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<document id="D6C50E83E6DCAF1001F88E037809BEEE" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE39BFF0EFF75F88CBAAEFB41" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Protozantaena duplicata Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="160" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="157" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:title id="40397C9D7EFC4C852D8870F8DF1D2203">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="DC37C771A93618B1FF839EFEBD6A9993">Perkins, Philip D.</mods:namePart>
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<mods:title id="34F42EFA588783AC766672F1AB832447">Zootaxa</mods:title>
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<mods:part id="0351EEF5701613790600DF0E5661D0E1">
<mods:date id="BD2C69CB9358B1647BFF113BDC9B473C">2017</mods:date>
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<mods:number id="40DB7C1B193552F467762894555AE507">2017-11-03</mods:number>
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<mods:number id="6610B725DAF1BBC5D60B3F821D5619D0">4342</mods:number>
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<mods:classification id="26C3EE577517B61B9F9A6CE8E9BE9A6A">journal article</mods:classification>
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<mods:identifier id="61B4F20C994C3C5CC762718461DAE1AD" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<treatment id="038587BBE39BFF0EFF75F88CBAAEFB41" ID-DOI="http://doi.org/10.5281/zenodo.5323688" ID-GBIF-Taxon="137233407" ID-Zenodo-Dep="5323688" LSID="urn:lsid:plazi:treatment:038587BBE39BFF0EFF75F88CBAAEFB41" httpUri="http://treatment.plazi.org/id/038587BBE39BFF0EFF75F88CBAAEFB41" lastPageId="159" lastPageNumber="160" pageId="156" pageNumber="157">
<subSubSection id="C3366526E39BFF0DFF75F88CBB98F8E1" pageId="156" pageNumber="157" type="nomenclature">
<paragraph id="8B9336ADE39BFF0DFF75F88CBB60F8C6" blockId="156.[151,676,1833,1893]" box="[151,604,1833,1859]" pageId="156" pageNumber="157">
<heading id="D0DB81C1E39BFF0DFF75F88CBB60F8C6" bold="true" box="[151,604,1833,1859]" fontSize="11" level="1" pageId="156" pageNumber="157" reason="1">
<emphasis id="B958EABFE39BFF0DFF75F88CBB60F8C6" bold="true" box="[151,604,1833,1859]" pageId="156" pageNumber="157">
<taxonomicName id="4C2C4D2EE39BFF0DFF75F88CB881F8C6" ID-CoL="4N7LZ" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,445,1833,1859]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="duplicata" status="sp. nov.">
<emphasis id="B958EABFE39BFF0DFF75F88CB881F8C6" bold="true" box="[151,445,1833,1859]" italics="true" pageId="156" pageNumber="157">Protozantaena duplicata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E39BFF0DFE2EF88CBB60F8C6" box="[460,604,1833,1859]" pageId="156" pageNumber="157" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE39BFF0DFF75F8E9BB98F8E1" blockId="156.[151,676,1833,1893]" box="[151,676,1868,1893]" pageId="156" pageNumber="157">
<figureCitation id="13172A28E39BFF0DFF75F8E9B9C3F8E1" box="[151,255,1868,1893]" captionStart="FIGURE 141" captionStartId="157.[151,250,1733,1755]" captionText="FIGURE 141. Protozantaena duplicata, holotype, habitus dorsal, lateral, and ventral." pageId="156" pageNumber="157">Figs. 141</figureCitation>
(habitus), 143 (aedeagus), 303 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E39BFF0CFF75F831BD8DFF51" lastPageId="157" lastPageNumber="158" pageId="156" pageNumber="157" type="materials_examined">
<paragraph id="8B9336ADE39BFF0CFF75F831BD8DFF51" blockId="156.[151,1437,1940,2037]" lastBlockId="157.[151,1436,151,429]" lastPageId="157" lastPageNumber="158" pageId="156" pageNumber="157">
<materialsCitation id="3B443CF0E39BFF0DFF75F831B8F3F871" ID-GBIF-Occurrence="1697157717" collectingDate="2012-01-24" collectionCode="NHRS" collectorName="T. Ranarilalatiana &amp; J. H. Randriamihaja" elevation="1904" latitude="-19.35364" location="Andongolongo" longLatPrecision="1" longitude="0.7883431" municipality="Vakinankaratra" pageId="156" pageNumber="157" specimenCount="2" stateProvince="Antananarivo" typeStatus="holotype">
<emphasis id="B958EABFE39BFF0DFF75F831B880F828" bold="true" box="[151,444,1940,1965]" pageId="156" pageNumber="157">
<typeStatus id="5497880FE39BFF0DFF75F831B9EEF828" box="[151,210,1940,1965]" pageId="156" pageNumber="157">Type</typeStatus>
Material.
<typeStatus id="5497880FE39BFF0DFEADF831B880F828" box="[335,444,1940,1965]" pageId="156" pageNumber="157" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE39BFF0DFDFEF831BBFEF828" box="[540,706,1940,1965]" country="Madagascar" name="Antananarivo" pageId="156" pageNumber="157">
<emphasis id="B958EABFE39BFF0DFDFEF831BBFEF828" bold="true" box="[540,706,1940,1965]" pageId="156" pageNumber="157">Antananarivo</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E39BFF0DFD32F831BA42F828" box="[720,894,1940,1965]" pageId="156" pageNumber="157">Vakinankaratra</collectingMunicipality>
: Manjakatompo Stn. Forestière:
<location id="8EF36076E39BFF0DFB0EF831BCA5F828" LSID="urn:lsid:plazi:treatment:038587BBE39BFF0EFF75F88CBAAEFB41:8EF36076E39BFF0DFB0EF831BCA5F828" box="[1260,1433,1940,1965]" latitude="-19.35364" longLatPrecision="1" longitude="0.7883431" municipality="Vakinankaratra" name="Andongolongo" pageId="156" pageNumber="157" stateProvince="Antananarivo">Andongolongo</location>
:
<geoCoordinate id="EE18506AE39BFF0DFF75F81DB81AF854" box="[151,294,1976,2001]" direction="south" orientation="latitude" pageId="156" pageNumber="157" precision="1" value="-19.35364">S 19.353639</geoCoordinate>
<geoCoordinate id="EE18506AE39BFF0DFECDF81CB8ECF854" box="[303,464,1976,2001]" direction="east" orientation="longitude" pageId="156" pageNumber="157" precision="1" value="0.7883431">E 0 47.300583</geoCoordinate>
, pool near source, elev.
<elevation id="0001D19EE39BFF0DFD04F81DBA03F855" box="[742,831,1976,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.904" pageId="156" pageNumber="157" unit="m" value="1904.0">
<quantity id="4CD49B48E39BFF0DFD04F81DBA03F855" box="[742,831,1976,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.904" pageId="156" pageNumber="157" unit="m" value="1904.0">1904 m</quantity>
</elevation>
,
<collectingDate id="EFD6E985E39BFF0DFCAEF81DBA80F855" box="[844,956,1976,2001]" pageId="156" pageNumber="157" value="2012-01-24">
<date id="FF92106DE39BFF0DFCAEF81DBA80F855" box="[844,956,1976,2001]" pageId="156" pageNumber="157" value="2012-01-24">24 i 2012</date>
</collectingDate>
,
<collectorName id="26D9537BE39BFF0DFC2BF81CBDAEF854" box="[969,1170,1976,2001]" pageId="156" pageNumber="157">T. Ranarilalatiana</collectorName>
&amp;
<collectorName id="26D9537BE39BFF0DFB5BF81CBCA0F854" box="[1209,1436,1976,2001]" pageId="156" pageNumber="157">J. H. Randriamihaja</collectorName>
(MADMJK12-12) (
<collectionCode id="ED3DAE68E39BFF0DFE94F879B8FBF871" box="[374,455,2012,2036]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="156" pageNumber="157">NHRS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B443CF0E39BFF0CFE3EF879BC1EFF2A" ID-GBIF-Occurrence="1697157758" collectionCode="NHRS" county="Same" elevation="2073" lastPageId="157" lastPageNumber="158" latitude="-19.344889" location="Anosiarivo" longLatPrecision="1" longitude="0.7884023" municipality="Vakinankaratra" pageId="156" pageNumber="157" specimenCount="2" stateProvince="Antananarivo" typeStatus="paratype">
<typeStatus id="5497880FE39BFF0DFE3EF879BB68F870" box="[476,596,2012,2037]" pageId="156" pageNumber="157" type="paratype">
<emphasis id="B958EABFE39BFF0DFE3EF879BB68F870" bold="true" box="[476,596,2012,2037]" pageId="156" pageNumber="157">Paratypes</emphasis>
</typeStatus>
(52):
<collectingCounty id="62F24E21E39BFF0DFD7CF879BBE0F871" box="[670,732,2012,2036]" pageId="156" pageNumber="157">Same</collectingCounty>
data as
<typeStatus id="5497880FE39BFF0DFCA2F879BA9EF870" box="[832,930,2012,2037]" pageId="156" pageNumber="157" type="holotype">holotype</typeStatus>
(6
<collectionCode id="ED3DAE68E39BFF0DFC2FF879BD21F871" box="[973,1053,2012,2036]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="156" pageNumber="157">NHRS</collectionCode>
);
<collectingRegion id="49E8F84FE39BFF0DFBD1F879BDE5F870" box="[1075,1241,2012,2037]" country="Madagascar" name="Antananarivo" pageId="156" pageNumber="157">
<emphasis id="B958EABFE39BFF0DFBD1F879BDE5F870" bold="true" box="[1075,1241,2012,2037]" pageId="156" pageNumber="157">Antananarivo</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BF7ACD7E39BFF0DFB09F879BCA5F870" box="[1259,1433,2012,2037]" pageId="156" pageNumber="157">Vakinankaratra</collectingMunicipality>
: Manjakatompo Stn. Forestière:
<location id="8EF36076E39AFF0CFE19FF32BB40FF35" LSID="urn:lsid:plazi:treatment:038587BBE39BFF0EFF75F88CBAAEFB41:8EF36076E39AFF0CFE19FF32BB40FF35" box="[507,636,151,176]" county="Same" latitude="-19.344889" longLatPrecision="1" longitude="0.7884023" municipality="Vakinankaratra" name="Anosiarivo" pageId="157" pageNumber="158" stateProvince="Antananarivo">Anosiarivo</location>
:
<geoCoordinate id="EE18506AE39AFF0CFD65FF3DBA29FF35" box="[647,789,152,176]" direction="south" orientation="latitude" pageId="157" pageNumber="158" precision="1" value="-19.344889">S 19.344889</geoCoordinate>
<geoCoordinate id="EE18506AE39AFF0CFCFFFF3DBA82FF35" box="[797,958,152,176]" direction="east" orientation="longitude" pageId="157" pageNumber="158" precision="1" value="0.7884023">E 0 47.304139</geoCoordinate>
, lake near source, elev.
<elevation id="0001D19EE39AFF0CFB28FF3DBC1EFF2A" box="[1226,1314,152,176]" metricMagnitude="3" metricUnit="m" metricValue="2.073" pageId="157" pageNumber="158" unit="m" value="2073.0">
<quantity id="4CD49B48E39AFF0CFB28FF3DBC1EFF2A" box="[1226,1314,152,176]" metricMagnitude="3" metricUnit="m" metricValue="2.073" pageId="157" pageNumber="158" unit="m" value="2073.0">2073 m</quantity>
</elevation>
</materialsCitation>
, 19° 21' S, 47° 18' E,
<date id="FF92106DE39AFF0CFEEEFF19B845FF51" box="[268,377,188,213]" pageId="157" pageNumber="158" value="2012-01-24">24 i 2012</date>
, T. Ranarilalatiana &amp; J. H. Randriamihaja (MADMJK12-13) (46 NHRS).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E39AFF0CFF25FF7ABB61FE28" pageId="157" pageNumber="158" type="diagnosis">
<paragraph id="8B9336ADE39AFF0CFF25FF7ABB61FE28" blockId="157.[151,1436,151,429]" pageId="157" pageNumber="158">
<emphasis id="B958EABFE39AFF0CFF25FF7AB8EBFF7D" bold="true" box="[199,471,223,248]" pageId="157" pageNumber="158">Differential Diagnosis.</emphasis>
Sharing with
<taxonomicName id="4C2C4D2EE39AFF0CFD60FF44BA2EFF7D" box="[642,786,223,248]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="ankaratra">
<emphasis id="B958EABFE39AFF0CFD60FF44BA2EFF7D" box="[642,786,223,248]" italics="true" pageId="157" pageNumber="158">P. ankaratra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2C4D2EE39AFF0CFCB0FF44BAE9FF7D" authorityName="Perkins" authorityYear="2017" box="[850,981,223,248]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B958EABFE39AFF0CFCB0FF44BAE9FF7D" box="[850,981,223,248]" italics="true" pageId="157" pageNumber="158">P. elongata</emphasis>
</taxonomicName>
the shape of the lateral margin of the pronotum, which is &quot;dual rimmed,&quot; having an extra ridge above the true (denticulate) lateral margin. Differentiated from both species by the smaller size (ca.
<quantity id="4CD49B48E39AFF0CFD67FE8DBBCAFEBA" box="[645,758,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="157" pageNumber="158" unit="mm" value="1.45">1. 45 mm</quantity>
vs.
<quantity id="4CD49B48E39AFF0CFCCBFE82BAA8FEC5" box="[809,916,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="157" pageNumber="158" unit="mm" value="1.75">1.75 mm</quantity>
). Also differentiated from
<taxonomicName id="4C2C4D2EE39AFF0CFB2BFE8CBC76FEC5" authorityName="Perkins" authorityYear="2017" box="[1225,1354,295,320]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="B958EABFE39AFF0CFB2BFE8CBC76FEC5" box="[1225,1354,295,320]" italics="true" pageId="157" pageNumber="158">P. elongata</emphasis>
</taxonomicName>
by the lighter color, less elongate elytra, and slightly differently shaped pronotum. Also differentiated from
<taxonomicName id="4C2C4D2EE39AFF0CFAEDFEE8BCA0FEE0" box="[1295,1436,332,357]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="ankaratra">
<emphasis id="B958EABFE39AFF0CFAEDFEE8BCA0FEE0" box="[1295,1436,332,357]" italics="true" pageId="157" pageNumber="158">P. ankaratra</emphasis>
</taxonomicName>
by the longer maxillary palpi (palpus length/head width ca. 0.86 vs.
<quantity id="4CD49B48E39AFF0CFC72FECABAC4FE0D" box="[912,1016,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="157" pageNumber="158" unit="mm" value="0.54">0.54 mm</quantity>
). The aedeagus of
<taxonomicName id="4C2C4D2EE39AFF0CFB25FED4BC71FE0D" authorityName="Perkins" authorityYear="2017" box="[1223,1357,367,392]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="duplicata">
<emphasis id="B958EABFE39AFF0CFB25FED4BC71FE0D" box="[1223,1357,367,392]" italics="true" pageId="157" pageNumber="158">P. duplicata</emphasis>
</taxonomicName>
differs distinctively from the compared species.
</paragraph>
</subSubSection>
<caption id="DF536625E39AFF0CFF75F960BAC0F95E" box="[151,1020,1733,1755]" pageId="157" pageNumber="158">
<paragraph id="8B9336ADE39AFF0CFF75F960BAC0F95E" blockId="157.[151,1020,1733,1755]" box="[151,1020,1733,1755]" pageId="157" pageNumber="158">
<emphasis id="B958EABFE39AFF0CFF75F960B810F95F" bold="true" box="[151,300,1733,1755]" pageId="157" pageNumber="158">FIGURE 141.</emphasis>
<taxonomicName id="4C2C4D2EE39AFF0CFED0F963BB10F95E" authorityName="Perkins" authorityYear="2017" box="[306,556,1733,1755]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="duplicata">
<emphasis id="B958EABFE39AFF0CFED0F963BB10F95E" box="[306,556,1733,1755]" italics="true" pageId="157" pageNumber="158">Protozantaena duplicata</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<subSubSection id="C3366526E39AFF0EFF25F8A7BD4BFBDD" lastPageId="159" lastPageNumber="160" pageId="157" pageNumber="158" type="description">
<paragraph id="8B9336ADE39AFF0CFF25F8A7BC40F8E6" blockId="157.[151,1436,1794,2035]" pageId="157" pageNumber="158">
<emphasis id="B958EABFE39AFF0CFF25F8A7B86DF89E" bold="true" box="[199,337,1794,1819]" pageId="157" pageNumber="158">Description</emphasis>
. Size:
<typeStatus id="5497880FE39AFF0CFE41F8A7BB39F89E" box="[419,517,1794,1819]" pageId="157" pageNumber="158" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.47/0.63; head width 0.44; pronotum 0.36/0.57, PA 0.46, PB 0.50; elytra 0.89/0.63. Dorsum dark brown except pronotum anteriorly and laterally lighter; legs light brown; maxillary palpi light brown to testaceous, distal mere acuminate, apex darker.
</paragraph>
<paragraph id="8B9336ADE39AFF0FFF25F8CBB9CDFE98" blockId="157.[151,1436,1794,2035]" lastBlockId="158.[151,1436,151,285]" lastPageId="158" lastPageNumber="159" pageId="157" pageNumber="158">
Head with eyes mod. large, in dorsal aspect ca.78 convex facets in longest series. Ocelli indistinct, round small shining, each located at base of very shallow oblique interocular fovea. Frons effacedly microreticulate, weakly shining, finely punctate, ca. 1xef on disc, finer anteriorly, interstices ca. 13xpd. Clypeus effacedly microreticulate, punctures similar to finest punctures of frons. Labrum dull, length subequal that of clypeus, apicomedially weakly emarginate, anterior margin narrowly upturned; laterally fringed with very short arcuate setae. Maxillary palpi moderately long, combined lengths of meres slightly less than width of head (ratio ca. 30/ 35); ratios of meres 24 ca.
<date id="FF92106DE399FF0FFE33FF45BB1CFF7D" box="[465,544,224,248]" pageId="158" pageNumber="159" value="2013-09-08">13/8/9.</date>
Mentum sparsely punctulate, submentum sparsely finely punctulate, interstices shining.
</paragraph>
<caption id="DF536625E399FF0FFF75F871BBD0F86E" box="[151,748,2004,2027]" pageId="158" pageNumber="159">
<paragraph id="8B9336ADE399FF0FFF75F871BBD0F86E" blockId="158.[151,748,2004,2027]" box="[151,748,2004,2027]" pageId="158" pageNumber="159">
<emphasis id="B958EABFE399FF0FFF75F871B851F86F" bold="true" box="[151,365,2004,2027]" pageId="158" pageNumber="159">FIGURES 142143.</emphasis>
<taxonomicName id="4C2C4D2EE399FF0FFE91F873BB3BF86E" box="[371,519,2006,2027]" class="Insecta" family="Hydraenidae" genus="Protozantaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="158" pageNumber="159" phylum="Arthropoda" rank="genus">
<emphasis id="B958EABFE399FF0FFE91F873BB3BF86E" box="[371,519,2006,2027]" italics="true" pageId="158" pageNumber="159">Protozantaena</emphasis>
</taxonomicName>
, aedeagi of holotypes.
</paragraph>
</caption>
<paragraph id="8B9336ADE398FF0EFF25FF3DBC2EFE29" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">Pronotum broad, anterior margin arcuate, ca. median 1/2 with narrow hyaline border. Anterior angles obtusangulate. Sides weakly rounded, widest at about midlength, weakly attenuate and sinuate behind middle, in lateral view margin with single rim. Posterior angle obtusangulate. Discal relief very finely sparsely punctate, punctures shallow, less than 0.1xef, interstices weakly shining, 36xpd; anterior and posterior impressions shallow, punctures much deeper and much larger than those on relief, some confluent, forming short transverse rows, each puncture with anterior margin extended posteriorly as narrow ridge, dividing puncture into a pair of two smaller punctures each of which ca. 12xef, short recumbent seta arising from posterior extreme of dividing ridge. Lacking impression at each posterior angle. Sculpture laterally subrugulose, dull, with some divided punctures.</paragraph>
<paragraph id="8B9336ADE398FF0EFF25FE12BBCFFD01" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">Elytra weakly shining, attenuate posteriorly, sides arcuate, explanate margins narrow, distinct from shoulder for 3/4 of length, apices separately rounded. Ten-seriate punctate, very shallowly, if at all, striate-impressed, punctures on disc small, round, ca. 12xef, interstices ca. 2xpd, anterior rim of each puncture with minute granule and minute recumbent seta, punctures becoming much smaller and shallower over posterior declivity; intervals ca. 23xpd, each with unilinear row of small punctures, much smaller than serial punctures, anterior margin of each with minute granule and minute short recumbent seta.</paragraph>
<paragraph id="8B9336ADE398FF0EFF25FD2ABA02FCD9" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">Metaventrite with moderately deep, oval impression in post 1/2; midlongitudinal carina between anterior extreme of groove and mesoventral intercoxal process. Metaventrite and abdominal ventrites 14 hydrofuge pubescent. Ventrites 5 and 6 moderately elongate, shining. Ventrite 5 hydrofuge pubescent in anterior 1/2, posterior 1/2 with very sparse pubescence in transverse band. Ventrite 6 very sparsely pubescent in transverse band across posterior 1/2. Last ventrite very sparsely pubescent, free margin markedly arcuate; last tergite with moderately wide apicomedian notch, and bearing moderately long setae.</paragraph>
<paragraph id="8B9336ADE398FF0EFF25FCCDBC03FC69" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">
Legs moderately short, femora moderately broad, metafemora width/metatarsus length ratio ca. 10/20; combined lengths of metatibia and metatarsus slightly greater than width of head (ratio ca. 25/21), ratio of lengths of metatibia/metatarsus ca. 30/20. Basal 3 protarsomeres, in male, with pad of suction setae. Wings fully developed. Females generally slightly larger than males: males ca.
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; females
<quantity id="4CD49B48E398FF0EFB76FC71BC06FC69" box="[1172,1338,980,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.64" metricValueMin="1.52" pageId="159" pageNumber="160" unit="mm" value="1.58" valueMax="1.64" valueMin="1.52">1.521.64 mm</quantity>
.
</paragraph>
<paragraph id="8B9336ADE398FF0EFF25FC52BD4BFBDD" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">
Aedeagus length ca.
<quantity id="4CD49B48E398FF0EFE56FC5DBB20FB8A" box="[436,540,1016,1040]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="159" pageNumber="160" unit="mm" value="0.43">0.43 mm</quantity>
; gonopore bearing process flagelliform, extending slightly beyond distal piece; distal piece bilobate; main piece in ventral view slightly arcuate distally, in lateral view markedly arcuate, tapering apically, with cluster of long setae; parameres very short, each with two setae (
<figureCitation id="13172A28E398FF0EFBEAFB9ABD56FBDD" box="[1032,1130,1087,1112]" captionStart="FIGURES 142 143" captionStartId="158.[151,264,2004,2027]" captionText="FIGURES 142 143. Protozantaena, aedeagi of holotypes." pageId="159" pageNumber="160">Fig. 143</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E398FF0EFF25FBC1BC3FFBF9" box="[199,1283,1124,1149]" pageId="159" pageNumber="160" type="etymology">
<paragraph id="8B9336ADE398FF0EFF25FBC1BC3FFBF9" blockId="159.[151,1437,151,1221]" box="[199,1283,1124,1149]" pageId="159" pageNumber="160">
<emphasis id="B958EABFE398FF0EFF25FBC1B872FBF8" bold="true" box="[199,334,1124,1149]" pageId="159" pageNumber="160">Etymology.</emphasis>
Named in reference to the dual rimmed prothorax and the dual punctured pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E398FF0EFF25FB22BAAEFB41" pageId="159" pageNumber="160" type="distribution">
<paragraph id="8B9336ADE398FF0EFF25FB22BAAEFB41" blockId="159.[151,1437,151,1221]" pageId="159" pageNumber="160">
<emphasis id="B958EABFE398FF0EFF25FB22B85CFB25" bold="true" box="[199,352,1159,1184]" pageId="159" pageNumber="160">Distribution.</emphasis>
Currently known from only the
<typeStatus id="5497880FE398FF0EFD31FB2CBA3FFB24" box="[723,771,1161,1185]" pageId="159" pageNumber="160">type</typeStatus>
locality and one nearby locality, Manjakatompo Station in eastern central
<collectingCountry id="F33B763DE398FF0EFEA2FB09B8F7FB40" box="[320,459,1196,1221]" name="Madagascar" pageId="159" pageNumber="160">Madagascar</collectingCountry>
; elevation 19042073 meters (
<figureCitation id="13172A28E398FF0EFCC1FB09BAB9FB41" box="[803,901,1196,1221]" captionStart="FIGURES 303 306" captionStartId="238.[151,264,1801,1824]" captionTargetBox="[331,1261,203,1770]" captionTargetId="figure@238.[319,1268,193,1780]" captionTargetPageId="238" captionText="FIGURES 303 306. Geographical distributions of Protozantaena species, Sicilicula species, Aulacochthebius species, and Ochthebius alluaudi." httpUri="https://zenodo.org/record/1041156/files/figure.png" pageId="159" pageNumber="160">Fig. 303</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>