665 lines
89 KiB
XML
665 lines
89 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.5.5.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333659" ID-ZooBank="0EBF1D41-2E9F-4D79-96F0-2F4CBA07587A" approvalRequired="6" approvalRequired_for_textStreams="5" approvalRequired_for_treatments="1" checkinTime="1668766408349" checkinUser="plazi" docAuthor="BOTERO-TRUJILLO, RICARDO, DAVIS, STEVEN R., MICHALIK, PETER & PRENDINI, LORENZO" docDate="2022" docId="038587CDB642FFAA0E12588FFBF3FDC5" docLanguage="en" docName="palaeoentomology.5.5.11.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Hirsutisoma grimaldii BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="500" masterDocId="FFBCFFB5B640FFAD0D385A22FFC5FFA9" masterDocTitle="Hirsutisoma grimaldii sp. nov., a ca. 99 - million-year-old ricinuleid (Primoricinulei, Hirsutisomidae) from Cretaceous Burmese amber with a corticolous, scansorial lifestyle" masterLastPageNumber="504" masterPageNumber="493" pageNumber="495" updateTime="1668766848702" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Hirsutisoma grimaldii sp. nov., a ca. 99 - million-year-old ricinuleid (Primoricinulei, Hirsutisomidae) from Cretaceous Burmese amber with a corticolous, scansorial lifestyle</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>BOTERO-TRUJILLO, RICARDO</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>DAVIS, STEVEN R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>MICHALIK, PETER</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>PRENDINI, LORENZO</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>493</mods:start>
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<mods:end>504</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.5.11</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.5.5.11</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7333659</mods:identifier>
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<mods:identifier type="ZooBank">0EBF1D41-2E9F-4D79-96F0-2F4CBA07587A</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:038587CDB642FFAA0E12588FFBF3FDC5" httpUri="http://treatment.plazi.org/id/038587CDB642FFAA0E12588FFBF3FDC5" lastPageId="7" lastPageNumber="500" pageId="2" pageNumber="495">
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<subSubSection box="[810,1176,684,711]" pageId="2" pageNumber="495" type="nomenclature">
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<paragraph blockId="2.[810,1176,684,746]" box="[810,1176,684,711]" pageId="2" pageNumber="495">
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<heading bold="true" box="[810,1176,684,711]" fontSize="10" level="2" pageId="2" pageNumber="495" reason="3">
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<emphasis bold="true" box="[810,1176,684,711]" pageId="2" pageNumber="495">
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†
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<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[824,1080,684,711]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis bold="true" box="[824,1080,684,711]" italics="true" pageId="2" pageNumber="495">Hirsutisoma grimaldii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1087,1176,685,711]" pageId="2" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection box="[810,1062,720,746]" pageId="2" pageNumber="495" type="description">
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<paragraph blockId="2.[810,1176,684,746]" box="[810,1062,720,746]" pageId="2" pageNumber="495">
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(
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<figureCitation box="[818,919,720,746]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[136,235,2008,2032]" captionStartId-1="3.[136,235,1998,2022]" captionStartId-2="5.[136,235,1522,1546]" captionStartId-3="6.[136,235,1709,1733]" captionTargetBox-0="[151,1435,1091,1982]" captionTargetBox-1="[151,1435,1083,1974]" captionTargetBox-2="[180,1417,200,1486]" captionTargetBox-3="[223,1401,205,1672]" captionTargetId-0="figure-296@2.[151,1435,1091,1983]" captionTargetId-1="figure-398@3.[151,1435,1083,1975]" captionTargetId-2="figure-119@5.[151,1436,189,1498]" captionTargetId-3="figure-85@6.[151,1436,183,1685]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." captionText-1="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." captionText-2="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." captionText-3="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi-0="http://doi.org/10.5281/zenodo.7333661" figureDoi-1="http://doi.org/10.5281/zenodo.7333663" figureDoi-2="http://doi.org/10.5281/zenodo.7333665" figureDoi-3="http://doi.org/10.5281/zenodo.7333667" httpUri-0="https://zenodo.org/record/7333661/files/figure.png" httpUri-1="https://zenodo.org/record/7333663/files/figure.png" httpUri-2="https://zenodo.org/record/7333665/files/figure.png" httpUri-3="https://zenodo.org/record/7333667/files/figure.png" pageId="2" pageNumber="495">Figs 1–4</figureCitation>
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;
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<tableCitation box="[931,1024,720,746]" captionStart="TABLE 1" captionStartId="4.[136,213,158,179]" captionText="TABLE 1. Measurements (mm) for holotype of Hirsutisoma grimaldii sp. nov. deposited in the American Museum of Natural History, New York.Abbreviations:L, length; W, width; H, height. Excludes measurements that could not be reliably obtained due to position or preservation of structure. 1 excludes pygidium or cucullus; 2 maximum width; 3 maximum height; 4 midline." pageId="2" pageNumber="495">Tables 1</tableCitation>
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,
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<tableCitation box="[1038,1053,720,746]" captionStart="TABLE 2" captionStartId="9.[136,207,159,178]" captionText="TABLE 2. Taxonomic distribution of selected characters by sex and life stage in species of Primoricinulei Wunderlich, 2015; Hirsutisoma acutiformis Wunderlich, 2017; Hirsutisoma bruckschi Wunderlich, 2017; Hirsutisoma dentata Wunderlich, 2017; Hirsutisoma grimaldii sp. nov.; Primoricinuleus pugio Wunderlich, 2015. Observations based on literature and published images (Wunderlich, 2015, 2017), except for H. bruckschi and H. grimaldii, the holotypes of which were examined. Abbreviations: j, juvenile; N, no; Y, yes." pageId="2" pageNumber="495">2</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="495" type="materials_examined">
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<paragraph blockId="2.[810,1452,790,1064]" pageId="2" pageNumber="495">
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<emphasis bold="true" box="[810,990,790,817]" pageId="2" pageNumber="495">Type material.</emphasis>
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<materialsCitation collectionCode="AMNH" country="Myanmar" location="Cretaceous Burmese" pageId="2" pageNumber="495" specimenCode="IZC 00357137" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus box="[1003,1107,790,816]" pageId="2" pageNumber="495">Holotype</typeStatus>
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<specimenCount box="[1121,1142,791,817]" pageId="2" pageNumber="495" type="male">♂</specimenCount>
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(
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<collectionCode box="[1163,1250,791,817]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="495">AMNH</collectionCode>
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_
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<specimenCode box="[1264,1440,790,817]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="495">IZC 00357137</specimenCode>
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),
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<collectingCountry box="[810,923,826,852]" name="Myanmar" pageId="2" pageNumber="495">Myanmar</collectingCountry>
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: embedded in an oval piece (18 × 12 ×
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<quantity box="[1376,1443,825,852]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="495" unit="mm" value="5.0">5 mm</quantity>
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) of
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<location LSID="urn:lsid:plazi:treatment:038587CDB642FFAA0E12588FFBF3FDC5:8EF36000B642FFAF0E70597FFBEAFCDE" box="[840,1071,861,887]" country="Myanmar" name="Cretaceous Burmese" pageId="2" pageNumber="495">Cretaceous Burmese</location>
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amber (
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<emphasis box="[1164,1190,862,887]" italics="true" pageId="2" pageNumber="495">ca</emphasis>
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. 99 Ma).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="495" type="etymology">
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||
<paragraph blockId="2.[810,1452,790,1064]" pageId="2" pageNumber="495">
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||
<emphasis bold="true" box="[863,998,896,922]" pageId="2" pageNumber="495">Etymology.</emphasis>
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||
The specific epithet is a patronym honoring AMNH Curator, Dr David A. Grimaldi, in recognition of his work on extinct and extant arthropods. Dr Grimaldi’s assistance was instrumental in making this fossil available for study.
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||
</paragraph>
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||
</subSubSection>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333661" ID-Zenodo-Dep="7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="2" pageNumber="495" startId="2.[136,235,2008,2032]" targetBox="[151,1435,1091,1982]" targetPageId="2">
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<paragraph blockId="2.[136,1451,2008,2068]" pageId="2" pageNumber="495">
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<emphasis bold="true" box="[136,261,2008,2032]" pageId="2" pageNumber="495">FIGURE 1.</emphasis>
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<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[268,495,2008,2032]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="2" pageNumber="495" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis box="[268,495,2008,2032]" italics="true" pageId="2" pageNumber="495">Hirsutisoma grimaldii</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[503,584,2008,2032]" pageId="2" pageNumber="495">
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<taxonomicNameLabel box="[503,584,2008,2032]" pageId="2" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype ♂ (AMNH_IZC 00357137).
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<emphasis bold="true" box="[991,1009,2008,2032]" pageId="2" pageNumber="495">A</emphasis>
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and
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<emphasis bold="true" box="[1061,1078,2008,2032]" pageId="2" pageNumber="495">B</emphasis>
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, Habitus, dorsal (
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<emphasis bold="true" box="[1262,1280,2008,2032]" pageId="2" pageNumber="495">A</emphasis>
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) and ventral (
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<emphasis bold="true" box="[1426,1443,2008,2032]" pageId="2" pageNumber="495">B</emphasis>
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) aspects.
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<emphasis bold="true" box="[222,240,2044,2068]" pageId="2" pageNumber="495">C</emphasis>
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, Prosoma, anterodorsal aspect.
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<emphasis bold="true" box="[562,580,2044,2068]" pageId="2" pageNumber="495">D</emphasis>
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, Coxosternal region.
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</paragraph>
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</caption>
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<subSubSection pageId="3" pageNumber="496" type="diagnosis">
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<paragraph blockId="3.[136,778,158,1048]" pageId="3" pageNumber="496">
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<emphasis bold="true" box="[189,312,158,184]" pageId="3" pageNumber="496">Diagnosis.</emphasis>
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<taxonomicName authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[349,628,158,184]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="grimaldii">
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<emphasis box="[349,628,158,184]" italics="true" pageId="3" pageNumber="496">Hirsutisoma grimaldii</emphasis>
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</taxonomicName>
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may be recognized by the presence of a distinct comb of 8–10 lamellate projections basally on the proventral surface of the pedipalp tarsus (
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<figureCitation box="[401,523,266,292]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="3" pageNumber="496">Fig. 3D–F</figureCitation>
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), a structure absent in the
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<typeStatus box="[180,228,303,329]" pageId="3" pageNumber="496">type</typeStatus>
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specimens of other congeners. It differs further from
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<taxonomicName authorityName="Wunderlich" authorityYear="2017" box="[197,339,338,364]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="bruckschi">
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<emphasis box="[197,339,338,364]" italics="true" pageId="3" pageNumber="496">H. bruckschi</emphasis>
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||
</taxonomicName>
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, the only other
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<taxonomicName box="[513,739,338,364]" pageId="3" pageNumber="496">
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||
<emphasis box="[513,651,339,364]" italics="true" pageId="3" pageNumber="496">Hirsutisoma</emphasis>
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species
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</taxonomicName>
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for which the adult male is known, in the morphology of the male leg III and copulatory apparatus: in the male of
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<taxonomicName authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="grimaldii">
|
||
<emphasis box="[750,770,411,436]" italics="true" pageId="3" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis box="[136,240,446,472]" italics="true" pageId="3" pageNumber="496">grimaldii</emphasis>
|
||
</taxonomicName>
|
||
, the prolateral surface of the metatarsus of leg III is sharply expanded distally, the first tarsomere of leg III bears a pronounced longitudinal ridge retrodorsally, and the horn-shaped part of the copulatory apparatus is relatively long, reaching well past the midline of the first tarsomere (
|
||
<figureCitation box="[262,390,626,653]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="3" pageNumber="496">Figs 2C, D</figureCitation>
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||
,
|
||
<figureCitation box="[402,441,626,652]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="3" pageNumber="496">4D</figureCitation>
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||
). In the male of
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<taxonomicName authorityName="Wunderlich" authorityYear="2017" box="[627,771,626,652]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="bruckschi">
|
||
<emphasis box="[627,771,626,652]" italics="true" pageId="3" pageNumber="496">H. bruckschi</emphasis>
|
||
</taxonomicName>
|
||
, the prolateral surface of the metatarsus of leg III is not expanded distally, no ridge is present retrodorsally on the first tarsomere of leg III, and the horn-shaped part of the copulatory apparatus is shorter, reaching to the midline of the first tarsomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="500" pageId="3" pageNumber="496" type="description">
|
||
<paragraph blockId="3.[136,778,158,1048]" pageId="3" pageNumber="496">
|
||
<emphasis bold="true" box="[189,335,842,868]" pageId="3" pageNumber="496">Description.</emphasis>
|
||
Male. Based on
|
||
<typeStatus box="[520,618,842,868]" pageId="3" pageNumber="496">holotype</typeStatus>
|
||
(AMNH_IZC 00357137).
|
||
</paragraph>
|
||
<paragraph blockId="3.[136,778,158,1048]" pageId="3" pageNumber="496">
|
||
<emphasis box="[189,351,915,940]" italics="true" pageId="3" pageNumber="496">Measurements</emphasis>
|
||
. Total length, approximately
|
||
<quantity box="[679,778,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="3.91" pageId="3" pageNumber="496" unit="mm" value="3.91">3.91 mm</quantity>
|
||
(
|
||
<tableCitation box="[143,227,950,976]" captionStart="TABLE 1" captionStartId="4.[136,213,158,179]" captionText="TABLE 1. Measurements (mm) for holotype of Hirsutisoma grimaldii sp. nov. deposited in the American Museum of Natural History, New York.Abbreviations:L, length; W, width; H, height. Excludes measurements that could not be reliably obtained due to position or preservation of structure. 1 excludes pygidium or cucullus; 2 maximum width; 3 maximum height; 4 midline." pageId="3" pageNumber="496">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[136,778,158,1048]" lastBlockId="3.[810,1452,158,1050]" pageId="3" pageNumber="496">
|
||
<emphasis box="[189,293,986,1012]" italics="true" pageId="3" pageNumber="496">Inclusion</emphasis>
|
||
. Complete specimen. Ventral aspect completely visible (
|
||
<figureCitation box="[381,478,1022,1048]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="3" pageNumber="496">Fig. 1B</figureCitation>
|
||
), amber matrix without significant imperfections; pedipalp coxae partially obscured by debris (
|
||
<figureCitation box="[1038,1127,194,220]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="3" pageNumber="496">Fig. 1D</figureCitation>
|
||
). Dorsal aspect of specimen, especially carapace, partly obscured by bubbles (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="3" pageNumber="496">Fig. 1A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[810,1452,158,1050]" pageId="3" pageNumber="496">
|
||
<emphasis box="[863,1180,302,328]" italics="true" pageId="3" pageNumber="496">Colouration and hardness</emphasis>
|
||
. Cuticle pale brown, uniformly coloured. Pedipalp tarsus and copulatory apparatus reddish. Colour of specimen and apparent compression of some leg segments (by pressure exerted by amber), suggest cuticle may be not heavily sclerotized.
|
||
</paragraph>
|
||
<paragraph blockId="3.[810,1452,158,1050]" lastBlockId="4.[136,778,1696,2093]" lastPageId="4" lastPageNumber="497" pageId="3" pageNumber="496">
|
||
<emphasis box="[863,955,483,509]" italics="true" pageId="3" pageNumber="496">Setation</emphasis>
|
||
. Coxosternal region, opisthosoma dorsal surface (posterior half) and, especially, ventral surface covered with short to medium-sized, rather spinose macrosetae. Opisthosoma dorsal surface of anterior one- or two-thirds densely hirsute, long macrosetae (some about half opisthosomal length) obscuring anterior dorsal sclerites (
|
||
<figureCitation box="[1096,1193,699,725]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="3" pageNumber="496">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[1208,1244,699,725]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="3" pageNumber="496">3C</figureCitation>
|
||
); unclear whether macrosetae originate along anterodorsal external margin of opisthosoma or from surface of anterior tergites; external border of opisthosoma provisioned along entire length with dense array of medium-sized macrosetae (apparently originating from ventral sclerites).
|
||
<taxonomicName box="[1352,1452,879,905]" class="Gastropoda" family="Conidae" genus="Cucullus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="497" phylum="Cnidaria" rank="genus">Cucullus</taxonomicName>
|
||
and legs covered with bristle-like macrosetae of variable length (
|
||
<figureCitation box="[894,964,951,977]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="3" pageNumber="496">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation box="[978,994,951,977]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="3" pageNumber="496">2</figureCitation>
|
||
); leg III, dorsal surface of first tarsomere with brush-like array of brownish macrosetae (apparently clavate) distally (
|
||
<figureCitation box="[999,1117,1023,1049]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="3" pageNumber="496">Fig. 2C, D</figureCitation>
|
||
). Pedipalps, setation similar to legs; tibia prolateral surface with distinct patch of dense, brownish macrosetae (
|
||
<figureCitation box="[402,496,1733,1760]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="4" pageNumber="497">Fig. 1C</figureCitation>
|
||
). Polygonal macrosetae absent.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333663" ID-Zenodo-Dep="7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="3" pageNumber="496" startId="3.[136,235,1998,2022]" targetBox="[151,1435,1083,1974]" targetPageId="3">
|
||
<paragraph blockId="3.[136,1451,1998,2058]" pageId="3" pageNumber="496">
|
||
<emphasis bold="true" box="[136,260,1998,2022]" pageId="3" pageNumber="496">FIGURE 2.</emphasis>
|
||
<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[267,493,1998,2022]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis box="[267,493,1998,2022]" italics="true" pageId="3" pageNumber="496">Hirsutisoma grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[500,580,1998,2022]" pageId="3" pageNumber="496">
|
||
<taxonomicNameLabel box="[500,580,1998,2022]" pageId="3" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (AMNH_IZC 00357137).
|
||
<emphasis bold="true" box="[983,1001,1998,2022]" pageId="3" pageNumber="496">A</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[1051,1068,1998,2022]" pageId="3" pageNumber="496">B</emphasis>
|
||
, Opisthosoma, dorsal (
|
||
<emphasis bold="true" box="[1305,1323,1998,2022]" pageId="3" pageNumber="496">A</emphasis>
|
||
) and ventral (
|
||
<emphasis bold="true" box="[144,161,2034,2058]" pageId="3" pageNumber="496">B</emphasis>
|
||
) aspects.
|
||
<emphasis bold="true" box="[262,280,2034,2058]" pageId="3" pageNumber="496">C</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[330,348,2034,2058]" pageId="3" pageNumber="496">D</emphasis>
|
||
, Sinistral leg III, prolateral aspect (
|
||
<emphasis bold="true" box="[708,726,2034,2058]" pageId="3" pageNumber="496">C</emphasis>
|
||
) and closeup (
|
||
<emphasis bold="true" box="[876,894,2034,2058]" pageId="3" pageNumber="496">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="4" pageNumber="497" startId="4.[136,213,158,179]">
|
||
<paragraph blockId="4.[136,1452,158,251]" pageId="4" pageNumber="497">
|
||
<emphasis bold="true" box="[136,237,158,179]" pageId="4" pageNumber="497">TABLE 1.</emphasis>
|
||
Measurements (mm) for holotype of
|
||
<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[591,794,158,180]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis box="[591,794,158,180]" italics="true" pageId="4" pageNumber="497">Hirsutisoma grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[802,878,158,179]" pageId="4" pageNumber="497">
|
||
<taxonomicNameLabel box="[802,878,158,179]" pageId="4" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
deposited in the American Museum of Natural History, New York. Abbreviations: L, length; W, width; H, height. Excludes measurements that could not be reliably obtained due to position or preservation of structure.
|
||
<superScript attach="right" box="[227,234,229,241]" fontSize="5" pageId="4" pageNumber="497">1</superScript>
|
||
excludes pygidium or cucullus;
|
||
<superScript attach="right" box="[529,536,229,241]" fontSize="5" pageId="4" pageNumber="497">2</superScript>
|
||
maximum width;
|
||
<superScript attach="right" box="[702,709,229,241]" fontSize="5" pageId="4" pageNumber="497">3</superScript>
|
||
maximum height;
|
||
<superScript attach="right" box="[880,887,229,241]" fontSize="5" pageId="4" pageNumber="497">4</superScript>
|
||
midline.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="4" pageNumber="497">
|
||
<table box="[311,1174,265,1647]" gridcols="3" gridrows="46" pageId="4" pageNumber="497">
|
||
<tr box="[311,1174,265,284]" gridrow="0" pageId="4" pageNumber="497" rowspan-0="1" rowspan-1="1">
|
||
<th box="[1071,1174,265,284]" gridcol="2" gridrow="0" pageId="4" pageNumber="497">Holotype ♂</th>
|
||
</tr>
|
||
<tr box="[311,1174,293,314]" gridrow="1" pageId="4" pageNumber="497" rowspan-1="1">
|
||
<th box="[311,471,293,314]" gridcol="0" gridrow="1" pageId="4" pageNumber="497">Total Body L1</th>
|
||
<td box="[1071,1174,293,314]" gridcol="2" gridrow="1" pageId="4" pageNumber="497">~3.91</td>
|
||
</tr>
|
||
<tr box="[311,1174,323,345]" gridrow="2" pageId="4" pageNumber="497">
|
||
<th box="[311,471,323,345]" gridcol="0" gridrow="2" pageId="4" pageNumber="497">
|
||
<taxonomicName box="[311,389,325,344]" class="Gastropoda" family="Conidae" genus="Cucullus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="497" phylum="Cnidaria" rank="genus">Cucullus</taxonomicName>
|
||
</th>
|
||
<td box="[641,876,323,345]" gridcol="1" gridrow="2" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,323,345]" gridcol="2" gridrow="2" pageId="4" pageNumber="497">0.58</td>
|
||
</tr>
|
||
<tr box="[311,1174,355,375]" gridrow="3" pageId="4" pageNumber="497">
|
||
<th box="[311,471,355,375]" gridcol="0" gridrow="3" pageId="4" pageNumber="497">Carapace</th>
|
||
<td box="[641,876,355,375]" gridcol="1" gridrow="3" pageId="4" pageNumber="497">L</td>
|
||
<td box="[1071,1174,355,375]" gridcol="2" gridrow="3" pageId="4" pageNumber="497">~1.25</td>
|
||
</tr>
|
||
<tr box="[311,1174,384,405]" gridrow="4" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,384,405]" gridcol="1" gridrow="4" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,384,405]" gridcol="2" gridrow="4" pageId="4" pageNumber="497">~0.90</td>
|
||
</tr>
|
||
<tr box="[311,1174,414,435]" gridrow="5" pageId="4" pageNumber="497">
|
||
<th box="[311,471,414,435]" gridcol="0" gridrow="5" pageId="4" pageNumber="497">Opisthosoma</th>
|
||
<td box="[641,876,414,435]" gridcol="1" gridrow="5" pageId="4" pageNumber="497">Total L1</td>
|
||
<td box="[1071,1174,414,435]" gridcol="2" gridrow="5" pageId="4" pageNumber="497">2.67</td>
|
||
</tr>
|
||
<tr box="[311,1174,445,466]" gridrow="6" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,445,466]" gridcol="1" gridrow="6" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,445,466]" gridcol="2" gridrow="6" pageId="4" pageNumber="497">1.94</td>
|
||
</tr>
|
||
<tr box="[311,1174,475,496]" gridrow="7" pageId="4" pageNumber="497">
|
||
<th box="[311,471,475,496]" gridcol="0" gridrow="7" pageId="4" pageNumber="497">Median Plate XII</th>
|
||
<td box="[641,876,475,496]" gridcol="1" gridrow="7" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,475,496]" gridcol="2" gridrow="7" pageId="4" pageNumber="497">1.20</td>
|
||
</tr>
|
||
<tr box="[311,1174,507,526]" gridrow="8" pageId="4" pageNumber="497">
|
||
<th box="[311,471,507,526]" gridcol="0" gridrow="8" pageId="4" pageNumber="497">Median Plate XIII</th>
|
||
<td box="[641,876,507,526]" gridcol="1" gridrow="8" pageId="4" pageNumber="497">L</td>
|
||
<td box="[1071,1174,507,526]" gridcol="2" gridrow="8" pageId="4" pageNumber="497">0.62</td>
|
||
</tr>
|
||
<tr box="[311,1174,535,557]" gridrow="9" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,535,557]" gridcol="1" gridrow="9" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,535,557]" gridcol="2" gridrow="9" pageId="4" pageNumber="497">1.16</td>
|
||
</tr>
|
||
<tr box="[311,1174,567,587]" gridrow="10" pageId="4" pageNumber="497">
|
||
<th box="[311,471,567,587]" gridcol="0" gridrow="10" pageId="4" pageNumber="497">Sternum</th>
|
||
<td box="[641,876,567,587]" gridcol="1" gridrow="10" pageId="4" pageNumber="497">L</td>
|
||
<td box="[1071,1174,567,587]" gridcol="2" gridrow="10" pageId="4" pageNumber="497">0.45</td>
|
||
</tr>
|
||
<tr box="[311,1174,596,617]" gridrow="11" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,596,617]" gridcol="1" gridrow="11" pageId="4" pageNumber="497">W2</td>
|
||
<td box="[1071,1174,596,617]" gridcol="2" gridrow="11" pageId="4" pageNumber="497">0.37</td>
|
||
</tr>
|
||
<tr box="[311,1174,628,647]" gridrow="12" pageId="4" pageNumber="497">
|
||
<th box="[311,471,628,647]" gridcol="0" gridrow="12" pageId="4" pageNumber="497">Pedipalp</th>
|
||
<td box="[641,876,628,647]" gridcol="1" gridrow="12" pageId="4" pageNumber="497">Femur L</td>
|
||
<td box="[1071,1174,628,647]" gridcol="2" gridrow="12" pageId="4" pageNumber="497">~0.26</td>
|
||
</tr>
|
||
<tr box="[311,1174,657,678]" gridrow="13" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,657,678]" gridcol="1" gridrow="13" pageId="4" pageNumber="497">Femur H3</td>
|
||
<td box="[1071,1174,657,678]" gridcol="2" gridrow="13" pageId="4" pageNumber="497">~0.19</td>
|
||
</tr>
|
||
<tr box="[311,1174,689,708]" gridrow="14" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,689,708]" gridcol="1" gridrow="14" pageId="4" pageNumber="497">Tibia L</td>
|
||
<td box="[1071,1174,689,708]" gridcol="2" gridrow="14" pageId="4" pageNumber="497">~0.22</td>
|
||
</tr>
|
||
<tr box="[311,1174,719,738]" gridrow="15" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,719,738]" gridcol="1" gridrow="15" pageId="4" pageNumber="497">Tarsus (Movable Finger) L</td>
|
||
<td box="[1071,1174,719,738]" gridcol="2" gridrow="15" pageId="4" pageNumber="497">0.15</td>
|
||
</tr>
|
||
<tr box="[311,1174,749,769]" gridrow="16" pageId="4" pageNumber="497">
|
||
<th box="[311,471,749,769]" gridcol="0" gridrow="16" pageId="4" pageNumber="497">Leg I</th>
|
||
<td box="[641,876,749,769]" gridcol="1" gridrow="16" pageId="4" pageNumber="497">Coxa L</td>
|
||
<td box="[1071,1174,749,769]" gridcol="2" gridrow="16" pageId="4" pageNumber="497">0.41</td>
|
||
</tr>
|
||
<tr box="[311,1174,779,799]" gridrow="17" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,779,799]" gridcol="1" gridrow="17" pageId="4" pageNumber="497">Trochanter L</td>
|
||
<td box="[1071,1174,779,799]" gridcol="2" gridrow="17" pageId="4" pageNumber="497">0.32</td>
|
||
</tr>
|
||
<tr box="[311,1174,810,829]" gridrow="18" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,810,829]" gridcol="1" gridrow="18" pageId="4" pageNumber="497">Femur L</td>
|
||
<td box="[1071,1174,810,829]" gridcol="2" gridrow="18" pageId="4" pageNumber="497">0.86</td>
|
||
</tr>
|
||
<tr box="[311,1174,838,859]" gridrow="19" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,838,859]" gridcol="1" gridrow="19" pageId="4" pageNumber="497">Femur W4</td>
|
||
<td box="[1071,1174,838,859]" gridcol="2" gridrow="19" pageId="4" pageNumber="497">0.09</td>
|
||
</tr>
|
||
<tr box="[311,1174,870,890]" gridrow="20" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,870,890]" gridcol="1" gridrow="20" pageId="4" pageNumber="497">Patella L</td>
|
||
<td box="[1071,1174,870,890]" gridcol="2" gridrow="20" pageId="4" pageNumber="497">0.37</td>
|
||
</tr>
|
||
<tr box="[311,1174,900,920]" gridrow="21" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,900,920]" gridcol="1" gridrow="21" pageId="4" pageNumber="497">Tibia L</td>
|
||
<td box="[1071,1174,900,920]" gridcol="2" gridrow="21" pageId="4" pageNumber="497">0.54</td>
|
||
</tr>
|
||
<tr box="[311,1174,931,950]" gridrow="22" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,931,950]" gridcol="1" gridrow="22" pageId="4" pageNumber="497">Metatarsus L</td>
|
||
<td box="[1071,1174,931,950]" gridcol="2" gridrow="22" pageId="4" pageNumber="497">0.64</td>
|
||
</tr>
|
||
<tr box="[311,1174,961,981]" gridrow="23" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,961,981]" gridcol="1" gridrow="23" pageId="4" pageNumber="497">Tarsus L</td>
|
||
<td box="[1071,1174,961,981]" gridcol="2" gridrow="23" pageId="4" pageNumber="497">0.25</td>
|
||
</tr>
|
||
<tr box="[311,1174,991,1011]" gridrow="24" pageId="4" pageNumber="497">
|
||
<th box="[311,471,991,1011]" gridcol="0" gridrow="24" pageId="4" pageNumber="497">Leg II</th>
|
||
<td box="[641,876,991,1011]" gridcol="1" gridrow="24" pageId="4" pageNumber="497">Coxa L</td>
|
||
<td box="[1071,1174,991,1011]" gridcol="2" gridrow="24" pageId="4" pageNumber="497">0.43</td>
|
||
</tr>
|
||
<tr box="[311,1174,1022,1041]" gridrow="25" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1022,1041]" gridcol="1" gridrow="25" pageId="4" pageNumber="497">Trochanter L</td>
|
||
<td box="[1071,1174,1022,1041]" gridcol="2" gridrow="25" pageId="4" pageNumber="497">0.37</td>
|
||
</tr>
|
||
<tr box="[311,1174,1052,1071]" gridrow="26" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1052,1071]" gridcol="1" gridrow="26" pageId="4" pageNumber="497">Metatarsus L</td>
|
||
<td box="[1071,1174,1052,1071]" gridcol="2" gridrow="26" pageId="4" pageNumber="497">0.97</td>
|
||
</tr>
|
||
<tr box="[311,1174,1082,1102]" gridrow="27" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1082,1102]" gridcol="1" gridrow="27" pageId="4" pageNumber="497">Tarsus L</td>
|
||
<td box="[1071,1174,1082,1102]" gridcol="2" gridrow="27" pageId="4" pageNumber="497">~1.25</td>
|
||
</tr>
|
||
<tr box="[311,1174,1113,1132]" gridrow="28" pageId="4" pageNumber="497">
|
||
<th box="[311,471,1113,1132]" gridcol="0" gridrow="28" pageId="4" pageNumber="497">Leg III</th>
|
||
<td box="[641,876,1113,1132]" gridcol="1" gridrow="28" pageId="4" pageNumber="497">Coxa L</td>
|
||
<td box="[1071,1174,1113,1132]" gridcol="2" gridrow="28" pageId="4" pageNumber="497">0.43</td>
|
||
</tr>
|
||
<tr box="[311,1174,1143,1162]" gridrow="29" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1143,1162]" gridcol="1" gridrow="29" pageId="4" pageNumber="497">Trochanter L</td>
|
||
<td box="[1071,1174,1143,1162]" gridcol="2" gridrow="29" pageId="4" pageNumber="497">0.32</td>
|
||
</tr>
|
||
<tr box="[311,1174,1173,1192]" gridrow="30" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1173,1192]" gridcol="1" gridrow="30" pageId="4" pageNumber="497">Femur L</td>
|
||
<td box="[1071,1174,1173,1192]" gridcol="2" gridrow="30" pageId="4" pageNumber="497">0.41</td>
|
||
</tr>
|
||
<tr box="[311,1174,1202,1223]" gridrow="31" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1202,1223]" gridcol="1" gridrow="31" pageId="4" pageNumber="497">Femur H4</td>
|
||
<td box="[1071,1174,1202,1223]" gridcol="2" gridrow="31" pageId="4" pageNumber="497">~0.30</td>
|
||
</tr>
|
||
<tr box="[311,1174,1234,1253]" gridrow="32" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1234,1253]" gridcol="1" gridrow="32" pageId="4" pageNumber="497">Patella L</td>
|
||
<td box="[1071,1174,1234,1253]" gridcol="2" gridrow="32" pageId="4" pageNumber="497">0.30</td>
|
||
</tr>
|
||
<tr box="[311,1174,1264,1283]" gridrow="33" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1264,1283]" gridcol="1" gridrow="33" pageId="4" pageNumber="497">Tibia L</td>
|
||
<td box="[1071,1174,1264,1283]" gridcol="2" gridrow="33" pageId="4" pageNumber="497">0.22</td>
|
||
</tr>
|
||
<tr box="[311,1174,1294,1314]" gridrow="34" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1294,1314]" gridcol="1" gridrow="34" pageId="4" pageNumber="497">Metatarsus L</td>
|
||
<td box="[1071,1174,1294,1314]" gridcol="2" gridrow="34" pageId="4" pageNumber="497">~0.43</td>
|
||
</tr>
|
||
<tr box="[311,1174,1324,1344]" gridrow="35" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1324,1344]" gridcol="1" gridrow="35" pageId="4" pageNumber="497">Tarsus L</td>
|
||
<td box="[1071,1174,1324,1344]" gridcol="2" gridrow="35" pageId="4" pageNumber="497">~0.86</td>
|
||
</tr>
|
||
<tr box="[311,1174,1355,1374]" gridrow="36" pageId="4" pageNumber="497">
|
||
<th box="[311,471,1355,1374]" gridcol="0" gridrow="36" pageId="4" pageNumber="497">Leg IV</th>
|
||
<td box="[641,876,1355,1374]" gridcol="1" gridrow="36" pageId="4" pageNumber="497">Coxa L</td>
|
||
<td box="[1071,1174,1355,1374]" gridcol="2" gridrow="36" pageId="4" pageNumber="497">0.47</td>
|
||
</tr>
|
||
<tr box="[311,1174,1385,1404]" gridrow="37" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1385,1404]" gridcol="1" gridrow="37" pageId="4" pageNumber="497">Trochanter 1 L</td>
|
||
<td box="[1071,1174,1385,1404]" gridcol="2" gridrow="37" pageId="4" pageNumber="497">0.43</td>
|
||
</tr>
|
||
<tr box="[311,1174,1415,1435]" gridrow="38" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1415,1435]" gridcol="1" gridrow="38" pageId="4" pageNumber="497">Trochanter 2 L</td>
|
||
<td box="[1071,1174,1415,1435]" gridcol="2" gridrow="38" pageId="4" pageNumber="497">0.41</td>
|
||
</tr>
|
||
<tr box="[311,1174,1446,1465]" gridrow="39" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1446,1465]" gridcol="1" gridrow="39" pageId="4" pageNumber="497">Femur L</td>
|
||
<td box="[1071,1174,1446,1465]" gridcol="2" gridrow="39" pageId="4" pageNumber="497">0.84</td>
|
||
</tr>
|
||
<tr box="[311,1174,1474,1495]" gridrow="40" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1474,1495]" gridcol="1" gridrow="40" pageId="4" pageNumber="497">Femur W4</td>
|
||
<td box="[1071,1174,1474,1495]" gridcol="2" gridrow="40" pageId="4" pageNumber="497">~0.15</td>
|
||
</tr>
|
||
<tr box="[311,1174,1504,1526]" gridrow="41" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1504,1526]" gridcol="1" gridrow="41" pageId="4" pageNumber="497">Femur H4</td>
|
||
<td box="[1071,1174,1504,1526]" gridcol="2" gridrow="41" pageId="4" pageNumber="497">0.15</td>
|
||
</tr>
|
||
<tr box="[311,1174,1536,1556]" gridrow="42" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1536,1556]" gridcol="1" gridrow="42" pageId="4" pageNumber="497">Patella L</td>
|
||
<td box="[1071,1174,1536,1556]" gridcol="2" gridrow="42" pageId="4" pageNumber="497">0.69</td>
|
||
</tr>
|
||
<tr box="[311,1174,1567,1586]" gridrow="43" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1567,1586]" gridcol="1" gridrow="43" pageId="4" pageNumber="497">Tibia L</td>
|
||
<td box="[1071,1174,1567,1586]" gridcol="2" gridrow="43" pageId="4" pageNumber="497">0.88</td>
|
||
</tr>
|
||
<tr box="[311,1174,1597,1616]" gridrow="44" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1597,1616]" gridcol="1" gridrow="44" pageId="4" pageNumber="497">Metatarsus L</td>
|
||
<td box="[1071,1174,1597,1616]" gridcol="2" gridrow="44" pageId="4" pageNumber="497">0.69</td>
|
||
</tr>
|
||
<tr box="[311,1174,1627,1647]" gridrow="45" pageId="4" pageNumber="497" rowspan-0="1">
|
||
<td box="[641,876,1627,1647]" gridcol="1" gridrow="45" pageId="4" pageNumber="497">Tarsus L</td>
|
||
<td box="[1071,1174,1627,1647]" gridcol="2" gridrow="45" pageId="4" pageNumber="497">~0.73</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="4.[136,778,1696,2093]" pageId="4" pageNumber="497">
|
||
<emphasis box="[189,625,1807,1834]" italics="true" pageId="4" pageNumber="497">Tegument surface macrosculpture</emphasis>
|
||
. Tegument completely smooth, soma and appendages without single visible granule; cuticular pits absent.
|
||
</paragraph>
|
||
<paragraph blockId="4.[136,778,1696,2093]" lastBlockId="4.[810,1452,1696,2093]" pageId="4" pageNumber="497">
|
||
<emphasis box="[189,476,1918,1944]" italics="true" pageId="4" pageNumber="497">Interlocking mechanism</emphasis>
|
||
. Absent. Prosoma and opisthosoma not interlocked (
|
||
<figureCitation box="[490,587,1955,1982]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="4" pageNumber="497">Fig. 3A</figureCitation>
|
||
); coxae of legs IV unmodified, not hooked to opisthosoma ventrally; opisthosoma dorsal and ventral surfaces visibly oval (
|
||
<figureCitation box="[144,270,2066,2092]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="4" pageNumber="497">Fig. 2A, B</figureCitation>
|
||
), anterior margins not truncate; anterior part of opisthosoma extending dorsally over posterior part of prosoma, slightly obscuring the latter; exposed pedicel presumably connecting both tagmata (
|
||
<figureCitation box="[1238,1328,1770,1797]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="4" pageNumber="497">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[810,1452,1696,2093]" pageId="4" pageNumber="497">
|
||
<emphasis box="[863,973,1808,1834]" italics="true" pageId="4" pageNumber="497">Carapace</emphasis>
|
||
. Carapace pear-shaped, narrower anteriorly (reminiscent of typical spider carapace), approximately as long as or slightly longer than wide; three pairs of ocelli in two triads laterally (
|
||
<figureCitation box="[1064,1151,1918,1945]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="4" pageNumber="497">Fig. 3B</figureCitation>
|
||
); sulci, if any, not visible.
|
||
</paragraph>
|
||
<paragraph blockId="4.[810,1452,1696,2093]" lastBlockId="5.[136,778,1823,2093]" lastPageId="5" lastPageNumber="498" pageId="4" pageNumber="497">
|
||
<taxonomicName box="[863,963,1955,1981]" class="Gastropoda" family="Conidae" genus="Cucullus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="497" phylum="Cnidaria" rank="genus">
|
||
<emphasis box="[863,963,1955,1981]" italics="true" pageId="4" pageNumber="497">Cucullus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName box="[981,1081,1955,1981]" class="Gastropoda" family="Conidae" genus="Cucullus" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="497" phylum="Cnidaria" rank="genus">Cucullus</taxonomicName>
|
||
trapezoidal, slightly wider than long (
|
||
<figureCitation box="[881,978,1992,2018]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="4" pageNumber="497">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation box="[997,1032,1992,2018]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="4" pageNumber="497">3B</figureCitation>
|
||
); lateral margins slightly diverging ventrally; ventrolateral margins rounded; ventral margin sublinear in ventral aspect, slightly bilobate in anterior aspect; anterior surface slightly concave, almost flat (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="5" pageNumber="498">Fig. 1C</figureCitation>
|
||
), without remarkable structures or modifications; sulci, if any, not visible.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333665" ID-Zenodo-Dep="7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="5" pageNumber="498" startId="5.[136,235,1522,1546]" targetBox="[180,1417,200,1486]" targetPageId="5">
|
||
<paragraph blockId="5.[136,1452,1522,1762]" pageId="5" pageNumber="498">
|
||
<emphasis bold="true" box="[136,260,1522,1546]" pageId="5" pageNumber="498">FIGURE 3.</emphasis>
|
||
<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[266,492,1522,1546]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis box="[266,492,1522,1546]" italics="true" pageId="5" pageNumber="498">Hirsutisoma grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[498,579,1522,1546]" pageId="5" pageNumber="498">
|
||
<taxonomicNameLabel box="[498,579,1522,1546]" pageId="5" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (AMNH_IZC 00357137).
|
||
<emphasis bold="true" box="[980,998,1522,1546]" pageId="5" pageNumber="498">A</emphasis>
|
||
, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel.
|
||
<emphasis bold="true" box="[778,795,1558,1582]" pageId="5" pageNumber="498">B</emphasis>
|
||
, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II.
|
||
<emphasis bold="true" box="[209,227,1594,1618]" pageId="5" pageNumber="498">C</emphasis>
|
||
, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII).
|
||
<emphasis bold="true" box="[1277,1295,1594,1618]" pageId="5" pageNumber="498">D</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[1345,1362,1594,1618]" pageId="5" pageNumber="498">E</emphasis>
|
||
, Dextral pedipalp, retroventral aspect (
|
||
<emphasis bold="true" box="[438,456,1630,1654]" pageId="5" pageNumber="498">D</emphasis>
|
||
), and closeup of tarsus (movable finger) and lamellate projections (
|
||
<emphasis bold="true" box="[1141,1158,1630,1654]" pageId="5" pageNumber="498">E</emphasis>
|
||
).
|
||
<emphasis bold="true" box="[1178,1193,1630,1654]" pageId="5" pageNumber="498">F</emphasis>
|
||
, Sinistral pedipalp, distal segments, dorsal aspect.
|
||
<emphasis bold="true" box="[390,410,1666,1690]" pageId="5" pageNumber="498">G</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[461,481,1666,1690]" pageId="5" pageNumber="498">H</emphasis>
|
||
, Dextral leg I femur, dorsal (
|
||
<emphasis bold="true" box="[782,802,1666,1690]" pageId="5" pageNumber="498">G</emphasis>
|
||
) and prodorsal (
|
||
<emphasis bold="true" box="[973,993,1666,1690]" pageId="5" pageNumber="498">H</emphasis>
|
||
) aspects.
|
||
<emphasis bold="true" box="[1095,1105,1666,1690]" pageId="5" pageNumber="498">I</emphasis>
|
||
, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[136,778,1823,2093]" lastBlockId="5.[810,1452,1823,2093]" pageId="5" pageNumber="498">
|
||
<emphasis box="[189,404,1944,1970]" italics="true" pageId="5" pageNumber="498">Coxosternal region</emphasis>
|
||
. Sternum large, situated centrally on prosoma ventral surface (
|
||
<figureCitation box="[464,559,1985,2011]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="5" pageNumber="498">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation box="[573,610,1985,2011]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="5" pageNumber="498">4C</figureCitation>
|
||
); drop-shaped, acute posteriorly; coxae of pedipalps and legs arranged concentrically around sternum; coxae of pedipalps abutting one another, forming floor of preoral cavity; coxae of legs I–IV short, separate from coxae of adjacent legs (
|
||
<emphasis box="[874,882,1905,1930]" italics="true" pageId="5" pageNumber="498">i</emphasis>
|
||
.
|
||
<emphasis box="[889,901,1905,1930]" italics="true" pageId="5" pageNumber="498">e</emphasis>
|
||
., lateral margins not fused along contact zone); sternum and coxae without remarkable structures or modifications.
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1452,1823,2093]" lastBlockId="6.[136,778,1938,2092]" lastPageId="6" lastPageNumber="499" pageId="5" pageNumber="498">
|
||
<emphasis box="[863,986,2026,2052]" italics="true" pageId="5" pageNumber="498">Chelicerae</emphasis>
|
||
. Chelicerae retracted into preoral cavity, not visible under light microscopy (micro-CT scanning revealed that fixed and movable fingers are long and similar in length, reminiscent of cheliceral fingers of Solifugae); finger dentition, if any, obscured; manus seemingly unmodified.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333667" ID-Zenodo-Dep="7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="6" pageNumber="499" startId="6.[136,235,1709,1733]" targetBox="[223,1401,205,1672]" targetPageId="6">
|
||
<paragraph blockId="6.[136,1452,1709,1877]" pageId="6" pageNumber="499">
|
||
<emphasis bold="true" box="[136,259,1709,1733]" pageId="6" pageNumber="499">FIGURE 4.</emphasis>
|
||
Legs and coxosternal region.
|
||
<emphasis bold="true" box="[559,577,1709,1733]" pageId="6" pageNumber="499">A</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[624,641,1709,1733]" pageId="6" pageNumber="499">B</emphasis>
|
||
, Legs, dorsal aspect, of modern ricinuleid (
|
||
<taxonomicName authority="Ewing, 1929" authorityName="Ewing" authorityYear="1929" box="[1074,1341,1709,1733]" class="Arachnida" family="Ricinoididae" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="499" phylum="Arthropoda" rank="family">Ricinoididae Ewing, 1929</taxonomicName>
|
||
, schematic representation) (
|
||
<emphasis bold="true" box="[307,325,1745,1769]" pageId="6" pageNumber="499">A</emphasis>
|
||
) and
|
||
<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[392,622,1745,1769]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis box="[392,622,1745,1769]" italics="true" pageId="6" pageNumber="499">Hirsutisoma grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[633,717,1745,1769]" pageId="6" pageNumber="499">
|
||
<taxonomicNameLabel box="[633,717,1745,1769]" pageId="6" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (AMNH_IZC 00357137), dextral leg IV (
|
||
<emphasis bold="true" box="[1308,1325,1745,1769]" pageId="6" pageNumber="499">B</emphasis>
|
||
).
|
||
<emphasis bold="true" box="[1351,1369,1745,1769]" pageId="6" pageNumber="499">C</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[1427,1445,1745,1769]" pageId="6" pageNumber="499">D</emphasis>
|
||
,
|
||
<taxonomicName authority="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI, 2022" authorityName="BOTERO-TRUJILLO & DAVIS & MICHALIK & PRENDINI" authorityYear="2022" box="[136,361,1781,1805]" class="Arachnida" family="Hirsutisomidae" genus="Hirsutisoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ricinulei" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
|
||
<emphasis box="[136,361,1781,1805]" italics="true" pageId="6" pageNumber="499">Hirsutisoma grimaldii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[367,446,1781,1805]" pageId="6" pageNumber="499">
|
||
<taxonomicNameLabel box="[367,446,1781,1805]" pageId="6" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ♂ (AMNH_IZC 00357137), coxosternal region (
|
||
<emphasis bold="true" box="[1044,1062,1781,1805]" pageId="6" pageNumber="499">C</emphasis>
|
||
) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (
|
||
<emphasis bold="true" box="[475,493,1817,1841]" pageId="6" pageNumber="499">D</emphasis>
|
||
) (note modified first tarsomere and unmodified second tarsomere). Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[810,1452,1938,2093]" lastBlockId="7.[136,778,158,2093]" lastPageId="7" lastPageNumber="500" pageId="6" pageNumber="499">
|
||
<emphasis box="[863,975,1938,1964]" italics="true" pageId="6" pageNumber="499">Pedipalps</emphasis>
|
||
. Pedipalps positioned anteriad (
|
||
<figureCitation box="[1351,1446,1938,1964]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="6" pageNumber="499">Figs 1C</figureCitation>
|
||
,
|
||
<figureCitation box="[810,845,1981,2007]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="6" pageNumber="499">3B</figureCitation>
|
||
); consisting of coxa, trochanters 1 and 2, femur, tibia, and tarsus; femur slightly longer than deep or wide; tibia length at most twice its maximum diameter, without fixed finger (not chelate); tarsus (movable finger) long and curved (
|
||
<figureCitation box="[232,355,194,220]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">Fig. 3D–F</figureCitation>
|
||
), about as long as tibia and at least half length of cucullo-carapacial commissure, laterally compressed, with sharp dorsal and ventral edges, basally with comb of 8–10 lamellate projections on proventral surface. Pedipalps without other remarkable structures or modifications.
|
||
</paragraph>
|
||
<paragraph blockId="7.[136,778,158,2093]" pageId="7" pageNumber="500">
|
||
<emphasis box="[189,242,411,436]" italics="true" pageId="7" pageNumber="500">Legs</emphasis>
|
||
. Legs comprising seven segments (coxa, one trochanter, femur, patella, tibia, metatarsus, and tarsus), except leg IV with eight (
|
||
<emphasis box="[432,440,483,508]" italics="true" pageId="7" pageNumber="500">i</emphasis>
|
||
.
|
||
<emphasis box="[447,459,483,508]" italics="true" pageId="7" pageNumber="500">e</emphasis>
|
||
., two trochanters). Plane of flexion of tibia similar to (aligned with) plane of flexion of patella (
|
||
<figureCitation box="[269,364,554,580]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="7" pageNumber="500">Fig. 4B</figureCitation>
|
||
), axis of flexion at articulation of tibia and patella slightly rotated or not rotated. Leg femora unmodified. Leg I femur very narrow proximally, longitudinal axis crooked (
|
||
<figureCitation box="[468,616,662,689]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="7" pageNumber="500">Figs 1C, D</figureCitation>
|
||
,
|
||
<figureCitation box="[638,764,662,688]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">3B, G, H</figureCitation>
|
||
); tarsus without dorsomedian lobe. Leg II longest; tarsus comprising five tarsomeres, all longer than wide and similar in length (
|
||
<figureCitation box="[346,439,770,796]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">Fig. 2C</figureCitation>
|
||
); laterodistal margins of first to fourth tarsomeres not overlapping adjacent tarsomere; fifth tarsomere without dorsomedian lobe. Leg III femur, patella, tibia, and metatarsus short and subspherical (
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">Figs 2C, D</figureCitation>
|
||
,
|
||
<figureCitation box="[231,270,914,940]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="7" pageNumber="500">4D</figureCitation>
|
||
); metatarsus prolateral surface with sharp, flat, glabrous expansion distally (
|
||
<figureCitation box="[538,636,950,976]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="7" pageNumber="500">Fig. 4D</figureCitation>
|
||
); metatarsal process forming curved, concave flap situated apically on metatarsus, near to and opposing tarsus; tarsus comprising four tarsomeres; first tarsomere markedly elongate, about four times longer than other tarsomeres, sinuous in dorsal aspect, shaped to accommodate copulatory apparatus that originates from its base, with pronounced retrodorsal longitudinal ridge; tarsomeres 2–4 short (each about as long as deep), similar in length and unmodified. Leg IV without remarkable structures or modifications; tarsus comprising five tarsomeres (
|
||
<figureCitation box="[473,557,1310,1336]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">Fig. 3I</figureCitation>
|
||
). Legs III and IV, terminal tarsomere apex margins not projected, ungues completely exposed in dorsal and lateral aspects (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">Figs 3I</figureCitation>
|
||
,
|
||
<figureCitation box="[173,212,1418,1444]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="7" pageNumber="500">4D</figureCitation>
|
||
); dorsomedian lobe absent. Ungues acuminate and sharp on all legs, longer on III and IV.
|
||
</paragraph>
|
||
<paragraph blockId="7.[136,778,158,2093]" pageId="7" pageNumber="500">
|
||
<emphasis box="[189,771,1490,1516]" italics="true" pageId="7" pageNumber="500">Copulatory apparatus and associated structures</emphasis>
|
||
. Comprising two parts not closely associated with one another (other components may be not visible) (
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">Figs 2C, D</figureCitation>
|
||
,
|
||
<figureCitation box="[170,209,1598,1624]" captionStart="FIGURE 4" captionStartId="6.[136,235,1709,1733]" captionTargetBox="[223,1401,205,1672]" captionTargetId="figure-85@6.[151,1436,183,1685]" captionTargetPageId="6" captionText="FIGURE 4. Legs and coxosternal region. A and B, Legs, dorsal aspect, of modern ricinuleid (Ricinoididae Ewing, 1929, schematic representation) (A) and Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), dextral leg IV (B). C and D, Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137), coxosternal region (C) (legs identified by Roman numerals); sinistral leg III, prolateral aspect (D) (note modified first tarsomere and unmodified second tarsomere).Abbreviations: cx, coxa; fe, femur; me, metatarsus; pa, patella; st, sternum; ti, tibia; tr, trochanter; tr1, trochanter 1; first and second, proximal tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.7333667" httpUri="https://zenodo.org/record/7333667/files/figure.png" pageId="7" pageNumber="500">4D</figureCitation>
|
||
); one part short, horn-shaped, lying prodorsally on first tarsomere; adjacent, in retrolateral position, hyaline, laterally compressed, lanceolate structure, apparently originating on metatarsus; other part, about twice length, narrower, and whip-shaped, curling around distal part of metatarsus. Based on morphology of modified leg III, horn-shaped part of copulatory apparatus presumably protected by various structures: ventrally by modified first tarsomere, dorsally by metatarsal process, prolaterally by metatarsal distal expansion, and retrolaterally by metatarsal(?) lanceolate structure.
|
||
</paragraph>
|
||
<paragraph blockId="7.[136,778,158,2093]" lastBlockId="7.[810,1452,158,620]" pageId="7" pageNumber="500">
|
||
<emphasis box="[189,337,1994,2020]" italics="true" pageId="7" pageNumber="500">Opisthosoma</emphasis>
|
||
. Opisthosoma oval, longer than wide (
|
||
<tableCitation box="[210,299,2030,2056]" captionStart="TABLE 1" captionStartId="4.[136,213,158,179]" captionText="TABLE 1. Measurements (mm) for holotype of Hirsutisoma grimaldii sp. nov. deposited in the American Museum of Natural History, New York.Abbreviations:L, length; W, width; H, height. Excludes measurements that could not be reliably obtained due to position or preservation of structure. 1 excludes pygidium or cucullus; 2 maximum width; 3 maximum height; 4 midline." pageId="7" pageNumber="500">Table 1</tableCitation>
|
||
), broadest at tergite XII. Dorsal surface divided transversally into separate tergites (
|
||
<figureCitation box="[629,724,2066,2092]" captionStart="FIGURE 1" captionStartId="2.[136,235,2008,2032]" captionTargetBox="[151,1435,1091,1982]" captionTargetId="figure-296@2.[151,1435,1091,1983]" captionTargetPageId="2" captionText="FIGURE 1. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Habitus, dorsal (A) and ventral (B) aspects. C, Prosoma, anterodorsal aspect. D, Coxosternal region." figureDoi="http://doi.org/10.5281/zenodo.7333661" httpUri="https://zenodo.org/record/7333661/files/figure.png" pageId="7" pageNumber="500">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation box="[736,773,2066,2092]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">2A</figureCitation>
|
||
,
|
||
<figureCitation box="[810,846,158,184]" captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">3C</figureCitation>
|
||
); tergites without submedian depressions. Tergite X not visible, obscured by long macrosetae and matrix imperfections, segmentation unclear; XI–XIII each comprising median and lateral sclerites (visible for XII and XIII, revealed by micro-CT scanning for XI) (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="5.[136,235,1522,1546]" captionTargetBox="[180,1417,200,1486]" captionTargetId="figure-119@5.[151,1436,189,1498]" captionTargetPageId="5" captionText="FIGURE 3. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A, Schematic representation of soma, showing how both tagmata presumably connected by exposed pedicel. B, Prosoma, dorsal aspect, showing raptorial pedipalp, and legs I and II. C, Opisthosoma, posterior part, dorsal aspect (note median and lateral sclerites of tergites XII and XIII). D and E, Dextral pedipalp, retroventral aspect (D), and closeup of tarsus (movable finger) and lamellate projections (E). F, Sinistral pedipalp, distal segments, dorsal aspect. G and H, Dextral leg I femur, dorsal (G) and prodorsal (H) aspects. I, Sinistral leg IV tarsus, prolateral aspect. Abbreviations: ca, carapace; cu, cucullus; fe, femur; me, metatarsus; pa, patella; ta, tarsus (movable finger); ti, tibia; tr, trochanter; tr2, trochanter 2." figureDoi="http://doi.org/10.5281/zenodo.7333665" httpUri="https://zenodo.org/record/7333665/files/figure.png" pageId="7" pageNumber="500">Fig. 3C</figureCitation>
|
||
); median sclerite of tergite XII only partially visible; of XIII cup-shaped, markedly wider than long (
|
||
<tableCitation box="[1349,1434,376,402]" captionStart="TABLE 1" captionStartId="4.[136,213,158,179]" captionText="TABLE 1. Measurements (mm) for holotype of Hirsutisoma grimaldii sp. nov. deposited in the American Museum of Natural History, New York.Abbreviations:L, length; W, width; H, height. Excludes measurements that could not be reliably obtained due to position or preservation of structure. 1 excludes pygidium or cucullus; 2 maximum width; 3 maximum height; 4 midline." pageId="7" pageNumber="500">Table 1</tableCitation>
|
||
), lateral margins parallel. Sternites XI–XIII entirely fused, forming uniform ventral sternal plate without irregularities (
|
||
<figureCitation box="[818,910,485,511]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">Fig. 2B</figureCitation>
|
||
); submedian depressions absent. Pygidium not exposed (
|
||
<figureCitation box="[914,1031,522,548]" captionStart="FIGURE 2" captionStartId="3.[136,235,1998,2022]" captionTargetBox="[151,1435,1083,1974]" captionTargetId="figure-398@3.[151,1435,1083,1975]" captionTargetPageId="3" captionText="FIGURE 2. Hirsutisoma grimaldii sp. nov., holotype ♂ (AMNH_IZC 00357137). A and B, Opisthosoma, dorsal (A) and ventral (B) aspects. C and D, Sinistral leg III, prolateral aspect (C) and closeup (D)." figureDoi="http://doi.org/10.5281/zenodo.7333663" httpUri="https://zenodo.org/record/7333663/files/figure.png" pageId="7" pageNumber="500">Fig. 2A, B</figureCitation>
|
||
), completely sheltered between dorsal and ventral opisthosomal sclerites, presumably retractile.
|
||
</paragraph>
|
||
<paragraph blockId="7.[810,1452,158,620]" box="[863,1078,594,620]" pageId="7" pageNumber="500">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |