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<document id="98702E6A7EA385ECA20E41F946BD787F" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE386FF12FF75FA77BA33FF7D" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena andranomena Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="132" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="130" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:titleInfo id="EA3D039357561715C0BC7C581A0D1412">
<mods:title id="3834E396C74655BBA5F8E70D07B29D71">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="D9DCE16EA6A2A49E8AAB97C41F8175F3" type="personal">
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<mods:roleTerm id="88CB83883F0021E061D7650085584A79">Author</mods:roleTerm>
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<mods:namePart id="9E3336C45BD39FD6830862807077203E">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="02305A425028FC55C56FD47CB3DD9057">text</mods:typeOfResource>
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<mods:title id="A522EB35A03C7E8ADFE8B839FB6702B2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF58188C94866A9AE05908FA0B52A8DE">
<mods:date id="6171389F12969EC4B6C3A2C86B525C61">2017</mods:date>
<mods:detail id="C7A4A975AA71CEC13865C80B0C6FE650" type="pubDate">
<mods:number id="1961776A12EF2E33F9DFAD96C41AF2E6">2017-11-03</mods:number>
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<mods:detail id="ABF6E02457873279AC938B8920221683" type="volume">
<mods:number id="827ABEEA12890EA9F687E5A564116575">4342</mods:number>
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<mods:classification id="344507D06319B64231C559E3849941A9">journal article</mods:classification>
<mods:identifier id="C5DC5FEA5C25847FCEAAC968EE88E765" type="CLB-Dataset">31639</mods:identifier>
<mods:identifier id="B99F892E13F1863182EDE84B1A225DC2" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
<mods:identifier id="301AA6C11DCE579B35C117761670955D" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
<mods:identifier id="6187399FD1F34A737F10838B38BE93A7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038587BBE386FF12FF75FA77BA33FF7D" ID-DOI="http://doi.org/10.5281/zenodo.5323640" ID-GBIF-Taxon="137233432" ID-Zenodo-Dep="5323640" LSID="urn:lsid:plazi:treatment:038587BBE386FF12FF75FA77BA33FF7D" httpUri="http://treatment.plazi.org/id/038587BBE386FF12FF75FA77BA33FF7D" lastPageId="131" lastPageNumber="132" pageId="129" pageNumber="130">
<subSubSection id="C3366526E386FF10FF75FA77BB9EF988" pageId="129" pageNumber="130" type="nomenclature">
<paragraph id="8B9336ADE386FF10FF75FA77BB62FA69" blockId="129.[151,674,1490,1549]" box="[151,606,1490,1516]" pageId="129" pageNumber="130">
<heading id="D0DB81C1E386FF10FF75FA77BB62FA69" bold="true" box="[151,606,1490,1516]" fontSize="11" level="1" pageId="129" pageNumber="130" reason="1">
<emphasis id="B958EABFE386FF10FF75FA77BB62FA69" bold="true" box="[151,606,1490,1516]" pageId="129" pageNumber="130">
<taxonomicName id="4C2C4D2EE386FF10FF75FA77B883FA69" ID-CoL="3N35H" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,447,1490,1516]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="andranomena" status="sp. nov.">
<emphasis id="B958EABFE386FF10FF75FA77B883FA69" bold="true" box="[151,447,1490,1516]" italics="true" pageId="129" pageNumber="130">Hydraena andranomena</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E386FF10FE2CFA77BB62FA69" box="[462,606,1490,1516]" pageId="129" pageNumber="130" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE386FF10FF75FA51BB9EF988" blockId="129.[151,674,1490,1549]" box="[151,674,1524,1549]" pageId="129" pageNumber="130">
<figureCitation id="13172A28E386FF10FF75FA51B9C2F988" box="[151,254,1524,1549]" captionStart="FIGURE 116" captionStartId="130.[151,250,1739,1761]" captionText="FIGURE 116. Hydraena andranomena, holotype, habitus dorsal, lateral, and ventral." pageId="129" pageNumber="130">Figs. 116</figureCitation>
(habitus), 118 (aedeagus), 284 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E386FF10FF75F999BAEEF9FC" pageId="129" pageNumber="130" type="materials_examined">
<paragraph id="8B9336ADE386FF10FF75F999BAEEF9FC" blockId="129.[151,1436,1596,2017]" pageId="129" pageNumber="130">
<materialsCitation id="3B443CF0E386FF10FF75F999BAEEF9FC" ID-GBIF-Occurrence="1697157679" collectingDate="1990-03-30" collectionCode="USNM" collectorName="W. E. Steiner &amp; C. Kremen &amp; V. Razafimahatratra" latitude="-19.883333" location="Andranomena" longLatPrecision="1270" longitude="45.066666" pageId="129" pageNumber="130" specimenCount="2" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="B958EABFE386FF10FF75F999B887F9D0" bold="true" box="[151,443,1596,1621]" pageId="129" pageNumber="130">
<typeStatus id="5497880FE386FF10FF75F999B9EEF9D0" box="[151,210,1596,1621]" pageId="129" pageNumber="130">Type</typeStatus>
Material.
<typeStatus id="5497880FE386FF10FEACF999B887F9D0" box="[334,443,1596,1621]" pageId="129" pageNumber="130" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE386FF10FDFBF999BB51F9D0" box="[537,621,1596,1621]" country="Madagascar" name="Toliara" pageId="129" pageNumber="130">
<emphasis id="B958EABFE386FF10FDFBF999BB51F9D0" bold="true" box="[537,621,1596,1621]" pageId="129" pageNumber="130">Toliara</emphasis>
</collectingRegion>
,
<location id="8EF36076E386FF10FD9FF999BA22F9D0" LSID="urn:lsid:plazi:treatment:038587BBE386FF12FF75FA77BA33FF7D:8EF36076E386FF10FD9FF999BA22F9D0" box="[637,798,1596,1621]" latitude="-19.883333" longLatPrecision="1270" longitude="45.066666" name="Andranomena" pageId="129" pageNumber="130" stateProvince="Toliara">Andranomena</location>
, at black light, dry forest near open marsh,
<geoCoordinate id="EE18506AE386FF10FAE0F998BC55F9D0" box="[1282,1385,1596,1621]" direction="south" orientation="latitude" pageId="129" pageNumber="130" precision="925" value="-19.883333">19° 53' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE386FF10FA97F999B9F8F9FD" direction="east" orientation="longitude" pageId="129" pageNumber="130" precision="925" value="45.066666">45° 4' E</geoCoordinate>
,
<collectingDate id="EFD6E985E386FF10FF32F9C4B870F9FC" box="[208,332,1632,1657]" pageId="129" pageNumber="130" value="1990-03-30">
<date id="FF92106DE386FF10FF32F9C4B870F9FC" box="[208,332,1632,1657]" pageId="129" pageNumber="130" value="1990-03-30">30 iii 1990</date>
</collectingDate>
,
<collectorName id="26D9537BE386FF10FEB5F9C4B8D1F9FC" box="[343,493,1632,1657]" pageId="129" pageNumber="130">W. E. Steiner</collectorName>
,
<collectorName id="26D9537BE386FF10FE1AF9C4BB4AF9FD" box="[504,630,1633,1657]" pageId="129" pageNumber="130">C. Kremen</collectorName>
,
<collectorName id="26D9537BE386FF10FD63F9C4BA5CF9FC" box="[641,864,1632,1657]" pageId="129" pageNumber="130">V. Razafimahatratra</collectorName>
(
<collectionCode id="ED3DAE68E386FF10FC8DF9C4BAFAF9FC" box="[879,966,1633,1657]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="129" pageNumber="130">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3366526E386FF10FF25F921BB8BF88C" pageId="129" pageNumber="130" type="diagnosis">
<paragraph id="8B9336ADE386FF10FF25F921BB8BF88C" blockId="129.[151,1436,1596,2017]" pageId="129" pageNumber="130">
<emphasis id="B958EABFE386FF10FF25F921B8E9F918" bold="true" box="[199,469,1668,1693]" pageId="129" pageNumber="130">Differential Diagnosis.</emphasis>
Somewhat similar in size (ca. 1.49 vs.
<quantity id="4CD49B48E386FF10FC45F920BD2EF918" box="[935,1042,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="129" pageNumber="130" unit="mm" value="1.39">1.39 mm</quantity>
), color, and shape to
<taxonomicName id="4C2C4D2EE386FF10FAEDF920BCA8F919" authorityName="Perkins" authorityYear="2017" box="[1295,1428,1669,1692]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="torquata">
<emphasis id="B958EABFE386FF10FAEDF920BCA8F919" box="[1295,1428,1669,1692]" italics="true" pageId="129" pageNumber="130">H. torquata</emphasis>
</taxonomicName>
; differing externally therefrom in having the pronotum proportionally longer, with much finer and sparser discal punctation, and shallower PF2 foveae; metaventral plaques are larger in
<taxonomicName id="4C2C4D2EE386FF10FC12F968BD4AF961" authorityName="Perkins" authorityYear="2017" box="[1008,1142,1741,1764]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="torquata">
<emphasis id="B958EABFE386FF10FC12F968BD4AF961" box="[1008,1142,1741,1764]" italics="true" pageId="129" pageNumber="130">H. torquata</emphasis>
</taxonomicName>
. The aedeagi of the two species are markedly dissimilar (
<figureCitation id="13172A28E386FF10FDE5F955BB4DF88C" box="[519,625,1776,1801]" captionStart="FIGURES 117 118" captionStartId="131.[151,264,1989,2012]" captionTargetBox="[354,1234,318,1942]" captionTargetId="figure@131.[332,1254,303,1966]" captionTargetPageId="131" captionText="FIGURES 117 118. Aedeagi of Hydraena regimbarti, non-type, and Hydraena andranomena, holotype." httpUri="https://zenodo.org/record/1041116/files/figure.png" pageId="129" pageNumber="130">Figs. 118</figureCitation>
,
<figureCitation id="13172A28E386FF10FD9FF954BB96F88C" box="[637,682,1777,1801]" captionStart="FIGURES 121 122" captionStartId="135.[151,264,1909,1932]" captionText="FIGURES 121 122. Hydraena, aedeagi of holotypes." pageId="129" pageNumber="130">121</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E386FF12FF25F8B1B847FF50" lastPageId="131" lastPageNumber="132" pageId="129" pageNumber="130" type="description">
<paragraph id="8B9336ADE386FF10FF25F8B1BA37F8D4" blockId="129.[151,1436,1596,2017]" pageId="129" pageNumber="130">
<emphasis id="B958EABFE386FF10FF25F8B1B86DF8A8" bold="true" box="[199,337,1812,1837]" pageId="129" pageNumber="130">Description</emphasis>
. Size:
<typeStatus id="5497880FE386FF10FE41F8B1BB39F8A8" box="[419,517,1812,1837]" pageId="129" pageNumber="130" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.49/0.61; head width 0.38; pronotum 0.37/0.51, PA 0.39, PB 0.45; elytra 0.90/0.61.
</paragraph>
<paragraph id="8B9336ADE386FF10FF25F8F9BDFEF81C" blockId="129.[151,1436,1596,2017]" pageId="129" pageNumber="130">Head and pronotal disc dark brown, almost piceous, remainder of dorsum dark brown, except anterior margin of pronotum lighter; legs brown; maxillary palpi brown, distal 0.5 of last palpomere not darker.</paragraph>
<paragraph id="8B9336ADE386FF13FF25F801BBCDFF50" blockId="129.[151,1436,1596,2017]" lastBlockId="130.[151,1437,151,464]" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">Head with frons moderately densely punctate, clypeus finely punctate, microreticulate, dull. Pronotum moderately densely moderately finely punctate in anterior and posterior, disc very finely and sparsely punctate, interstices 36xpd, strongly shining. Mentum finely punctate; postmentum punctate. Genae roundly raised, medially smooth and shining, without posterior ridge.</paragraph>
<paragraph id="8B9336ADE385FF13FF25FF45BAEFFEC5" blockId="130.[151,1437,151,464]" pageId="130" pageNumber="131">Pronotum slightly sinuate laterally, very slightly emarginate between anterior angle and widest part of pronotum; anterior margin straight behind eyes, very slightly emarginate behind frons; PF1 absent; PF2 moderately deep, very shallowly, if at all, confluent medially; PF3 deep; PF4 shallow.</paragraph>
<paragraph id="8B9336ADE385FF13FF25FEE9BB60FE55" blockId="130.[151,1437,151,464]" pageId="130" pageNumber="131">Elytra weakly arcuate laterally; summit of posterior declivity near midlength; lateral explanate margins narrow; serial punctures round, separated by ca. 1xpd, slightly larger than largest pronotal punctures. Intervals not raised, ca. 12xpd, weakly shining. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<caption id="DF536625E385FF13FF75F96EBAC5F964" box="[151,1017,1739,1761]" pageId="130" pageNumber="131">
<paragraph id="8B9336ADE385FF13FF75F96EBAC5F964" blockId="130.[151,1017,1739,1761]" box="[151,1017,1739,1761]" pageId="130" pageNumber="131">
<emphasis id="B958EABFE385FF13FF75F96EB817F965" bold="true" box="[151,299,1739,1761]" pageId="130" pageNumber="131">FIGURE 116.</emphasis>
<taxonomicName id="4C2C4D2EE385FF13FED3F96EBB15F964" authorityName="Perkins" authorityYear="2017" box="[305,553,1739,1761]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="andranomena">
<emphasis id="B958EABFE385FF13FED3F96EBB15F964" box="[305,553,1739,1761]" italics="true" pageId="130" pageNumber="131">Hydraena andranomena</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<paragraph id="8B9336ADE385FF12FF25F8ADB847FF50" blockId="130.[151,1437,1800,2005]" lastBlockId="131.[151,1436,151,248]" lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.5/2/1. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt and raised slightly above metaventral intercoxal process. Plaques small, narrow and indistinct, partially obscured by pubescence, converging slightly anteriorly, not raised, located at sides of median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca. 1.5x P2. Protibia very slightly arcuate, almost straight, slightly widened at ca. distal ¼. Mesotibia straight, widest at ca. midlength. Metatibia straight, slender. Metafemur slender, not markedly arcuate on lateral and medial margins. Abdominal apex with deep asymmetrical notch.</paragraph>
</subSubSection>
<subSubSection id="C3366526E384FF12FF25FF7ABA33FF7D" box="[199,783,223,248]" pageId="131" pageNumber="132" type="etymology">
<paragraph id="8B9336ADE384FF12FF25FF7ABA33FF7D" blockId="131.[151,1436,151,248]" box="[199,783,223,248]" pageId="131" pageNumber="132">
<emphasis id="B958EABFE384FF12FF25FF7AB872FF7D" bold="true" box="[199,334,223,248]" pageId="131" pageNumber="132">Etymology.</emphasis>
Named in reference to the
<typeStatus id="5497880FE384FF12FD9CFF45BB92FF7D" box="[638,686,224,248]" pageId="131" pageNumber="132">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>