<emphasis id="B958EAF06826FFE44F4A6159FE21F911" bold="true" box="[336,476,1654,1681]" italics="true" pageId="5" pageNumber="515">Colouration</emphasis>
.
</emphasis>
Body above dark brown to black, underside black, sometimes either dorsally or ventrally with yellow pattern; membrane brownish; compound eyes silver or dark brown; ocelli yellowish or reddish; rostrum yellowish; antennae and legs always paler, yellowish to brown. Abdominal spiracles concolorous with surrounding surface of ventrites.
</paragraph>
<paragraph id="8B9336E26826FFE44EDD6028FD2CF80D" blockId="5.[151,1437,1654,2041]" pageId="5" pageNumber="515">
<emphasis id="B958EAF06826FFE44EDD6028FE0DF8A1" bold="true" box="[199,496,1798,1825]" italics="true" pageId="5" pageNumber="515">Integument and vestiture.</emphasis>
Body above matt or slightly shining with only clavus and corium matt; ventral side shining except of matt evaporatorium. Body above and below covered with dense, coarse punctures; posterior portion of pronotal disc with very coarse punctures; punctures on scutellum partly confluent to short transverse wrinkles; abdominal sternites with finer punctures.
</paragraph>
<paragraph id="8B9336E26826FFE74EDD60B8FC6DFF79" blockId="5.[151,1437,1654,2041]" lastBlockId="6.[151,1437,151,1941]" lastPageId="6" lastPageNumber="516" pageId="5" pageNumber="515">
Body glabrous, only antennomeres I to IV with short, erect, brown setae, antennomeres III and IV in addition with short, semierect, golden setae. Labium with short, sparse golden setae, femora with short sparse brown setae, tibiae with dense, short, erect setae, tarsi dorsally with comparatively long, semierect sparse setae and short, dense mat of golden setae ventrally. Genital capsule along ventral and lateral rim and ventral rim infolding with dense, semierect, golden brown setae (
<figureCitation id="13172A676825FFE74C1C6794FDA3FF55" box="[518,606,187,213]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="6" pageNumber="516">Fig. 37</figureCitation>
). External female genitalia (valvifers VIII on dorsal side and on mesial margins, valvifers IX and laterotergite IX) with sparse golden setae.
</paragraph>
<paragraph id="8B9336E26825FFE74EDD662CFD8FFEC1" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD662CFE80FE9D" box="[199,381,259,285]" italics="true" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD662CFEC6FE9D" bold="true" box="[199,315,259,285]" italics="true" pageId="6" pageNumber="516">Structure.</emphasis>
Body
</emphasis>
robust, 
<quantity id="4CD49B076825FFE74FC2662CFDACFE9D" box="[472,593,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.4" metricValueMin="1.3" pageId="6" pageNumber="516" unit="mm" value="18.5" valueMax="24.0" valueMin="13.0">13–24 mm</quantity>
long, widely oval, widest in posterior half (across abdominal segment III), biconvex, more strongly vaulted ventrally.
</paragraph>
<paragraph id="8B9336E26825FFE74EDD6664FA6BFCDD" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD6664FEF9FEE5" box="[199,260,331,357]" italics="true" pageId="6" pageNumber="516">Head</emphasis>
(
<figureCitation id="13172A676825FFE74F0B6664FE7FFEE6" box="[273,386,331,358]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6–13</figureCitation>
) slightly shorter than wide across compound eyes (head length: width ratio 0.73–0.99); dorsum of head more or less flat, lateral margins concave in front of eyes, more or less reflexed upwards including apex of mandibular plates. Mandibular plates explanate, wider and longer than clypeus, each of them separately rounded apically, meeting (
<figureCitation id="13172A676825FFE74F776698FE48FE52" box="[365,437,439,466]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 6</figureCitation>
) or nearly meeting (
<figureCitation id="13172A676825FFE74CB26698FCBCFE51" box="[680,833,439,465]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 7, 9, 10</figureCitation>
) in front of clypeus, leaving narrow V-shaped cleft anteriad of clypeus apex, or not meeting (
<figureCitation id="13172A676825FFE74C4566F4FD4CFE75" box="[607,689,475,501]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 12</figureCitation>
); clypeus correspondingly narrowing and anteriorly closed (
<figureCitation id="13172A676825FFE74B5166F4FA73FE76" box="[1355,1422,475,502]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 6</figureCitation>
), nearly closed (
<figureCitation id="13172A676825FFE74F2666D0FE30FD99" box="[316,461,511,537]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 7, 9, 10</figureCitation>
), or apically free (
<figureCitation id="13172A676825FFE74C8466D0FD0FFD99" box="[670,754,511,537]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 12</figureCitation>
). Compound eyes moderately large, rounded, protruding out of the head outline in most of their width (
<figureCitation id="13172A676825FFE74C6C650CFCB3FDBD" box="[630,846,547,573]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6–7, 9–10, 12</figureCitation>
). Ocelli small, situated posteriad of compound eyes, close to anterior margin of pronotum. Round, smooth, unpunctured spot present between compound eye and ocellus (
<figureCitation id="13172A676825FFE74E856544FE8EFD05" box="[159,371,619,645]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6–7, 9–10, 12</figureCitation>
). Postgenae behind eyes narrow. Base of antennae including antenniferous tubercle partly (
<figureCitation id="13172A676825FFE74B706544FF3EFD2A" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6–7</figureCitation>
) to completely visible (
<figureCitation id="13172A676825FFE74FD165A0FDE1FD29" box="[459,540,655,681]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 12</figureCitation>
) from above. Antennae 5-segmented, slender. Scape (I) cylindrical, shortest and stoutest; basipedicellite (IIa) shortest; antennomeres IIa to IV slender and uniformly cylindrical. Bucculae (
<figureCitation id="13172A676825FFE74B4A659CFA6EFD4D" box="[1360,1427,691,717]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Fig. 8</figureCitation>
) short, low, anteriorly obtusangulate, with small tooth-like projection anteroventrally, posteriorly widely rounded, leaving apex of labiomere I free. Labrum (
<figureCitation id="13172A676825FFE74C6165D4FD06FC95" box="[635,763,763,789]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 11, 13</figureCitation>
) flat and narrow, reaching base of labiomere II. Lengths of labiomeres from shortest to longest: IV ≤ I &lt;III &lt;II. Labiomere I moderately stout, short, nearly reaching or slightly surpassing posterior margin of bucculae (
<figureCitation id="13172A676825FFE74C7F646CFD18FCDD" box="[613,741,835,861]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 11, 13</figureCitation>
); apex of labium reaching between mesocoxae to metacoxae.
</paragraph>
<paragraph id="8B9336E26825FFE74EDD6448FD55FB21" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD6448FECBFC00" box="[199,310,871,896]" italics="true" pageId="6" pageNumber="516">Pronotum</emphasis>
(
<figureCitation id="13172A676825FFE74F536448FE6EFC01" box="[329,403,871,897]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="6" pageNumber="516">Figs 1</figureCitation>
, 
<figureCitation id="13172A676825FFE74FBF6448FE3EFC01" box="[421,451,871,897]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="6" pageNumber="516">64</figureCitation>
, 
<figureCitation id="13172A676825FFE74FC96448FE0CFC01" box="[467,497,871,897]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="6" pageNumber="516">88</figureCitation>
, 
<figureCitation id="13172A676825FFE74C1B6448FD97FC01" box="[513,618,871,897]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="6" pageNumber="516">108, 110</figureCitation>
). Anterior pronotal margin medially incised, concave, forming arcuate to quadrangular concavity receiving the postocular portion of head (
<figureCitation id="13172A676825FFE74DB864A4FBE4FC25" box="[930,1049,907,933]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6, 10</figureCitation>
, 
<figureCitation id="13172A676825FFE74A3D64A4FBBAFC26" box="[1063,1095,907,934]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="6" pageNumber="516">64</figureCitation>
); anterolateral angles truncate behind eyes, each laterally with one minute, laterally directed denticle (
<figureCitation id="13172A676825FFE74DA26480FBABFC49" box="[952,1110,943,969]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="6" pageNumber="516">Figs 6, 10, 12</figureCitation>
); anterolateral margins sharp but not emarginate, slightly reflexed and minutely to distinctly crenulate (
<figureCitation id="13172A676825FFE74DC564FCFBCAFC6D" box="[991,1079,979,1005]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="6" pageNumber="516">Figs 64</figureCitation>
, 
<figureCitation id="13172A676825FFE74A5F64FCFB9FFC6D" box="[1093,1122,979,1005]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="6" pageNumber="516">88</figureCitation>
) in anterior half, smooth in posterior half, shallowly concave in middle. Humeri rectangulate to slightly obtusangulate with small tubercle present sublaterally; posterolateral margin of pronotum straight or slightly convex, directed obliquely towards anterolateral angles of scutellum, posterolateral angles of pronotum widely rounded; posterior margin of pronotum slightly concave to nearly straight. Anterior part of pronotal disc (in front of humeri) declivent and flat; posterior part of pronotal disc slightly convex (
<figureCitation id="13172A676825FFE74C2663A8FD61FB21" box="[572,668,1159,1185]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="6" pageNumber="516">Fig. 111</figureCitation>
).
</paragraph>
<paragraph id="8B9336E26825FFE74EDD6384FAECFAB1" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD6384FEC8FB45" box="[199,309,1195,1221]" italics="true" pageId="6" pageNumber="516">Scutellum</emphasis>
(
<figureCitation id="13172A676825FFE74F5C6384FE53FB45" box="[326,430,1195,1221]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="6" pageNumber="516">Figs 1, 3</figureCitation>
, 
<figureCitation id="13172A676825FFE74FA76384FE26FB45" box="[445,475,1195,1221]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="6" pageNumber="516">88</figureCitation>
, 
<figureCitation id="13172A676825FFE74FF36384FDE9FB45" box="[489,532,1195,1221]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="6" pageNumber="516">108</figureCitation>
) triangular, longer than wide at base, lateral margins nearly straight or slightly convex at frenal portion, slightly incised at apices of frena (in about two thirds of scutellar length); apex of scutellum parabolic, not surpassing anterodistal angles of corium, narrowly to more broadly rounded. Disc of scutellum anteriorly convex, posteriorly flat, in some species with saddle-like concavity in front of frenal apices.
</paragraph>
<paragraph id="8B9336E26825FFE74EDD6214FAE0FA41" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD6214FEC0FAD5" box="[199,317,1339,1365]" italics="true" pageId="6" pageNumber="516">Hemelytra</emphasis>
(
<figureCitation id="13172A676825FFE74F516214FE53FAD5" box="[331,430,1339,1365]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="6" pageNumber="516">Figs 1, 3</figureCitation>
, 
<figureCitation id="13172A676825FFE74FA06214FE25FAD5" box="[442,472,1339,1365]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="6" pageNumber="516">88</figureCitation>
, 
<figureCitation id="13172A676825FFE74FF96214FDF3FAD5" box="[483,526,1339,1365]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="6" pageNumber="516">108</figureCitation>
). Clavus narrowly triangular, reaching only two thirds of scutellar length. Corium laterally more or less arcuate, anterodistal angles acutangulate, narrowly rounded apically, much surpassing apex of scutellum, reaching about middle of segment V. Membrane translucent, infumate, widely rounded apically, reaching or slightly surpassing apex of abdomen. Membrane bearing 7‒12 prominent and simple veins (
<figureCitation id="13172A676825FFE74AA66288FAECFA41" box="[1212,1297,1446,1473]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="6" pageNumber="516">Fig. 65</figureCitation>
).
</paragraph>
<paragraph id="8B9336E26825FFE74EDD62E4FB99F8A9" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD62E4FDC0FA64" box="[199,573,1483,1509]" italics="true" pageId="6" pageNumber="516">Thoracic pleuron and sternum.</emphasis>
Mesosternum with narrow, median longitudinal carina (
<figureCitation id="13172A676825FFE74AF762E4FAB7FA65" box="[1261,1354,1483,1509]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="6" pageNumber="516">Fig. 26</figureCitation>
: mec). Metasternum with lateral margins raised, narrow (
<figureCitation id="13172A676825FFE74CD662C0FCE3F989" box="[716,798,1519,1545]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="6" pageNumber="516">Fig. 23</figureCitation>
) to wide, flat (
<figureCitation id="13172A676825FFE74DDC62C0FBB7F989" box="[966,1098,1518,1545]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="6" pageNumber="516">Figs 24–25</figureCitation>
: mts). Ostiole (
<figureCitation id="13172A676825FFE74AEC62C0FA84F989" box="[1270,1401,1519,1545]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Figs 14–19</figureCitation>
: o) situated between outer margins of meso- and metacetabulum, elongate oval, directed posterolaterad, accompanied with short and narrow, slightly elevated, spout-shaped peritreme, reaching about middle of metapleuron width; median furrow distinct, at least in basal half of peritreme; periostiolar depression shallow. Evaporatorium very large (
<figureCitation id="13172A676825FFE74EC56150FE9EF919" box="[223,355,1662,1689]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Figs 14, 15</figureCitation>
, 
<figureCitation id="13172A676825FFE74F6B6150FE6DF91A" box="[369,400,1663,1690]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="6" pageNumber="516">68</figureCitation>
), covering most of metapleuron (except its posterior and lateral areas), prolonged medially between meso- and metacoxae, laterally delimited by low longitudinal ridge; on mesopleuron covering its posterior half (mesepimeron), but leaving smooth ca. triangular spot in middle of lateral margin (
<figureCitation id="13172A676825FFE74A6A61E8FB38F961" box="[1136,1221,1735,1761]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Fig. 20</figureCitation>
: sms). Gyrification developed, fine and shallow (
<figureCitation id="13172A676825FFE74FC161C4FDA7F885" box="[475,602,1771,1797]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Figs 14, 21</figureCitation>
). Metathoracic spiracle wide, long and narrow, visible in ventral view (
<figureCitation id="13172A676825FFE74B7161C4FEF6F8A9" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Figs 14–15, 21</figureCitation>
: sp), protected by anastomosing, reticulate mycoid filter processes (
<figureCitation id="13172A676825FFE74A196020FBAAF8A9" box="[1027,1111,1807,1833]" captionStart="FIGURES 14–22" captionStartId="9.[151,264,1779,1803]" captionTargetBox="[151,1436,181,1755]" captionTargetId="figure-29@9.[151,1436,181,1755]" captionTargetPageId="9" captionText="FIGURES 14–22. External scent efferent system of metathoracic scent gland. 14—Agathocles normalis (Distant, 1901), male, Laos: Phou Pane Mt. (23×). 15—A. joceliae sp. nov., female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16—A. limbatus Stål, 1876 (90×); 17—A. flavipes sp. nov. (100×); 18—A. joceliae sp. nov. (75×); 19—A. normalis (Distant, 1901) (85×). 20—A. joceliae sp. nov., evaporatorium laterally on mesopleuron (75×). 21—A. normalis, metathoracic spiracle (42×). 22—A. limbatus, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="6" pageNumber="516">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="8B9336E26825FFE74EDD601CFBA3F815" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="516">
<emphasis id="B958EAF06825FFE74EDD601CFF01F8CC" box="[199,252,1843,1868]" italics="true" pageId="6" pageNumber="516">Legs</emphasis>
(
<figureCitation id="13172A676825FFE74F11601CFEA9F8CD" box="[267,340,1843,1869]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="6" pageNumber="516">Figs 2</figureCitation>
, 
<figureCitation id="13172A676825FFE74F7A601CFE83F8CD" box="[352,382,1843,1869]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="6" pageNumber="516">89</figureCitation>
, 
<figureCitation id="13172A676825FFE74F91601CFE4BF8CD" box="[395,438,1843,1869]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="6" pageNumber="516">109</figureCitation>
). Femora cylindrical, unarmed, rounded in cross section, slightly longer and thicker than appropriate tibiae. All tibiae shallowly sulcate on dorsal surface. All tarsi with tarsomere II shortest, I longest, nearly as long as II and III combined; all tarsomeres dorsally regularly rounded, not grooved.
</paragraph>
<caption id="DF53666A6824FFE64E8D60B8FD70F877" ID-DOI="http://doi.org/10.5281/zenodo.4692373" ID-Zenodo-Dep="4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="7" pageNumber="517" startId="7.[151,264,1943,1967]" targetBox="[203,1384,183,1912]" targetPageId="7">
<paragraph id="8B9336E26824FFE64E8D60B8FD70F877" blockId="7.[151,1437,1943,2040]" pageId="7" pageNumber="517">
<emphasis id="B958EAF06824FFE64E8D60B8FEC1F82F" bold="true" box="[151,316,1943,1967]" pageId="7" pageNumber="517">FIGURES 1–5.</emphasis>
<taxonomicName id="4C2C4D616824FFE64F5960B7FD78F82F" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[323,645,1943,1968]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="517" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF06824FFE64F5960B7FDEFF830" box="[323,530,1944,1968]" italics="true" pageId="7" pageNumber="517">Agathocles limbatus</emphasis>
<bibRefCitation id="EFBD4B136824FFE64C0060B8FD78F82F" author="Stal, C." box="[538,645,1943,1967]" pageId="7" pageNumber="517" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of 
<taxonomicName id="4C2C4D616824FFE64C3E6093FC7BF854" authority="Zhang &amp; Lin, 1984" authorityName="Zhang &amp; Lin" authorityYear="1984" box="[548,902,1980,2004]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="517" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B958EAF06824FFE64C3E6093FD40F853" box="[548,701,1980,2003]" italics="true" pageId="7" pageNumber="517">A. yunnanensis</emphasis>
Zhang &amp; 
<bibRefCitation id="EFBD4B136824FFE64D386093FC7BF854" author="Lin, Y. - J. &amp; Zhang, S. - M." box="[802,902,1980,2004]" pageId="7" pageNumber="517" pagination="267 - 270" refId="ref31784" refString="Lin, Y. - J. &amp; Zhang, S. - M. (1984) Three new species of Pentatomidae from Yunnan, China. Entomotaxonomia, 6 (4), 267 - 270. [in Chinese, English summary]" type="journal article" year="1984">Lin, 1984</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B958EAF06824FFE64D8A6093FC10F854" bold="true" box="[912,1005,1980,2004]" pageId="7" pageNumber="517">
<taxonomicNameLabel id="A26B578B6824FFE64D8A6093FC10F854" box="[912,1005,1980,2004]" pageId="7" pageNumber="517">syn. nov.</taxonomicNameLabel>
</emphasis>
, female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5).
</paragraph>
</caption>
<caption id="DF53666A682BFFE94E8D603CFABBF85F" ID-DOI="http://doi.org/10.5281/zenodo.4692377" ID-Zenodo-Dep="4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="8" pageNumber="518" startId="8.[151,264,1811,1835]" targetBox="[215,1372,181,1787]" targetPageId="8">
<paragraph id="8B9336E2682BFFE94E8D603CFABBF85F" blockId="8.[151,1437,1811,2016]" pageId="8" pageNumber="518">
<emphasis id="B958EAF0682BFFE94E8D603CFEBAF8AB" bold="true" box="[151,327,1811,1835]" pageId="8" pageNumber="518">FIGURES 6–13.</emphasis>
Head. 6–8— 
<taxonomicName id="4C2C4D61682BFFE94FD7603CFCE0F8AB" authority="Stal, 1876: 6" authorityName="Stal" authorityPageNumber="6" authorityYear="1876" box="[461,797,1811,1835]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="518" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF0682BFFE94FD7603CFD67F8AB" box="[461,666,1811,1835]" italics="true" pageId="8" pageNumber="518">Agathocles limbatus</emphasis>
<bibRefCitation id="EFBD4B13682BFFE94CBA603CFCE0F8AB" author="Stal, C." box="[672,797,1811,1835]" pageId="8" pageNumber="518" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876: 6</bibRefCitation>
</taxonomicName>
—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9— 
<taxonomicName id="4C2C4D61682BFFE94A686017FA65F8D0" authority="(Distant, 1901 a)" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[1138,1432,1847,1872]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="8" pageNumber="518" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0682BFFE94A686017FB14F8CF" box="[1138,1257,1847,1871]" italics="true" pageId="8" pageNumber="518">A. normalis</emphasis>
(
<bibRefCitation id="EFBD4B13682BFFE94AED6018FA6DF8D0" author="Distant, W. L." box="[1271,1424,1847,1872]" pageId="8" pageNumber="518" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901a</bibRefCitation>
)
</taxonomicName>
, male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11— 
<taxonomicName id="4C2C4D61682BFFE94CE66073FC9AF8F3" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[764,871,1883,1907]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="518" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF0682BFFE94CE66073FC9AF8F3" box="[764,871,1883,1907]" italics="true" pageId="8" pageNumber="518">A. flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682BFFE94D746073FC43F8F4" bold="true" box="[878,958,1884,1908]" pageId="8" pageNumber="518">
<taxonomicNameLabel id="A26B578B682BFFE94D746073FC43F8F4" box="[878,958,1884,1908]" pageId="8" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13— 
<taxonomicName id="4C2C4D61682BFFE94CAB60AFFCE0F817" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[689,797,1919,1943]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="518" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682BFFE94CAB60AFFCE0F817" box="[689,797,1919,1943]" italics="true" pageId="8" pageNumber="518">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682BFFE94D3E60AFFC89F818" bold="true" box="[804,884,1920,1944]" pageId="8" pageNumber="518">
<taxonomicNameLabel id="A26B578B682BFFE94D3E60AFFC89F818" box="[804,884,1920,1944]" pageId="8" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment.
</paragraph>
</caption>
<caption id="DF53666A682AFFE84E8D61DCFC90F863" ID-DOI="http://doi.org/10.5281/zenodo.4692379" ID-Zenodo-Dep="4692379" httpUri="https://zenodo.org/record/4692379/files/figure.png" pageId="9" pageNumber="519" startId="9.[151,264,1779,1803]" targetBox="[151,1436,181,1755]" targetPageId="9">
<paragraph id="8B9336E2682AFFE84E8D61DCFC90F863" blockId="9.[151,1437,1779,2020]" pageId="9" pageNumber="519">
<emphasis id="B958EAF0682AFFE84E8D61DCFEA4F88B" bold="true" box="[151,345,1779,1803]" pageId="9" pageNumber="519">FIGURES 14–22.</emphasis>
External scent efferent system of metathoracic scent gland. 14— 
<taxonomicName id="4C2C4D61682AFFE84A0D61DCFA65F88C" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[1047,1432,1779,1804]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0682AFFE84A0D61DCFB10F88B" box="[1047,1261,1779,1803]" italics="true" pageId="9" pageNumber="519">Agathocles normalis</emphasis>
(Distant, 1901)
</taxonomicName>
, male, Laos: Phou Pane Mt. (23×). 15— 
<taxonomicName id="4C2C4D61682AFFE84C216037FD57F8AF" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[571,682,1815,1839]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682AFFE84C216037FD57F8AF" box="[571,682,1815,1839]" italics="true" pageId="9" pageNumber="519">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682AFFE84CAE6037FCF5F8B0" bold="true" box="[692,776,1816,1840]" pageId="9" pageNumber="519">
<taxonomicNameLabel id="A26B578B682AFFE84CAE6037FCF5F8B0" box="[692,776,1816,1840]" pageId="9" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, Malaysia: Kelantan: Kampong Timor. 16–19—ostiole and peritreme: 16— 
<taxonomicName id="4C2C4D61682AFFE84F736013FDAEF8D3" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[361,595,1851,1875]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF0682AFFE84F736013FE22F8D3" box="[361,479,1851,1875]" italics="true" pageId="9" pageNumber="519">A. limbatus</emphasis>
<bibRefCitation id="EFBD4B13682AFFE84FF36014FDAEF8D3" author="Stal, C." box="[489,595,1851,1875]" pageId="9" pageNumber="519" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
(90×); 17— 
<taxonomicName id="4C2C4D61682AFFE84CC26013FCBBF8D3" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[728,838,1851,1875]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF0682AFFE84CC26013FCBBF8D3" box="[728,838,1851,1875]" italics="true" pageId="9" pageNumber="519">A. flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682AFFE84D556013FC5EF8D4" bold="true" box="[847,931,1852,1876]" pageId="9" pageNumber="519">
<taxonomicNameLabel id="A26B578B682AFFE84D556013FC5EF8D4" box="[847,931,1852,1876]" pageId="9" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(100×); 18— 
<taxonomicName id="4C2C4D61682AFFE84A2F6013FB59F8D3" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1077,1188,1851,1875]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682AFFE84A2F6013FB59F8D3" box="[1077,1188,1851,1875]" italics="true" pageId="9" pageNumber="519">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682AFFE84AB76013FAFCF8D4" bold="true" box="[1197,1281,1852,1876]" pageId="9" pageNumber="519">
<taxonomicNameLabel id="A26B578B682AFFE84AB76013FAFCF8D4" box="[1197,1281,1852,1876]" pageId="9" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(75×); 19— 
<taxonomicName id="4C2C4D61682AFFE84B9C6013FE6AF8F8" authority="(Distant, 1901)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0682AFFE84B9C6013FF0CF8F7" italics="true" pageId="9" pageNumber="519">A. normalis</emphasis>
(Distant, 1901)
</taxonomicName>
(85×). 20— 
<taxonomicName id="4C2C4D61682AFFE84C06604FFD76F8F7" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[540,651,1887,1911]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682AFFE84C06604FFD76F8F7" box="[540,651,1887,1911]" italics="true" pageId="9" pageNumber="519">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682AFFE84C8F604FFD15F8F8" bold="true" box="[661,744,1888,1912]" pageId="9" pageNumber="519">
<taxonomicNameLabel id="A26B578B682AFFE84C8F604FFD15F8F8" box="[661,744,1888,1912]" pageId="9" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, evaporatorium laterally on mesopleuron (75×). 21— 
<taxonomicName id="4C2C4D61682AFFE84B07604FFA6BF8F7" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[1309,1430,1887,1911]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0682AFFE84B07604FFA6BF8F7" box="[1309,1430,1887,1911]" italics="true" pageId="9" pageNumber="519">A. normalis</emphasis>
</taxonomicName>
, metathoracic spiracle (42×). 22— 
<taxonomicName id="4C2C4D61682AFFE84FF560ABFD9EF81B" authorityName="Stal" authorityYear="1876" box="[495,611,1923,1947]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="519" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF0682AFFE84FF560ABFD9EF81B" box="[495,611,1923,1947]" italics="true" pageId="9" pageNumber="519">A. limbatus</emphasis>
</taxonomicName>
, detail of metathoracic spiracle with reticulate filter processes (320×). Lettering: ev—evaporatorium, mf—median furrow, o—ostiole, pe—peritreme, sms—smooth spot on mesopleuron, sp—spiracle. Scale bars: 1 mm (14, 15, 21), 0.5 mm (16–20), 0.1 mm (22). Orig. P. Kment.
</paragraph>
</caption>
<caption id="DF53666A6829FFEB4E8D62B9FB69F9BE" ID-DOI="http://doi.org/10.5281/zenodo.4692381" ID-Zenodo-Dep="4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="10" pageNumber="520" startId="10.[151,264,1430,1454]" targetBox="[151,1436,181,1406]" targetPageId="10">
<paragraph id="8B9336E26829FFEB4E8D62B9FB69F9BE" blockId="10.[151,1437,1430,1598]" pageId="10" pageNumber="520">
<emphasis id="B958EAF06829FFEB4E8D62B9FEA9FA2E" bold="true" box="[151,340,1430,1454]" pageId="10" pageNumber="520">FIGURES 23–28.</emphasis>
Morphology of thorax and abdomen. 23–25—metasternum: 23— 
<taxonomicName id="4C2C4D616829FFEB4DF762B9FB4EFA2E" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1005,1203,1430,1454]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="520" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF06829FFEB4DF762B9FB4EFA2E" box="[1005,1203,1430,1454]" italics="true" pageId="10" pageNumber="520">Agathocles flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF06829FFEB4AA262B9FAF5FA2E" bold="true" box="[1208,1288,1430,1454]" pageId="10" pageNumber="520">
<taxonomicNameLabel id="A26B578B6829FFEB4AA262B9FAF5FA2E" box="[1208,1288,1430,1454]" pageId="10" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(50×); 24— 
<taxonomicName id="4C2C4D616829FFEB4B9C62B8FF1AFA52" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="520" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF06829FFEB4B9C62B8FF1AFA52" italics="true" pageId="10" pageNumber="520">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF06829FFEB4EEA6295FEB9FA52" bold="true" box="[240,324,1466,1490]" pageId="10" pageNumber="520">
<taxonomicNameLabel id="A26B578B6829FFEB4EEA6295FEB9FA52" box="[240,324,1466,1490]" pageId="10" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(55×); 25— 
<taxonomicName id="4C2C4D616829FFEB4FD36294FD4FFA52" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[457,690,1466,1490]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="520" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF06829FFEB4FD36294FDC2FA52" box="[457,575,1466,1490]" italics="true" pageId="10" pageNumber="520">A. limbatus</emphasis>
<bibRefCitation id="EFBD4B136829FFEB4C526295FD4FFA52" author="Stal, C." box="[584,690,1466,1490]" pageId="10" pageNumber="520" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
(37×). 26–28— 
<taxonomicName id="4C2C4D616829FFEB4D416294FC2CFA52" authorityName="Stal" authorityYear="1876" box="[859,977,1466,1490]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="520" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF06829FFEB4D416294FC2CFA52" box="[859,977,1466,1490]" italics="true" pageId="10" pageNumber="520">A. limbatus</emphasis>
</taxonomicName>
: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment.
</paragraph>
</caption>
<paragraph id="8B9336E26829FFEB4EDD61BFFCFDF802" blockId="10.[151,1437,1680,1922]" pageId="10" pageNumber="520">
<emphasis id="B958EAF06829FFEB4EDD61BFFE57F92A" box="[199,426,1680,1706]" italics="true" pageId="10" pageNumber="520">Pregenital abdomen</emphasis>
slightly wider than pronotum across humeri (
<figureCitation id="13172A676829FFEB4DBC61BFFC10F92B" box="[934,1005,1680,1707]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="10" pageNumber="520">Figs 1</figureCitation>
, 
<figureCitation id="13172A676829FFEB4DE061BEFBE5F92B" box="[1018,1048,1681,1707]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="10" pageNumber="520">88</figureCitation>
, 
<figureCitation id="13172A676829FFEB4A3E61BEFBAFF92B" box="[1060,1106,1681,1707]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="10" pageNumber="520">108</figureCitation>
). Connexivum wide, exposed laterally and slightly lamellate (
<figureCitation id="13172A676829FFEB4FE4619BFDBDF94F" box="[510,576,1716,1743]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="10" pageNumber="520">Fig. 1</figureCitation>
); posterolateral angles of each laterotergite III–VII about rectangulate, only slightly protruding from outline of the abdomen. Abdominal venter uniformly convex, neither grooved, nor keeled medially; ventrite III without spinose projection or depression medially (
<figureCitation id="13172A676829FFEB4DC261D3FBDEF897" box="[984,1059,1788,1815]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="10" pageNumber="520">Figs 2</figureCitation>
, 
<figureCitation id="13172A676829FFEB4A2B61D2FBB2F897" box="[1073,1103,1789,1815]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="10" pageNumber="520">89</figureCitation>
, 
<figureCitation id="13172A676829FFEB4A4461D2FB71F897" box="[1118,1164,1789,1815]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="10" pageNumber="520">109</figureCitation>
). Spiracle on ventrite II covered by metapleuron or its posterior margin more or less exposed. Each ventrite III–VII laterally with spiracle and pair of trichobothria situated posterolaterad of spiracle (
<figureCitation id="13172A676829FFEB4D2E606BFC4AF8DF" box="[820,951,1860,1887]" captionStart="FIGURES 23–28" captionStartId="10.[151,264,1430,1454]" captionTargetBox="[151,1436,181,1406]" captionTargetId="figure-19@10.[151,1436,181,1406]" captionTargetPageId="10" captionText="FIGURES 23–28. Morphology of thorax and abdomen. 23–25—metasternum: 23—Agathocles flavipes sp. nov. (50×); 24—A. joceliae sp. nov. (55×); 25—A. limbatus Stål, 1876 (37×). 26–28—A. limbatus: 26—mesosternum (50×); 27—spiracle and trichobothria on abdominal segment VI (170×); 28—apex of abdomen and genital capsule, lateral view (35×). Lettering: cx2— mesocoxa, cx3—metacoxa, gc—genital capsule, la2‒4—labiomere II‒IV; msc—mesosternal carina; mts—metasternum; sp— abdominal spiracle; tr—trichobothrium. Scale bars: 1 mm (23–26, 28), 0.3 mm (27). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692381" httpUri="https://zenodo.org/record/4692381/files/figure.png" pageId="10" pageNumber="520">Figs 27–28</figureCitation>
); inner trichobothrium situated in and outer trichobothrium laterad of the imaginary spiracular line.
</paragraph>
<caption id="DF53666A6828FFEA4E8D64D5FE0FFBDA" ID-DOI="http://doi.org/10.5281/zenodo.4692383" ID-Zenodo-Dep="4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="11" pageNumber="521" startId="11.[151,264,1018,1042]" targetBox="[151,1432,181,994]" targetPageId="11">
<paragraph id="8B9336E26828FFEA4E8D64D5FE0FFBDA" blockId="11.[151,1437,1018,1114]" pageId="11" pageNumber="521">
<emphasis id="B958EAF06828FFEA4E8D64D5FEAAFB92" bold="true" box="[151,343,1018,1042]" pageId="11" pageNumber="521">FIGURES 29–32.</emphasis>
External female genitalia. 29–30— 
<taxonomicName id="4C2C4D616828FFEA4CD764D5FBEFFB92" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[717,1042,1018,1042]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="521" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF06828FFEA4CD764D5FC63FB92" box="[717,926,1018,1042]" italics="true" pageId="11" pageNumber="521">Agathocles limbatus</emphasis>
<bibRefCitation id="EFBD4B136828FFEA4DB264D5FBEFFB92" author="Stal, C." box="[936,1042,1018,1042]" pageId="11" pageNumber="521" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
(29—27×); 31— 
<taxonomicName id="4C2C4D616828FFEA4AD064D4FAC5FB92" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1226,1336,1018,1042]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="521" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF06828FFEA4AD064D4FAC5FB92" box="[1226,1336,1018,1042]" italics="true" pageId="11" pageNumber="521">A. flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF06828FFEA4B5B64D5FA68FB92" bold="true" box="[1345,1429,1018,1042]" pageId="11" pageNumber="521">
<taxonomicNameLabel id="A26B578B6828FFEA4B5B64D5FA68FB92" box="[1345,1429,1018,1042]" pageId="11" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 32— 
<taxonomicName id="4C2C4D616828FFEA4ED06330FECAFBB6" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[202,311,1054,1078]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="521" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF06828FFEA4ED06330FECAFBB6" box="[202,311,1054,1078]" italics="true" pageId="11" pageNumber="521">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF06828FFEA4F256331FE6CFBB6" bold="true" box="[319,401,1054,1078]" pageId="11" pageNumber="521">
<taxonomicNameLabel id="A26B578B6828FFEA4F256331FE6CFBB6" box="[319,401,1054,1078]" pageId="11" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment.
</paragraph>
</caption>
<paragraph id="8B9336E26828FFEA4EDD63A7FD6BF887" blockId="11.[151,1437,1160,2015]" pageId="11" pageNumber="521">
<emphasis id="B958EAF06828FFEA4EDD63A7FE8DFB22" box="[199,368,1160,1186]" italics="true" pageId="11" pageNumber="521">Male genitalia.</emphasis>
Genital capsule (
<figureCitation id="13172A676828FFEA4C3663A7FD56FB23" box="[556,683,1160,1187]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="11" pageNumber="521">Figs 37–41</figureCitation>
, 
<figureCitation id="13172A676828FFEA4CAE63A7FD01FB22" box="[692,764,1160,1186]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">69–74</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D1F63A6FCB3FB23" box="[773,846,1161,1187]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">90–94</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D4D63A7FC44FB23" box="[855,953,1160,1187]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="11" pageNumber="521">113–116</figureCitation>
) nearly quadrangular in dorsal/ventral view, longer than wide. Dorsal rim (
<figureCitation id="13172A676828FFEA4FF66383FDB9FB47" box="[492,580,1196,1223]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">Figs 72</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C4A6382FD8DFB47" box="[592,624,1197,1223]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">92</figureCitation>
: dr) deeply and concavely excavated with median emargination. Ventral rim undulated, slightly (
<figureCitation id="13172A676828FFEA4FB063FFFDFEFB6B" box="[426,515,1232,1259]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="11" pageNumber="521">Figs 41</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C0A63FEFDD0FB6B" box="[528,557,1233,1259]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">73</figureCitation>
) to deeply (
<figureCitation id="13172A676828FFEA4CA363FFFCECFB6B" box="[697,785,1232,1259]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">Figs 93</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D0563FFFCB4FB6B" box="[799,841,1232,1259]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="11" pageNumber="521">115</figureCitation>
) incised medially; ventral rim infolding unmodified (
<figureCitation id="13172A676828FFEA4EBA63DBFF0EFA8F" box="[160,243,1268,1295]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">Fig. 91</figureCitation>
), medially with short median process perpendicular to ventral rim (
<figureCitation id="13172A676828FFEA4DE863DBFB8AFA8F" box="[1010,1143,1268,1295]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">Figs 71, 74</figureCitation>
), or with wide depression (
<figureCitation id="13172A676828FFEA4E856237FEDDFAB3" box="[159,288,1304,1331]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="11" pageNumber="521">Figs 37–39</figureCitation>
, 
<figureCitation id="13172A676828FFEA4F306236FE72FAB3" box="[298,399,1305,1331]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="11" pageNumber="521">113–114</figureCitation>
). Superior processes of dorsal rim usually present (
<figureCitation id="13172A676828FFEA4DDF6237FB93FAB3" box="[965,1134,1304,1331]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="11" pageNumber="521">Figs 37–39, 43</figureCitation>
, 
<figureCitation id="13172A676828FFEA4A636236FB16FAB2" box="[1145,1259,1304,1331]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">90–91, 96</figureCitation>
, 
<figureCitation id="13172A676828FFEA4AEC6236FA72FAB3" box="[1270,1423,1305,1331]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="11" pageNumber="521">113–114, 117</figureCitation>
), in 
<taxonomicName id="4C2C4D616828FFEA4EAE6212FED6FAD6" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[180,299,1340,1367]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="521" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF06828FFEA4EAE6212FED6FAD6" box="[180,299,1340,1367]" italics="true" pageId="11" pageNumber="521">A. flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF06828FFEA4F296212FE70FAD7" bold="true" box="[307,397,1341,1367]" pageId="11" pageNumber="521">
<taxonomicNameLabel id="A26B578B6828FFEA4F296212FE70FAD7" box="[307,397,1341,1367]" pageId="11" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
absent. Parameral socket open (
<figureCitation id="13172A676828FFEA4CE26213FCB2FAD7" box="[760,847,1340,1367]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="11" pageNumber="521">Figs 38</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D466212FC87FAD7" box="[860,890,1341,1367]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="11" pageNumber="521">91</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D9C6212FC4EFAD7" box="[902,947,1341,1367]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="11" pageNumber="521">114</figureCitation>
: ps) or in 
<taxonomicName id="4C2C4D616828FFEA4A3F6212FB66FAD6" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1061,1179,1340,1367]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="521" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B958EAF06828FFEA4A3F6212FB66FAD6" box="[1061,1179,1340,1367]" italics="true" pageId="11" pageNumber="521">A. flavipes</emphasis>
</taxonomicName>
closed, kidney-shaped (
<figureCitation id="13172A676828FFEA4E85624FFED9FAFB" box="[159,292,1376,1403]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">Figs 71, 74</figureCitation>
: ps). Paramere (
<figureCitation id="13172A676828FFEA4FC1624FFDA3FAFB" box="[475,606,1376,1403]" captionStart="FIGURES 44–53" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[154,1433,181,1919]" captionTargetId="figure-19@16.[154,1433,181,1919]" captionTargetPageId="16" captionText="FIGURES 44–53. Paramere of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (44–46, 50, 52) and China: Guangxi (47–49, 51, 53) (five different views, 70×). Lettering: ap—apical process of paramere, bp—basal process of paramere, map— projection of mesal angle of paramere stem. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692391" httpUri="https://zenodo.org/record/4692391/files/figure.png" pageId="11" pageNumber="521">Figs 44–53</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C71624FFD49FAFB" box="[619,692,1376,1403]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">75–77</figureCitation>
, 
<figureCitation id="13172A676828FFEA4CDA624EFCF5FAFB" box="[704,776,1377,1403]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="11" pageNumber="521">84–87</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D0F624EFC91FAFB" box="[789,876,1377,1403]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="11" pageNumber="521">97–101</figureCitation>
, 
<figureCitation id="13172A676828FFEA4D62624EFC26FAFB" box="[888,987,1377,1403]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="11" pageNumber="521">118–124</figureCitation>
) with main axis of parameral crown Cshaped in lateral view; basal process (bp) foliaceous, roughly semicircular, apically with row of few to many long setae (
<figureCitation id="13172A676828FFEA4EC56287FE44FA43" box="[223,441,1448,1475]" captionStart="FIGURES 44–53" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[154,1433,181,1919]" captionTargetId="figure-19@16.[154,1433,181,1919]" captionTargetPageId="16" captionText="FIGURES 44–53. Paramere of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (44–46, 50, 52) and China: Guangxi (47–49, 51, 53) (five different views, 70×). Lettering: ap—apical process of paramere, bp—basal process of paramere, map— projection of mesal angle of paramere stem. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692391" httpUri="https://zenodo.org/record/4692391/files/figure.png" pageId="11" pageNumber="521">Figs 44, 47–48, 51</figureCitation>
, 
<figureCitation id="13172A676828FFEA4FDC6287FDF2FA43" box="[454,527,1448,1475]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">75–76</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C016286FD8EFA43" box="[539,627,1449,1475]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="11" pageNumber="521">97, 101</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C9A6286FD1BFA43" box="[640,742,1449,1475]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="11" pageNumber="521">118, 121</figureCitation>
); stem (st) short and broad; apical process (ap) hook-shaped, apically more or less obtuse (
<figureCitation id="13172A676828FFEA4FC662E3FDCEFA66" box="[476,563,1484,1510]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">Figs 76</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C2462E2FD7BFA67" box="[574,646,1485,1511]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="11" pageNumber="521">84–87</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C8B62E2FD27FA67" box="[657,730,1485,1511]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="11" pageNumber="521">97–99</figureCitation>
, 
<figureCitation id="13172A676828FFEA4CFF62E2FCBBFA67" box="[741,838,1485,1511]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="11" pageNumber="521">118, 122</figureCitation>
) or triangular, more pointed apically (
<figureCitation id="13172A676828FFEA4AF462E3FEF7F98B" captionStart="FIGURES 44–53" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[154,1433,181,1919]" captionTargetId="figure-19@16.[154,1433,181,1919]" captionTargetPageId="16" captionText="FIGURES 44–53. Paramere of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (44–46, 50, 52) and China: Guangxi (47–49, 51, 53) (five different views, 70×). Lettering: ap—apical process of paramere, bp—basal process of paramere, map— projection of mesal angle of paramere stem. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692391" httpUri="https://zenodo.org/record/4692391/files/figure.png" pageId="11" pageNumber="521">Figs 44, 46–47, 49, 52–53</figureCitation>
), always provided with an elongate patch of unpigmented, scaled texture (
<figureCitation id="13172A676828FFEA4A5262DFFB57F98B" box="[1096,1194,1520,1547]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="11" pageNumber="521">Fig. 124</figureCitation>
); mesal angle of stem with or without an additional projection variable in shape (
<figureCitation id="13172A676828FFEA4D38613BFBE2F9AF" box="[802,1055,1556,1583]" captionStart="FIGURES 44–53" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[154,1433,181,1919]" captionTargetId="figure-19@16.[154,1433,181,1919]" captionTargetPageId="16" captionText="FIGURES 44–53. Paramere of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (44–46, 50, 52) and China: Guangxi (47–49, 51, 53) (five different views, 70×). Lettering: ap—apical process of paramere, bp—basal process of paramere, map— projection of mesal angle of paramere stem. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692391" httpUri="https://zenodo.org/record/4692391/files/figure.png" pageId="11" pageNumber="521">Figs 44, 46–47, 50–51</figureCitation>
, 
<figureCitation id="13172A676828FFEA4A30613AFB89F9AF" box="[1066,1140,1557,1583]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="11" pageNumber="521">84–87</figureCitation>
). Phallus (
<figureCitation id="13172A676828FFEA4AFD613BFA9AF9AF" box="[1255,1383,1556,1583]" captionStart="FIGURES 54–59" captionStartId="17.[151,264,1598,1622]" captionTargetBox="[158,1432,190,1567]" captionTargetId="figure-29@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURES 54–59. Penis of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (54–56—non inflated, 57–59—inflated): 54, 57—dorsal view; 55, 58—lateral view; 56—ventral view; 59—apical view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival process, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692393" httpUri="https://zenodo.org/record/4692393/files/figure.png" pageId="11" pageNumber="521">Figs 54–59</figureCitation>
, 
<figureCitation id="13172A676828FFEA4B68613AFF48F9D3" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">78– 80</figureCitation>
, 
<figureCitation id="13172A676828FFEA4EDB6116FEF7F9D3" box="[193,266,1593,1619]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="11" pageNumber="521">82–83</figureCitation>
, 
<figureCitation id="13172A676828FFEA4F0F6116FE81F9D3" box="[277,380,1593,1619]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="11" pageNumber="521">102–107</figureCitation>
): Articulatory apparatus composed of basal plate (bpt) and support bridge complex surrounded by intersegmental membrane (ism); 1+1 capitate processes (cp) connected to support bridge complex (sbc) by means of dorsal connectives (dc) (
<figureCitation id="13172A676828FFEA4C0361AFFD61F91B" box="[537,668,1664,1691]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="11" pageNumber="521">Figs 82–83</figureCitation>
). Phallotheca short, barrel shaped, constricted at both ends, broadest in middle. A pair of dorsal conjunctival processes, transversely placed and ending in sclerotised processes (
<figureCitation id="13172A676828FFEA4B5B618BFF48F963" captionStart="FIGURES 54–59" captionStartId="17.[151,264,1598,1622]" captionTargetBox="[158,1432,190,1567]" captionTargetId="figure-29@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURES 54–59. Penis of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (54–56—non inflated, 57–59—inflated): 54, 57—dorsal view; 55, 58—lateral view; 56—ventral view; 59—apical view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival process, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692393" httpUri="https://zenodo.org/record/4692393/files/figure.png" pageId="11" pageNumber="521">Figs 57, 59</figureCitation>
, 
<figureCitation id="13172A676828FFEA4ED861E6FEF6F963" box="[194,267,1737,1763]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">78–79</figureCitation>
, 
<figureCitation id="13172A676828FFEA4F0C61E7FE81F963" box="[278,380,1736,1763]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="11" pageNumber="521">105–107</figureCitation>
); median penial plates fused basally, outer margins broadly and thickly sclerotized, encircling short, curved aedeagus (
<figureCitation id="13172A676828FFEA4FBC61C3FE00F887" box="[422,509,1772,1799]" captionStart="FIGURES 54–59" captionStartId="17.[151,264,1598,1622]" captionTargetBox="[158,1432,190,1567]" captionTargetId="figure-29@17.[151,1436,181,1574]" captionTargetPageId="17" captionText="FIGURES 54–59. Penis of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (54–56—non inflated, 57–59—inflated): 54, 57—dorsal view; 55, 58—lateral view; 56—ventral view; 59—apical view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival process, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692393" httpUri="https://zenodo.org/record/4692393/files/figure.png" pageId="11" pageNumber="521">Figs 57</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C1361C2FDDAF887" box="[521,551,1773,1799]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="11" pageNumber="521">80</figureCitation>
, 
<figureCitation id="13172A676828FFEA4C2961C3FD9CF887" box="[563,609,1772,1799]" captionStart="FIGURES 102–107" captionStartId="28.[151,264,1763,1787]" captionTargetBox="[161,1424,189,1732]" captionTargetId="figure-19@28.[151,1436,181,1740]" captionTargetPageId="28" captionText="FIGURES 102–107. Penis of Agathocles species. 102–105—A. joceliae sp. nov. (102–104—not inflated, 105—inflated): 102—dorsal view, 103—lateral view, 104—ventral view, 105—apical view. 106–107—A. normalis (Distant, 1901), inflated: 106—dorsal view, 107—ventral view. Lettering: ad—aedeagus, dlc—dorsolateral conjunctival processes, mpp—median penial plate, pt—phallotheca. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692409" httpUri="https://zenodo.org/record/4692409/files/figure.png" pageId="11" pageNumber="521">105</figureCitation>
: ad).
</paragraph>
<paragraph id="8B9336E26828FFED4EDD603FFED5FE51" blockId="11.[151,1437,1160,2015]" lastBlockId="12.[151,1437,151,465]" lastPageId="12" lastPageNumber="522" pageId="11" pageNumber="521">
<emphasis id="B958EAF06828FFEA4EDD603FFE78F8AA" box="[199,389,1808,1834]" italics="true" pageId="11" pageNumber="521">Female genitalia</emphasis>
(
<figureCitation id="13172A676828FFEA4F88603FFDEEF8AB" box="[402,531,1808,1835]" captionStart-0="FIGURES 29–32" captionStart-1="FIGURES 33–36" captionStartId-0="11.[151,264,1018,1042]" captionStartId-1="12.[151,264,1778,1802]" captionTargetBox-0="[151,1432,181,994]" captionTargetBox-1="[161,1426,495,1755]" captionTargetId-0="figure-29@11.[151,1436,181,994]" captionTargetId-1="figure-163@12.[151,1436,489,1755]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." captionText-1="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi-0="http://doi.org/10.5281/zenodo.4692383" figureDoi-1="http://doi.org/10.5281/zenodo.4692385" httpUri-0="https://zenodo.org/record/4692383/files/figure.png" httpUri-1="https://zenodo.org/record/4692385/files/figure.png" pageId="11" pageNumber="521">Figs 29–36</figureCitation>
). Laterotergite VIII subtriangular. Laterotergite IX broadest preapically, gradually narrowing apicad, apically more or less rounded, medially touching each other or not (
<figureCitation id="13172A676828FFEA4A69601BFB0BF8CF" box="[1139,1270,1844,1871]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="11" pageNumber="521">Figs 29–32</figureCitation>
, 
<figureCitation id="13172A676828FFEA4B19601AFADEF8CF" box="[1283,1315,1845,1871]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="11" pageNumber="521">34</figureCitation>
). Valvifers VIII quadrant with median margins juxtaposed and dorsal margins slightly concave to slightly convex (
<figureCitation id="13172A676828FFEA4B0D6077FA65F8F3" box="[1303,1432,1880,1907]" captionStart="FIGURES 29–32" captionStartId="11.[151,264,1018,1042]" captionTargetBox="[151,1432,181,994]" captionTargetId="figure-29@11.[151,1436,181,994]" captionTargetPageId="11" captionText="FIGURES 29–32. External female genitalia. 29–30—Agathocles limbatus Stål, 1876 (29—27×); 31—A. flavipes sp. nov.; 32—A. joceliae sp. nov. Lettering: lt8–9—laterotergites VIII–IX, t8—tergite VIII, vf8–9—valvifers VIII–IX, x—segment X. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692383" httpUri="https://zenodo.org/record/4692383/files/figure.png" pageId="11" pageNumber="521">Figs 29–32</figureCitation>
, 
<figureCitation id="13172A676828FFEA4E8D6052FF4AF817" box="[151,183,1917,1943]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="11" pageNumber="521">34</figureCitation>
). Valvifers IX fused medially with median fusion line well apparent or not. Spermathecal opening associated with a small annular sclerite which has a pair of oblique projections surrounding spermathecal opening laterally, and a larger, arch-shaped sclerite surrounding the previous sclerite (
<figureCitation id="13172A676828FFEA4D4260EBFC24F85E" box="[856,985,1988,2015]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="11" pageNumber="521">Figs 33, 36</figureCitation>
: so); ring sclerites present, unsclerotized (
<figureCitation id="13172A67682FFFED4EBA67B8FF08FF31" box="[160,245,151,177]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Fig. 36</figureCitation>
: rs) or slightly sclerotized (
<figureCitation id="13172A67682FFFED4C3467B8FD49FF31" box="[558,692,151,177]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 34</figureCitation>
: rs). Spermatheca: proximal duct (
<figureCitation id="13172A67682FFFED4A2767B8FB6EFF31" box="[1085,1171,151,177]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Fig. 33</figureCitation>
: pd) thicker and longer than distal duct; dilation long (
<figureCitation id="13172A67682FFFED4C1A6794FD4AFF55" box="[512,695,186,213]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 35, 36</figureCitation>
: dil), distal invagination (‘sclerotized rod’) distinctly broadened distally (
<figureCitation id="13172A67682FFFED4EE667F0FE78FF79" box="[252,389,222,249]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 35</figureCitation>
: div), subparallel in most of its length, basally either straight and broadened (
<figureCitation id="13172A67682FFFED4B1467F0FA69FF79" box="[1294,1428,222,249]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 35</figureCitation>
) or thin and strongly curved (
<figureCitation id="13172A67682FFFED4FFC662CFDC3FE9E" box="[486,574,259,286]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Fig. 36</figureCitation>
); distal duct (
<figureCitation id="13172A67682FFFED4CC4662CFC68FE9D" box="[734,917,258,285]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 35, 36</figureCitation>
: dd) short and thin, much shorter than distal invagination; proximal flange (
<figureCitation id="13172A67682FFFED4FE36608FD51FEC1" box="[505,684,294,321]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 35, 36</figureCitation>
: pf) narrower than distal flange; intermediate part of spermatheca narrow, distal flange either separated (
<figureCitation id="13172A67682FFFED4C4C6664FD56FEE5" box="[598,683,330,357]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Fig. 35</figureCitation>
) or tightly attached to apical receptacle (
<figureCitation id="13172A67682FFFED4A946664FAE5FEE5" box="[1166,1304,331,357]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 36</figureCitation>
: df); apical receptacle subglobular (
<figureCitation id="13172A67682FFFED4FB96640FE09FE09" box="[419,500,366,393]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Fig. 35</figureCitation>
) or subhemispherical (
<figureCitation id="13172A67682FFFED4CEF6640FC8AFE09" box="[757,887,367,393]" captionStart="FIGURES 33–36" captionStartId="12.[151,264,1778,1802]" captionTargetBox="[161,1426,495,1755]" captionTargetId="figure-163@12.[151,1436,489,1755]" captionTargetPageId="12" captionText="FIGURES 33–36. External and internal female genitalia. 33—Agathocles limbatus Stål, 1876; 34–35—A. flavipes sp. nov.; 36—A. joceliae sp. nov. Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35)." figureDoi="http://doi.org/10.5281/zenodo.4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522">Figs 33, 36</figureCitation>
: ar), with one long projection directed proximad, not reaching to surpassing proximal flange, and one to four short processes of various orientation, not surpassing distal flange.
</paragraph>
</subSubSection>
<caption id="DF53666A682FFFED4E8D61DDFB03F8F6" ID-DOI="http://doi.org/10.5281/zenodo.4692385" ID-Zenodo-Dep="4692385" httpUri="https://zenodo.org/record/4692385/files/figure.png" pageId="12" pageNumber="522" startId="12.[151,264,1778,1802]" targetBox="[161,1426,495,1755]" targetPageId="12">
<paragraph id="8B9336E2682FFFED4E8D61DDFB03F8F6" blockId="12.[151,1436,1778,1910]" pageId="12" pageNumber="522">
<emphasis id="B958EAF0682FFFED4E8D61DDFEABF88A" bold="true" box="[151,342,1778,1802]" pageId="12" pageNumber="522">FIGURES 33–36.</emphasis>
External and internal female genitalia. 33— 
<taxonomicName id="4C2C4D61682FFFED4D3E61DDFB94F88A" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[804,1129,1778,1802]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="522" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF0682FFFED4D3E61DDFC09F88A" box="[804,1012,1778,1802]" italics="true" pageId="12" pageNumber="522">Agathocles limbatus</emphasis>
<bibRefCitation id="EFBD4B13682FFFED4DE761DDFB94F88A" author="Stal, C." box="[1021,1129,1778,1802]" pageId="12" pageNumber="522" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
; 34–35— 
<taxonomicName id="4C2C4D61682FFFED4AD761DDFAC7F88A" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1229,1338,1778,1802]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="522" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF0682FFFED4AD761DDFAC7F88A" box="[1229,1338,1778,1802]" italics="true" pageId="12" pageNumber="522">A. flavipes</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682FFFED4B5861DDFA68F88A" bold="true" box="[1346,1429,1778,1802]" pageId="12" pageNumber="522">
<taxonomicNameLabel id="A26B578B682FFFED4B5861DDFA68F88A" box="[1346,1429,1778,1802]" pageId="12" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 36— 
<taxonomicName id="4C2C4D61682FFFED4ED06039FECBF8AE" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[202,310,1814,1838]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="522" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682FFFED4ED06039FECBF8AE" box="[202,310,1814,1838]" italics="true" pageId="12" pageNumber="522">A. joceliae</emphasis>
</taxonomicName>
<emphasis id="B958EAF0682FFFED4F266039FE71F8AE" bold="true" box="[316,396,1814,1838]" pageId="12" pageNumber="522">
<taxonomicNameLabel id="A26B578B682FFFED4F266039FE71F8AE" box="[316,396,1814,1838]" pageId="12" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Lettering: ar—apical receptacle, dd—distal duct of spermatheca, df—dorsal flange, dil—spermathecal dilation, div—distal invagination of spermathecal duct, ip—intermediate part, pd—proximal duct of spermatheca, pf—proximal flange, rs—ring sclerite, so—spermathecal opening. Scale bars: 1 mm. Orig. P. Kment (33, 36), Salini (34–35).
</paragraph>
</caption>
<subSubSection id="C3366569682EFFEC4EDD67B9FEBCFC25" pageId="13" pageNumber="523" type="diagnosis">
<paragraph id="8B9336E2682EFFEC4EDD67B9FD2DFEE5" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
<emphasis id="B958EAF0682EFFEC4EDD67B9FE36FF30" bold="true" box="[199,459,150,176]" pageId="13" pageNumber="523">Differential diagnosis.</emphasis>
The genus 
<taxonomicName id="4C2C4D61682EFFEC4C5E67B8FD3DFF31" authorityName=": Atkinson" authorityYear="1888" box="[580,704,151,177]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C5E67B8FD3DFF31" box="[580,704,151,177]" italics="true" pageId="13" pageNumber="523">Agathocles</emphasis>
</taxonomicName>
is well recognizable with the following diagnostic characters: large (body length 
<quantity id="4CD49B07682EFFEC4F356794FE50FF55" box="[303,429,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.4" metricValueMin="1.3" pageId="13" pageNumber="523" unit="mm" value="18.5" valueMax="24.0" valueMin="13.0">13–24 mm</quantity>
), with robust body, widest across abdominal segment III; head only slightly broader than long with mandibular plates explanate and distinctly longer than clypeus, meeting or not in front of it; anterolateral margins of pronotum crenulate to slightly serrate; mesosternum carinate; labium reaching mesocoxae or slightly beyond, at most to metacoxae; all femora unarmed; peritreme spout-shaped, well developed; and abdominal sternites convex regularly without any tubercles or furrows.
</paragraph>
<paragraph id="8B9336E2682EFFEC4EDD6640FF22FD99" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
Among SE Asian genera of 
<taxonomicName id="4C2C4D61682EFFEC4FE76640FD9AFE09" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[509,615,367,393]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="554" phylum="Arthropoda" rank="tribe" tribe="Caystrini">Caystrini</taxonomicName>
, 
<taxonomicName id="4C2C4D61682EFFEC4C6B6640FD10FE09" authorityName=": Atkinson" authorityYear="1888" box="[625,749,367,393]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C6B6640FD10FE09" box="[625,749,367,393]" italics="true" pageId="13" pageNumber="523">Agathocles</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C2C4D61682EFFEC4DA06640FBE7FE09" authorityName="Distant" authorityYear="1901" box="[954,1050,367,393]" class="Insecta" genus="Valescus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4DA06640FBE7FE09" box="[954,1050,367,393]" italics="true" pageId="13" pageNumber="523">Valescus</emphasis>
</taxonomicName>
(also having anterolateral margins of pronotum crenulate; 
<quantity id="4CD49B07682EFFEC4F8566BCFDE4FE2D" box="[415,537,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="13" pageNumber="523" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
) and 
<taxonomicName id="4C2C4D61682EFFEC4C4D66BCFD18FE2C" authorityName="Distant" authorityYear="1901" box="[599,741,403,428]" class="Insecta" family="Pentatomidae" genus="Praetextatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C4D66BCFD18FE2C" box="[599,741,403,428]" italics="true" pageId="13" pageNumber="523">Praetextatus</emphasis>
</taxonomicName>
(anterolateral margins of pronotum smooth, 
<quantity id="4CD49B07682EFFEC4AC166BCFABBFE2E" box="[1243,1350,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="13" pageNumber="523" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
)—both are smaller, their scape does not reach apex of head, and mesosternum is furrowed (cf. 
<bibRefCitation id="EFBD4B13682EFFEC4A626698FAF1FE51" author="Distant, W. L." box="[1144,1292,439,465]" pageId="13" pageNumber="523" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant 1902</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4B036698FF2FFE75" author="Hsiao, T. - Y. &amp; Zheng, S. - Z." pageId="13" pageNumber="523" pagination="53 - 158" refId="ref30844" refString="Hsiao, T. - Y. &amp; Zheng, S. - Z. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera - Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158. [in Chinese with English summary]." type="book chapter" year="1977">
Hsiao 
<emphasis id="B958EAF0682EFFEC4B796698FA60FE51" box="[1379,1437,439,465]" italics="true" pageId="13" pageNumber="523">et al.</emphasis>
1977
</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4EC466F4FE25FE75" author="Ahmad, I. &amp; Afzal, M." box="[222,472,475,501]" pageId="13" pageNumber="523" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal 1979</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4FFC66F4FD98FE75" author="Chen, F. - Y." box="[486,613,475,501]" pageId="13" pageNumber="523" pagination="43 - 46" refId="ref29207" refString="Chen, F. - Y. (1983) New species of Pentatomidae from Guizhou, China (Hemiptera: Pentatomidae). Entomotaxonomia, 5 (1), 43 - 46." type="journal article" year="1983">Chen 1983</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4C6B66F4FC44FE75" author="Ahmad, I. &amp; Kamaluddin, S." box="[625,953,475,501]" pageId="13" pageNumber="523" pagination="169 - 183" refId="ref28067" refString="Ahmad, I. &amp; Kamaluddin, S. (1989) A revision of the tribe Caystrini (Hemiptera: Pentatomidae: Pentatominae) from Indo- Pakistan subcontinent with description of two new species from Pakistan and their cladistic analysis. Proceedings of the Pakistan Congress of Zoology, 9, 169 - 183." type="journal article" year="1989">Ahmad &amp; Kamaluddin 1989</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4DDF66F4FB2FFE75" author="Ahmad, I. &amp; Zahid, M." box="[965,1234,475,501]" pageId="13" pageNumber="523" pagination="1 - 5" refId="ref28233" refString="Ahmad, I. &amp; Zahid, M. (2006 b) Redescription of the caysrine genus Praetextatus Distant along with its type species P. typicus Distant (Hemiptera: Pentatomidae) with references to their unknown characters of metathoracic scent auricle and genitalia and key to its known species and their relationships. International Journal of Biology and Biotechnology, 3 (1), 1 - 5." type="journal article" year="2006">Ahmad &amp; Zahid 2006b</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4AC566F4FA9BFE75" author="Zahid, M." box="[1247,1382,475,501]" pageId="13" pageNumber="523" refId="ref34295" refString="Zahid, M. (2006) A revision of the tribes Caystrini Stal and Myrocheini Stal (Heteroptera: Pentatomidae: Pentatominae) with a revision of their type genera Caystrus Stal and Myrochea Stal of the World with special reference to their cladistic analysis. Unpublished Ph. D. Thesis. Department of Zoology, University of Karachi, Karachi, xxii + 464 pp." type="book" year="2006">Zahid 2006</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4B6866F4FF2EFD99" author="Fan, Z. - H." pageId="13" pageNumber="523" refId="ref29913" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Unpublished Ph. D., Nankai University, Tianjin, 614 pp." type="book" year="2011">Fan 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8B9336E2682EFFEC4EDD650CFD9BFD06" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
The enigmatic genus 
<taxonomicName id="4C2C4D61682EFFEC4FB5650CFDA7FDBC" authorityName="sensu Stal" authorityYear="1854" box="[431,602,547,572]" class="Insecta" family="Pentatomidae" genus="Paramecocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4FB5650CFDA7FDBC" box="[431,602,547,572]" italics="true" pageId="13" pageNumber="523">Paramecocoris</emphasis>
</taxonomicName>
(body length 6 linien = 
<quantity id="4CD49B07682EFFEC4D40650CFC30FDBD" box="[858,973,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.317" pageId="13" pageNumber="523" unit="mm" value="13.17">13.17 mm</quantity>
), known only from its original description, differs in having the head longer than wide and basipedicellite (IIa) as long as antennomeres III and IV combined (
<bibRefCitation id="EFBD4B13682EFFEC4E856544FED7FD05" author="Fieber, F. X." box="[159,298,618,645]" pageId="13" pageNumber="523" pagination="426 - 488" refId="ref29953" refString="Fieber, F. X. (1851) Rhynchotographien. Calve, Pragae, 64 pp. [preprint from Abhandlungen der Bohmischen Gesellschaft der Wissenschaften, Funfter Folge, 7, 426 - 488 (1852)]" type="book chapter" year="1851">Fieber 1851</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4F2C6544FE34FD05" author="Distant, W. L." box="[310,457,619,645]" pageId="13" pageNumber="523" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant 1902</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4FCF6544FDA7FD06" author="Zahid, M." box="[469,602,619,646]" pageId="13" pageNumber="523" refId="ref34295" refString="Zahid, M. (2006) A revision of the tribes Caystrini Stal and Myrocheini Stal (Heteroptera: Pentatomidae: Pentatominae) with a revision of their type genera Caystrus Stal and Myrochea Stal of the World with special reference to their cladistic analysis. Unpublished Ph. D. Thesis. Department of Zoology, University of Karachi, Karachi, xxii + 464 pp." type="book" year="2006">Zahid 2006</bibRefCitation>
).
</paragraph>
<paragraph id="8B9336E2682EFFEC4EDD65A0FABDFCB9" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
The remaining genera differ in having anterolateral margins of pronotum smooth and heads distinctly wider than long with mandibular plates not explanate, continually narrowing in front of eyes, as long as or only slightly longer than clypeus, clypeus apically free in 
<taxonomicName id="4C2C4D61682EFFEC4C5965F8FD3AFD70" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[579,711,727,752]" class="Insecta" family="Pentatomidae" genus="Heissocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C5965F8FD3AFD70" box="[579,711,727,752]" italics="true" pageId="13" pageNumber="523">Heissocoris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2C4D61682EFFEC4CCC65F8FC88FD70" authorityName="Ahmad &amp; Zahid" authorityYear="2012" box="[726,885,727,752]" class="Insecta" family="Pentatomidae" genus="Neoriazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4CCC65F8FC88FD70" box="[726,885,727,752]" italics="true" pageId="13" pageNumber="523">Neoriazocoris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2C4D61682EFFEC4D9E65F8FC17FD71" authorityName="Distant" authorityYear="1900" box="[900,1002,727,753]" class="Insecta" genus="Parodius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4D9E65F8FC17FD71" box="[900,1002,727,753]" italics="true" pageId="13" pageNumber="523">Parodius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2C4D61682EFFEC4A3865F8FB67FD70" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[1058,1178,727,752]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4A3865F8FB67FD70" box="[1058,1178,727,752]" italics="true" pageId="13" pageNumber="523">Riazocoris</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBD4B13682EFFEC4AB365F8FAC2FD71" author="Distant, W. L." box="[1193,1343,727,753]" pageId="13" pageNumber="523" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant 1902</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4B5065F8FEC9FC95" author="Ahmad, I. &amp; Afzal, M." pageId="13" pageNumber="523" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal 1979</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4F2465D4FD7FFC95" author="Ahmad, I. &amp; Kamaluddin, S." box="[318,642,763,789]" pageId="13" pageNumber="523" pagination="169 - 183" refId="ref28067" refString="Ahmad, I. &amp; Kamaluddin, S. (1989) A revision of the tribe Caystrini (Hemiptera: Pentatomidae: Pentatominae) from Indo- Pakistan subcontinent with description of two new species from Pakistan and their cladistic analysis. Proceedings of the Pakistan Congress of Zoology, 9, 169 - 183." type="journal article" year="1989">Ahmad &amp; Kamaluddin 1989</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4C9465D4FCEEFC95" author="Zahid, M." box="[654,787,763,789]" pageId="13" pageNumber="523" refId="ref34295" refString="Zahid, M. (2006) A revision of the tribes Caystrini Stal and Myrocheini Stal (Heteroptera: Pentatomidae: Pentatominae) with a revision of their type genera Caystrus Stal and Myrochea Stal of the World with special reference to their cladistic analysis. Unpublished Ph. D. Thesis. Department of Zoology, University of Karachi, Karachi, xxii + 464 pp." type="book" year="2006">Zahid 2006</bibRefCitation>
; Ahmad &amp; 
<bibRefCitation id="EFBD4B13682EFFEC4D8E65D4FBEAFC95" author="Zahid, M. &amp; Ahmad, I." box="[916,1047,763,789]" pageId="13" pageNumber="523" pagination="1653 - 1656" refId="ref34733" refString="Zahid, M. &amp; Ahmad, I. (2012) Metathoracic scent auricle and genitalia of pentatomine stink bug Caystrus pallipes (Stal) (Hemiptera: Pentatomidae) and their bearing on its phylogenetic relationship. Pakistan Journal of Zoology, 44 (6), 1653 - 1656." type="journal article" year="2012">Zahid 2012</bibRefCitation>
) or closed by mandibular plates in 
<taxonomicName id="4C2C4D61682EFFEC4E8D6430FF06FCB9" authorityName="Stal" authorityYear="1861" box="[151,251,799,825]" class="Insecta" family="Pentatomidae" genus="Caystrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4E8D6430FF06FCB9" box="[151,251,799,825]" italics="true" pageId="13" pageNumber="523">Caystrus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2C4D61682EFFEC4F2B6430FE4AFCB8" authorityName="Bergroth" authorityYear="1906" box="[305,439,799,824]" class="Insecta" genus="Hippotiscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4F2B6430FE4AFCB8" box="[305,439,799,824]" italics="true" pageId="13" pageNumber="523">Hippotiscus</emphasis>
</taxonomicName>
(e.g., 
<bibRefCitation id="EFBD4B13682EFFEC4FE46430FD6CFCB9" author="Distant, W. L." box="[510,657,799,825]" pageId="13" pageNumber="523" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant 1902</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4C816430FC5CFCB9" author="Ahmad, I. &amp; Zahid, M." box="[667,929,799,825]" pageId="13" pageNumber="523" pagination="271 - 275" refId="ref28178" refString="Ahmad, I. &amp; Zahid, M. (2006 a) Redescription of a serious caystrine stink bug bamboo pest Hippotiscus dorsalis Stal (Hemiptera: Pentatomidae) along with its genus and their phylogenetic relationships. International Journal of Biology and Biotechnology, 3 (2), 271 - 275." type="journal article" year="2006">Ahmad &amp; Zahid 2006a</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4DB76430FBCCFCB9" author="Zahid, M." box="[941,1073,799,825]" pageId="13" pageNumber="523" refId="ref34295" refString="Zahid, M. (2006) A revision of the tribes Caystrini Stal and Myrocheini Stal (Heteroptera: Pentatomidae: Pentatominae) with a revision of their type genera Caystrus Stal and Myrochea Stal of the World with special reference to their cladistic analysis. Unpublished Ph. D. Thesis. Department of Zoology, University of Karachi, Karachi, xxii + 464 pp." type="book" year="2006">Zahid 2006</bibRefCitation>
; 
<bibRefCitation id="EFBD4B13682EFFEC4A266430FAC8FCB9" author="Zahid, M. &amp; Ahmad, I." box="[1084,1333,799,825]" pageId="13" pageNumber="523" pagination="33 - 40" refId="ref34656" refString="Zahid, M. &amp; Ahmad, I. (2011) A review of caystrine subgroup Caystrus langei Breddin (1899) with special reference to the species described by Linnavuori i. e., C. hipponax, C. niokanus and C. pseudobrunnescens (Hemiptera: Pentatomidae: Pentatominae) from Ethiopian and Palaearctic Regions. International Journal of Biology and Biotechnology, 8 (1), 33 - 40." type="journal article" year="2011">Zahid &amp; Ahmad 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8B9336E2682EFFEC4EDD646CFEBCFC25" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
The 
<taxonomicName id="4C2C4D61682EFFEC4EE3646CFE6CFCDD" box="[249,401,835,861]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="511" phylum="Arthropoda" rank="tribe" tribe="Rolstoniellini">Rolstoniellini</taxonomicName>
genera 
<taxonomicName id="4C2C4D61682EFFEC4FF2646CFDA0FCDD" authorityName="Distant" authorityYear="1902" box="[488,605,835,861]" class="Insecta" genus="Exithemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4FF2646CFDA0FCDD" box="[488,605,835,861]" italics="true" pageId="13" pageNumber="523">Exithemus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2C4D61682EFFEC4C88646CFD0FFCDD" authorityName="Stal" authorityYear="1868" box="[658,754,835,861]" class="Insecta" genus="Critheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C88646CFD0FFCDD" box="[658,754,835,861]" italics="true" pageId="13" pageNumber="523">Critheus</emphasis>
</taxonomicName>
differs markedly by laterally prolonged and narrowing, rugashaped peritreme, while species of 
<taxonomicName id="4C2C4D61682EFFEC4C366448FD43FC01" authorityName="Rider" authorityYear="1997" box="[556,702,871,897]" class="Insecta" family="Pentatomidae" genus="Rolstoniellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4C366448FD43FC01" box="[556,702,871,897]" italics="true" pageId="13" pageNumber="523">Rolstoniellus</emphasis>
</taxonomicName>
differs by conspicuously produced humeral angles of pronotum (
<bibRefCitation id="EFBD4B13682EFFEC4E8564A4FEC9FC25" author="Distant, W. L." box="[159,308,907,933]" pageId="13" pageNumber="523" pagination="1 - 438" refId="ref29587" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Series Including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xxxvii + 1 - 438." type="book chapter" year="1902">Distant 1902</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366569682EFFEC4EDD6480FAAFFBDA" pageId="13" pageNumber="523" type="etymology">
<paragraph id="8B9336E2682EFFEC4EDD6480FAB3FC6D" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
<emphasis id="B958EAF0682EFFEC4EDD6480FEB3FC49" bold="true" box="[199,334,943,969]" pageId="13" pageNumber="523">Etymology.</emphasis>
<taxonomicName id="4C2C4D61682EFFEC4F4F6480FE2CFC49" authorityName=": Atkinson" authorityYear="1888" box="[341,465,943,969]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4F4F6480FE2CFC49" box="[341,465,943,969]" italics="true" pageId="13" pageNumber="523">Agathocles</emphasis>
</taxonomicName>
(= 
<emphasis id="B958EAF0682EFFEC4FE36480FD8FFC49" box="[505,626,943,969]" italics="true" pageId="13" pageNumber="523">ἈγαθΟΚλῆς</emphasis>
) is a Greek name, e.g. of several ancient Greek philosophers and rulers. The name is derived from 
<emphasis id="B958EAF0682EFFEC4FA564FCFDE5FC6D" box="[447,536,979,1005]" italics="true" pageId="13" pageNumber="523">agathos</emphasis>
(= 
<emphasis id="B958EAF0682EFFEC4C2564FDFD72FC6D" box="[575,655,978,1005]" italics="true" pageId="13" pageNumber="523">ἀγαθός</emphasis>
) meaning good and 
<emphasis id="B958EAF0682EFFEC4D6F64FCFC52FC6D" box="[885,943,979,1005]" italics="true" pageId="13" pageNumber="523">kleos</emphasis>
(= 
<emphasis id="B958EAF0682EFFEC4DCC64FDFBE9FC6D" box="[982,1044,978,1005]" italics="true" pageId="13" pageNumber="523">ΚλέΟς</emphasis>
) meaning glory; masculine.
</paragraph>
<paragraph id="8B9336E2682EFFEC4EDD64D8FAAFFBDA" blockId="13.[151,1437,150,1114]" pageId="13" pageNumber="523">
Neither the etymology nor the gender of the name 
<taxonomicName id="4C2C4D61682EFFEC4CE764D8FCABFB91" authorityName=": Distant" authorityYear="1902" box="[765,854,1015,1041]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4CE764D8FCABFB91" box="[765,854,1015,1041]" italics="true" pageId="13" pageNumber="523">Surenus</emphasis>
</taxonomicName>
were explicitly given in the original description. As we failed to trace it in the Latin or Greek dictionaries, the name must be considered masculine according to the Article 30.2.4 of 
<bibRefCitation id="EFBD4B13682EFFEC4F196310FE65FBD9" author="ICZN" box="[259,408,1087,1113]" pageId="13" pageNumber="523" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
. The name is also a hemihomonym with the plant genus 
<taxonomicName id="4C2C4D61682EFFEC4A0E6310FB90FBD9" authorityName=": Distant" authorityYear="1902" box="[1044,1133,1087,1113]" class="Insecta" family="Pentatomidae" genus="Surenus" kingdom="Plantae" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4A0E6310FB90FBD9" box="[1044,1133,1087,1113]" italics="true" pageId="13" pageNumber="523">Surenus</emphasis>
</taxonomicName>
(family 
<taxonomicName id="4C2C4D61682EFFEC4AD46310FABBFBDA" box="[1230,1350,1087,1114]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="523" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366569682EFFEC4E8D6384FE09FB45" box="[151,500,1194,1221]" pageId="13" pageNumber="523" type="multiple">
<paragraph id="8B9336E2682EFFEC4E8D6384FE09FB45" blockId="13.[151,500,1194,1221]" box="[151,500,1194,1221]" pageId="13" pageNumber="523">
<heading id="D0DB818E682EFFEC4E8D6384FE09FB45" bold="true" box="[151,500,1194,1221]" fontSize="11" level="1" pageId="13" pageNumber="523" reason="1">
<emphasis id="B958EAF0682EFFEC4E8D6384FE09FB45" bold="true" box="[151,500,1194,1221]" pageId="13" pageNumber="523">
Key to species of 
<taxonomicName id="4C2C4D61682EFFEC4F746385FE09FB45" authorityName=": Atkinson" authorityYear="1888" box="[366,500,1194,1221]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF0682EFFEC4F746385FE09FB45" bold="true" box="[366,500,1194,1221]" italics="true" pageId="13" pageNumber="523">Agathocles</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3366569682EFFEF4E8D63DFFA61FF67" lastPageId="14" lastPageNumber="524" pageId="13" pageNumber="523" type="key">
<keyStep id="30D82E47682EFFEC4E8D63DFFA61F98D" pageId="13" pageNumber="523">
<paragraph id="8B9336E2682EFFEC4E8D63DFFA61FA17" blockId="13.[151,1437,1263,2019]" pageId="13" pageNumber="523">
<keyLead id="30DD95D7682EFFEC4E8D63DFFA61FA17" pageId="13" pageNumber="523">
1 Small species (body length 
<quantity id="4CD49B07682EFFEC4FEE63C0FD78FA85" box="[500,645,1263,1285]" metricMagnitude="-2" metricUnit="m" metricValue="1.455" metricValueMax="1.55" metricValueMin="1.3599999999999999" pageId="13" pageNumber="523" unit="mm" value="14.55" valueMax="15.5" valueMin="13.6">13.6–15.5 mm</quantity>
). Body dorsally and ventrally with uniform dark brown colouration with yellow callosities instead of patterns (
<figureCitation id="13172A67682EFFEC4C166222FD82FAA3" box="[524,639,1293,1315]" captionStart="FIGURES 60–63" captionStartId="21.[151,264,1954,1978]" captionTargetBox="[205,1389,192,1928]" captionTargetId="figure-29@21.[195,1393,181,1930]" captionTargetPageId="21" captionText="FIGURES 60–63. Agathocles flavipes sp. nov., habitus. 60–61—male (60—dorsal view, 61—ventral view); 62–63—female (62—dorsal view, 63—ventral view). Scale bars: 1 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692395" httpUri="https://zenodo.org/record/4692395/files/figure.png" pageId="13" pageNumber="523">Figs 60–63</figureCitation>
). Head shorter (head length: width ratio 0.73), continually narrowing in front of eyes (
<figureCitation id="13172A67682EFFEC4F006205FE9CFAC0" box="[282,353,1322,1344]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="13" pageNumber="523">Fig. 10</figureCitation>
). Scape (I) surpassing apex of head about half of its length (
<figureCitation id="13172A67682EFFEC4DBA6205FC15FAC0" box="[928,1000,1322,1344]" captionStart="FIGURES 64–68" captionStartId="22.[151,264,988,1012]" captionTargetBox="[157,1436,181,964]" captionTargetId="figure-19@22.[151,1436,181,964]" captionTargetPageId="22" captionText="FIGURES 64–68. Agathocles flavipes sp. nov. 64—head and pronotum; 65—membrane with longitudinal venation; 66—apex of male abdomen with genital capsule in postero-ventral view; 67—apex of female abdomen with external genitalia in posteroventral view; 68—external scent efferent system. Lettering: pe—peritreme, sc—scape. Scale bars: 1 mm Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692397" httpUri="https://zenodo.org/record/4692397/files/figure.png" pageId="13" pageNumber="523">Fig. 64</figureCitation>
). Basipedicellite (IIa) short, about half length of distipedicellite (IIb). Genital capsule lacking superior processes of dorsal rim; ventral rim infolding with distinct median process (
<figureCitation id="13172A67682EFFEC4F2D624AFE5BFAFB" box="[311,422,1381,1403]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="13" pageNumber="523">Figs 71, 74</figureCitation>
). Apical process of paramere hook-shaped, mesal angle with additional projection (
<figureCitation id="13172A67682EFFEC4ADC624AFACBFAFB" box="[1222,1334,1381,1403]" captionStart="FIGURES 69–80" captionStartId="23.[151,264,1829,1853]" captionTargetBox="[225,1358,187,1799]" captionTargetId="figure-29@23.[222,1366,181,1805]" captionTargetPageId="23" captionText="FIGURES 69–80. Male genitalia of Agathocles flavipes sp. nov.: 69–74—genital capsule: 69, 72—dorsal view; 70, 73— ventral view; 71, 74—caudal view. 75–77—paramere (75, 77—lateral views, 76—ventral view). 78–80—penis (78—ventral view, 79—dorsal view, 80—lateral view). Lettering: ad—aedeagus, ap—apical process of paramere, ba—basal apodeme, bp— basal process of paramere, dlc—dorsolateral conjunctival processes, dr—dorsal rim, map—medial angle of parameral crown, mp—medial process of ventral rim, pa—processes of aedeagus, ps—paramere socket, pt—phallotheca, st—stem of paramere, vr—ventral rim. Scale bars: 0.2 mm (Figs 75–80), 0.5 mm (Figs 69–74). Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692399" httpUri="https://zenodo.org/record/4692399/files/figure.png" pageId="13" pageNumber="523">Figs 75–77</figureCitation>
, 
<figureCitation id="13172A67682EFFEC4B25624AFA80FAFB" box="[1343,1405,1381,1403]" captionStart="FIGURES 81–87" captionStartId="24.[151,264,1119,1143]" captionTargetBox="[158,1430,189,1092]" captionTargetId="figure-19@24.[151,1436,181,1095]" captionTargetPageId="24" captionText="FIGURES 81–87. Male genitalia of Agathocles flavipes sp. nov. 81—proctiger; 82–83—articulatory apparatus (82—dorsal view, 83—ventral view). 84–87—paramere, lateral view (arrow showing the variation in the mesal angle). Lettering: as—anal segment; at—anal tube; bpt—basal plate; cp—capitate process; dc—dorsal connective; hi—hinge; ism—intersternal membrane of segments IX‒X; map—medial angle of parameral crown; sbc—support bridge complex. Scale bars: 0.2 mm. Orig. Salini." figureDoi="http://doi.org/10.5281/zenodo.4692401" httpUri="https://zenodo.org/record/4692401/files/figure.png" pageId="13" pageNumber="523">84–87</figureCitation>
)....................................................................................... 
<emphasis id="B958EAF0682EFFEC4AF862ADFA61FA17" bold="true" box="[1250,1436,1410,1432]" pageId="13" pageNumber="523">
<taxonomicName id="4C2C4D61682EFFEC4AF862ADFAB7FA18" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1250,1354,1410,1432]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B958EAF0682EFFEC4AF862ADFAB7FA18" bold="true" box="[1250,1354,1410,1432]" italics="true" pageId="13" pageNumber="523">A. flavipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B578B682EFFEC4B4A62ADFA61FA17" box="[1360,1436,1410,1431]" pageId="13" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9336E2682EFFEC4E8D628FFA61F98D" blockId="13.[151,1437,1263,2019]" pageId="13" pageNumber="523">
<keyLead id="30DD95D7682EFFEC4E8D628FFA61F98D" pageId="13" pageNumber="523">
– Large species (
<quantity id="4CD49B07682EFFEC4F69628FFDF8FA35" box="[371,517,1440,1461]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.42" metricValueMin="1.88" pageId="13" pageNumber="523" unit="mm" value="21.5" valueMax="24.2" valueMin="18.8">18.8–24.2 mm</quantity>
). Body dorsally and ventrally with dark brown to blackish colouration with yellow patterns on scutellum or connexivum. Head longer (head length: width ratio 0.79–0.90), parallel-sided anteapically. Basipedicellite (IIa) long, only slightly shorter than distipedicellite (IIb). Genital capsule with superior processes of dorsal rim developed; ventral rim infolding without median process..................................................................... 2
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</paragraph>
</keyStep>
<keyStep id="30D82E47682EFFEC4E8D613AFA61F8B2" pageId="13" pageNumber="523">
<paragraph id="8B9336E2682EFFEC4E8D613AFA61F95B" blockId="13.[151,1437,1263,2019]" pageId="13" pageNumber="523">
<keyLead id="30DD95D7682EFFEC4E8D613AFA61F95B" pageId="13" pageNumber="523">
2 Connexivum ventrally, sometimes also dorsally, yellow to orange; scutellum unicolorous except very small and scattered yellowish callosities (
<figureCitation id="13172A67682EFFEC4FA1611DFDEBF9C8" box="[443,534,1586,1608]" captionStart="FIGURES 1–5" captionStartId="7.[151,264,1943,1967]" captionTargetBox="[203,1384,183,1912]" captionTargetId="figure-29@7.[196,1392,179,1917]" captionTargetPageId="7" captionText="FIGURES 1–5. Agathocles limbatus Stål, 1876. 1–2—habitus of male from Laos: Ban Saluey → Phou Pane Mt. (22.2 mm). 3–5—habitus and labels of holotype of A. yunnanensis Zhang &amp; Lin, 1984, syn. nov., female, China: Yunnan: Cang-yuan (scale bar 1 mm). Orig. P. Kment (1–2), D. Rédei (3–5)." figureDoi="http://doi.org/10.5281/zenodo.4692373" httpUri="https://zenodo.org/record/4692373/files/figure.png" pageId="13" pageNumber="523">Figs 1–4</figureCitation>
). Scape reaching about apex of head (
<figureCitation id="13172A67682EFFEC4DBB611DFC1DF9C8" box="[929,992,1586,1608]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="13" pageNumber="523">Fig. 6</figureCitation>
). Genital capsule with subquadrate superior processes of dorsal rim (
<figureCitation id="13172A67682EFFEC4FD5617FFD99F9E6" box="[463,612,1616,1638]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="13" pageNumber="523">Figs 37–39, 43</figureCitation>
); ventral rim shallowly bisinuate in ventral view (
<figureCitation id="13172A67682EFFEC4A5D617FFB6DF9E6" box="[1095,1168,1616,1638]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="13" pageNumber="523">Fig. 41</figureCitation>
); ventral rim infolding with large semicircular impression medially (
<figureCitation id="13172A67682EFFEC4C6B6142FD18F903" box="[625,741,1645,1667]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="13" pageNumber="523">Figs 37–39</figureCitation>
); ventral rim produced ventrally (see in lateral view, 
<figureCitation id="13172A67682EFFEC4AE56142FAB7F903" box="[1279,1354,1645,1667]" captionStart="FIGURES 37–43" captionStartId="15.[151,264,1722,1746]" captionTargetBox="[151,1436,181,1698]" captionTargetId="figure-29@15.[151,1436,181,1698]" captionTargetPageId="15" captionText="FIGURES 37–43. Male genital capsule of Agathocles limbatus Stål, 1876 (37, 40, 41—Laos: Phou Pane Mt.; 38, 42–43— Cambodia: Pailin env.; 39—China: Guangxi): 37—caudal view, not dissected (27×); 38–39—caudal view, dissected (27×); 40—lateral view (27×); 41—ventral view (19×); 42—proctiger (75×); 43—superior process of dorsal rim (110×). Lettering: pa –paramere, plp—posterolateral projection, ps—paramere socket, sp—superior process of dorsal rim, vif—ventral rim infolding, x—proctiger. Scale bars: 1 mm (37–41), 0.5 mm (42–43). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692387" httpUri="https://zenodo.org/record/4692387/files/figure.png" pageId="13" pageNumber="523">Fig. 40</figureCitation>
). Apical process of paramere triangular, mesal angle with additional projection (
<figureCitation id="13172A67682EFFEC4D8961A5FB6AF920" box="[915,1175,1674,1696]" captionStart="FIGURES 44–53" captionStartId="16.[151,264,1943,1967]" captionTargetBox="[154,1433,181,1919]" captionTargetId="figure-19@16.[154,1433,181,1919]" captionTargetPageId="16" captionText="FIGURES 44–53. Paramere of Agathocles limbatus Stål, 1876 from Cambodia: Pailin env. (44–46, 50, 52) and China: Guangxi (47–49, 51, 53) (five different views, 70×). Lettering: ap—apical process of paramere, bp—basal process of paramere, map— projection of mesal angle of paramere stem. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692391" httpUri="https://zenodo.org/record/4692391/files/figure.png" pageId="13" pageNumber="523">Figs 44, 46–47, 49, 52–53</figureCitation>
). Body length 
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................................................................................................................................................................................... 
<taxonomicName id="4C2C4D61682EFFEC4ADE61EAFA61F95B" authority="Stal, 1876" authorityName="Stal" authorityYear="1876" box="[1220,1436,1732,1755]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B958EAF0682EFFEC4ADE61EAFAC8F95A" bold="true" box="[1220,1333,1732,1754]" italics="true" pageId="13" pageNumber="523">A. limbatus</emphasis>
<bibRefCitation id="EFBD4B13682EFFEC4B2161EAFA61F95B" author="Stal, C." box="[1339,1436,1733,1755]" pageId="13" pageNumber="523" pagination="1 - 162" refId="ref33749" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9336E2682EFFEC4E8D61CCFA61F8B2" blockId="13.[151,1437,1263,2019]" pageId="13" pageNumber="523">
<keyLead id="30DD95D7682EFFEC4E8D61CCFA61F8B2" pageId="13" pageNumber="523">
– Connexivum dark, concolorous with surrounding abdominal terga and sterna; scutellum with yellowish spot on apex (
<figureCitation id="13172A67682EFFEC4B6861CDFF01F895" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="13" pageNumber="523">Figs 88</figureCitation>
, 
<figureCitation id="13172A67682EFFEC4F1C602FFEA0F896" box="[262,349,1792,1814]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="13" pageNumber="523">108, 110</figureCitation>
). Ventral rim of genital capsule with single, deep incision medially (
<figureCitation id="13172A67682EFFEC4DF7602FFBC5F895" box="[1005,1080,1792,1814]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="13" pageNumber="523">Figs 93</figureCitation>
, 
<figureCitation id="13172A67682EFFEC4A5861D0FB95F895" box="[1090,1128,1791,1813]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="13" pageNumber="523">115</figureCitation>
), not produced ventrally. Apical process of paramere hook-shaped, mesal angle without additional projection (
<figureCitation id="13172A67682EFFEC4DD16032FBBBF8B2" box="[971,1094,1821,1843]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="13" pageNumber="523">Figs 97–101</figureCitation>
, 
<figureCitation id="13172A67682EFFEC4A4A6032FB5BF8B3" box="[1104,1190,1821,1843]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="13" pageNumber="523">118–123</figureCitation>
)................... 3
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</paragraph>
</keyStep>
<paragraph id="8B9336E2682EFFEC4E8D6015FA61F846" blockId="13.[151,1437,1263,2019]" pageId="13" pageNumber="523">
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3 Scutellum with small, yellowish, punctate spot on apex (
<figureCitation id="13172A67682EFFEC4CE56015FC7AF8D0" box="[767,903,1850,1872]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="13" pageNumber="523">Figs 108, 110</figureCitation>
). Lateral pronotal margin anteriorly minutely crenulate. Scape about reaching apex of head (
<figureCitation id="13172A67682EFFEC4C4B6077FD55F8EE" box="[593,680,1880,1902]" captionStart="FIGURES 108–112" captionStartId="30.[151,264,1652,1676]" captionTargetBox="[153,1428,193,1628]" captionTargetId="figure-19@30.[151,1436,181,1628]" captionTargetPageId="30" captionText="FIGURES 108–112. Agathocles normalis (Distant, 1901), habitus. 108–109—male, Laos, Phou Pane Mt. (body length 18.78 mm) (108—dorsal view, 109—ventral view).110–112—holotype of Surenus normalis Distant, 1901, female (110—dorsal view, 111—lateral view, 112—labels). Orig. P. Kment (108–109), M. Tavano (110–112)." figureDoi="http://doi.org/10.5281/zenodo.4692411" httpUri="https://zenodo.org/record/4692411/files/figure.png" pageId="13" pageNumber="523">Fig. 108</figureCitation>
). Genital capsule with small, subtriangular superior processes of dorsal rim (
<figureCitation id="13172A67682EFFEC4EF3605AFE5DF80B" box="[233,416,1909,1931]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="13" pageNumber="523">Figs 113–114, 117</figureCitation>
); ventral rim infolding with large crescent-shaped impression medially (
<figureCitation id="13172A67682EFFEC4A78605AFB16F80B" box="[1122,1259,1909,1931]" captionStart="FIGURES 113–117" captionStartId="31.[151,264,896,920]" captionTargetBox="[306,1282,181,873]" captionTargetId="figure-29@31.[306,1282,181,873]" captionTargetPageId="31" captionText="FIGURES 113–117. Male genital capsule of Agathocles normalis (Distant, 1901): 113—caudal view (27×); 114—caudal view, dissected (32×); 115—ventral view (21×); 116—lateral view (27×); 117—superior process of dorsal rim (150×). Lettering: plp—posterolateral projection of genital capsule, sp—superior process of dorsal rim. Scale bars: 1 mm (113–116), 0.1 mm (117). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692413" httpUri="https://zenodo.org/record/4692413/files/figure.png" pageId="13" pageNumber="523">Figs 113–114</figureCitation>
). Basal process of paramere oriented more vertically, bearing row of many setae (
<figureCitation id="13172A67682EFFEC4D2760BDFC38F828" box="[829,965,1938,1960]" captionStart="FIGURES 118–124" captionStartId="31.[151,264,1924,1948]" captionTargetBox="[311,1277,1053,1901]" captionTargetId="figure-85@31.[311,1277,1053,1901]" captionTargetPageId="31" captionText="FIGURES 118–124. Paramere of Agathocles normalis (Distant, 1901), five different views (118–123, 100×) and detail of microsculpture (124, 150×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 1 mm (118–123), 0.25 mm (124). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692415" httpUri="https://zenodo.org/record/4692415/files/figure.png" pageId="13" pageNumber="523">Figs 118–123</figureCitation>
). Body length 18.80–20.00 mm.......................................................................................... 
<taxonomicName id="4C2C4D61682EFFEC4A9C609FFA61F846" authority="(Distant, 1901 a)" authorityName="Salini &amp; Kment" authorityYear="2021" baseAuthorityName="Distant" baseAuthorityYear="1901" box="[1158,1436,1967,1990]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="523" phylum="Arthropoda" rank="species" species="normalis">
<emphasis id="B958EAF0682EFFEC4A9C609FFB07F845" bold="true" box="[1158,1274,1967,1989]" italics="true" pageId="13" pageNumber="523">A. normalis</emphasis>
(
<bibRefCitation id="EFBD4B13682EFFEC4B1D609FFA68F846" author="Distant, W. L." box="[1287,1429,1968,1990]" pageId="13" pageNumber="523" pagination="99 - 114" refId="ref29459" refString="Distant, W. L. (1901 a) Enumeration of the Heteroptera (Rhynchota) collected by Signor Leonardo Fea in Burma and its vicinity. Part I. Family Pentatomidae. Transactions of the Entomological Society of London, 1901 (1), 99 - 114. https: // doi. org / 10.1111 / j. 1365 - 2311.1901. tb 02734. x" type="journal article" year="1901">Distant, 1901a</bibRefCitation>
)
</taxonomicName>
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</keyLead>
</paragraph>
<paragraph id="8B9336E2682EFFEF4E8D60E2FA61FF67" blockId="13.[151,1437,1263,2019]" lastBlockId="14.[226,1436,151,232]" lastPageId="14" lastPageNumber="524" pageId="13" pageNumber="523">
<keyLead id="30DD95D7682EFFEC4E8D60E2FA61F862" box="[151,1436,1997,2019]" pageId="13" pageNumber="523">
– Scutellum with large yellowish impunctate spot on apex (
<figureCitation id="13172A67682EFFEC4D0C60E2FC9DF863" box="[790,864,1997,2019]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="13" pageNumber="523">Fig. 88</figureCitation>
). Lateral pronotal margin anteriorly slightly serrate. Scape
</keyLead>
surpassing apex of head about half of its length (
<figureCitation id="13172A67682DFFEF4CB767B8FD0AFF2D" box="[685,759,151,173]" captionStart="FIGURES 6–13" captionStartId="8.[151,264,1811,1835]" captionTargetBox="[215,1372,181,1787]" captionTargetId="figure-19@8.[215,1372,181,1787]" captionTargetPageId="8" captionText="FIGURES 6–13. Head. 6–8—Agathocles limbatus Stål, 1876: 6—dorsal view, Vietnam: Phia Oac NP (magnification 20×); 7— dorsal view, Thailand: Phu Kradung NP (20×); 8—lateral view, Laos: Phou Pane Mt. (30×). 9—A. normalis (Distant, 1901a), male, dorsal view, Laos: Phou Pane Mt. (23× mm). 10–11—A. flavipes sp. nov., female, dorsal (10, 30×) and ventral (11, 40×) view, India: Maharashtra: 70 km N of Pune. 12–13—A. joceliae sp. nov.: 12—holotype, male, dorsal view, Malaysia: Kelantan (25×); 13—female, ventral view, Malaysia: Kelantan: Kampong Sungai Om (27×). Lettering: at—antenniferous tubercle, bu— buccula, cl—clypeus, la—labium, lr—labrum, mp—mandibular plate, oc—ocellus. Scale bars: 1 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692377" httpUri="https://zenodo.org/record/4692377/files/figure.png" pageId="14" pageNumber="524">Figs 12</figureCitation>
, 
<figureCitation id="13172A67682DFFEF4D1A67B7FCE6FF2E" box="[768,795,152,174]" captionStart="FIGURES 88–89" captionStartId="25.[151,264,1912,1936]" captionTargetBox="[157,1428,892,1887]" captionTargetId="figure-264@25.[151,1436,885,1889]" captionTargetPageId="25" captionText="FIGURES 88–89. Agathocles joceliae sp. nov., habitus of holotype, male, body length 19.27 mm (88—dorsal view, 89— ventral view). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692403" httpUri="https://zenodo.org/record/4692403/files/figure.png" pageId="14" pageNumber="524">88</figureCitation>
). Genital capsule with crescent-shaped superior processes of dorsal rim (
<figureCitation id="13172A67682DFFEF4F08679AFE54FF4B" box="[274,425,181,203]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="14" pageNumber="524">Figs 90–91, 96</figureCitation>
); ventral rim sharp, without apparent infolding (
<figureCitation id="13172A67682DFFEF4D9F679AFC05FF4B" box="[901,1016,181,203]" captionStart="FIGURES 90–96" captionStartId="26.[151,264,1564,1588]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@26.[151,1436,181,1541]" captionTargetPageId="26" captionText="FIGURES 90–96. Male genital capsule of Agathocles joceliae sp. nov.: 90—caudal view (42×); 91—caudal view, dissected (42×); 92—dorsal view (30×); 93—ventral view (27×); 94—lateral view (37×); 95—proctiger (85×); 96—superior process of dorsal rim (130×). Lettering: plp—posterolateral projection of genital capsule, ps—paramere socket, sp—superior process of dorsal rim. Scale bars: 1 mm (92–94), 0.5 mm (90–91, 95), 0.2 mm (96). Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692405" httpUri="https://zenodo.org/record/4692405/files/figure.png" pageId="14" pageNumber="524">Figs 90–91</figureCitation>
). Basal process of paramere oriented more horizontally, bearing group of few setae (
<figureCitation id="13172A67682DFFEF4C7767FDFD14FF68" box="[621,745,210,232]" captionStart="FIGURES 97–101" captionStartId="27.[151,264,1687,1711]" captionTargetBox="[151,1436,453,1664]" captionTargetId="figure-147@27.[151,1436,453,1664]" captionTargetPageId="27" captionText="FIGURES 97–101. Paramere of Agathocles joceliae sp. nov., five different views (100×). Lettering: ap—apical process of paramere, bp—basal process of paramere. Scale bars: 0.5 mm. Orig. P. Kment." figureDoi="http://doi.org/10.5281/zenodo.4692407" httpUri="https://zenodo.org/record/4692407/files/figure.png" pageId="14" pageNumber="524">Figs 97–101</figureCitation>
). Body length 
<quantity id="4CD49B07682DFFEF4D6C67FDFBE1FF68" box="[886,1052,210,232]" metricMagnitude="-2" metricUnit="m" metricValue="1.9939999999999998" metricValueMax="2.098" metricValueMin="1.89" pageId="14" pageNumber="524" unit="mm" value="19.939999999999998" valueMax="20.98" valueMin="18.9">18.90–20.98 mm</quantity>
................ 
<emphasis id="B958EAF0682DFFEF4AF867FDFA61FF67" bold="true" box="[1250,1436,210,232]" pageId="14" pageNumber="524">
<taxonomicName id="4C2C4D61682DFFEF4AF867FDFAB7FF68" authority="Salini &amp; Kment, 2021" authorityName="Salini &amp; Kment" authorityYear="2021" box="[1250,1354,210,232]" class="Insecta" genus="Agathocles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="524" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B958EAF0682DFFEF4AF867FDFAB7FF68" bold="true" box="[1250,1354,210,232]" italics="true" pageId="14" pageNumber="524">A. joceliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B578B682DFFEF4B4A67FDFA61FF67" box="[1360,1436,210,231]" pageId="14" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>