319 lines
51 KiB
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319 lines
51 KiB
XML
<document id="F8CFA0AB764B44371E7B96B4046FCD8E" ID-CLB-Dataset="5866" ID-DOI="10.5852/ejt.2021.757.1415" ID-GBIF-Dataset="9950dbcc-5627-479c-b9e7-abe6b8a0ddd1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5047683" ID-ZooBank="F95D404B-0A7C-429F-BC4F-AAEDF9DC870C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625079024571" checkinUser="felipe" docAuthor="Bordera, Santiago & Broad, Gavin R." docDate="2021" docId="03858793FFD7FFA9DAB7FEB67926FB9E" docLanguage="en" docName="ejt-757_80-101.pdf" docOrigin="European Journal of Taxonomy 757" docSource="http://dx.doi.org/10.5852/ejt.2021.757.1415" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Handaoia fritzi Bordera & Broad 2021, sp. nov." docType="treatment" docUuid="3FE16228-86F1-4A8C-8FA0-227999E65A4F" docUuidSource="ZooBank" docVersion="8" lastPageNumber="88" masterDocId="FFBCFFEBFFD1FFA1D83CFFBE781FFF84" masterDocTitle="Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World" masterLastPageNumber="101" masterPageNumber="80" pageNumber="86" updateTime="1699021642505" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DFEE4E4034C85540457AA38AA7DB6D0C">Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World</mods:title>
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<mods:namePart id="14FE24FFDE47922F86379231A508F84E">Bordera, Santiago</mods:namePart>
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<mods:affiliation id="5E74383AB1C3DDF229CFB841ACBCBCD3">Departamento de Ciencias Ambientales y Recursos Naturales, University of Alicante, San Vicente del Raspeig, E- 03690 Alicante, Spain.</mods:affiliation>
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<mods:nameIdentifier id="929817F56DBE1BBCEF72A6370C64DC84" type="email">s.bordera@ua.es</mods:nameIdentifier>
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<mods:namePart id="91395E03F6E9A3060D9790EAA8E82635">Broad, Gavin R.</mods:namePart>
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<mods:affiliation id="9EB6765AC6ED6D308235498DFE4714A8">Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:nameIdentifier id="21C3AAAA02B43F9AA4839354958EB3F9" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="06113A1FF25745EF07D577E7C0C05458">European Journal of Taxonomy</mods:title>
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<mods:date id="67E5DB04D20E9EC459E46AFF1B0F6E63">2021</mods:date>
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<treatment id="03858793FFD7FFA9DAB7FEB67926FB9E" ID-DOI="http://doi.org/10.5281/zenodo.5056068" ID-GBIF-Taxon="183231745" ID-Zenodo-Dep="5056068" LSID="urn:lsid:zoobank.org:act:3FE16228-86F1-4A8C-8FA0-227999E65A4F" httpUri="http://treatment.plazi.org/id/03858793FFD7FFA9DAB7FEB67926FB9E" lastPageId="8" lastPageNumber="88" pageId="6" pageNumber="86">
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<subSubSection id="C336650EFFD7FFA7DAB7FEB67BB6FEA7" box="[651,937,264,291]" pageId="6" pageNumber="86" type="nomenclature">
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<paragraph id="8B933685FFD7FFA7DAB7FEB67BB6FEA7" blockId="6.[375,1212,264,361]" box="[651,937,264,291]" pageId="6" pageNumber="86">
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<heading id="D0DB81E9FFD7FFA7DAB7FEB67BB6FEA7" box="[651,937,264,291]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="86" reason="2">
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<taxonomicName id="4C2C4D06FFD7FFA7DAB7FEB67B58FEA6" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[651,839,264,291]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="86" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
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<emphasis id="B958EA97FFD7FFA7DAB7FEB67B58FEA6" bold="true" box="[651,839,264,291]" italics="true" pageId="6" pageNumber="86">Handaoia fritzi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD7FFA7DB72FEB77BB6FEA7" box="[846,937,265,291]" pageId="6" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C336650EFFD7FFA7D94BFE927B70FEED" pageId="6" pageNumber="86" type="description">
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<paragraph id="8B933685FFD7FFA7D94BFE927CA3FEC3" blockId="6.[375,1212,264,361]" box="[375,1212,300,327]" pageId="6" pageNumber="86">
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<uri id="FFBD3A87FFD7FFA7D94BFE927CA3FEC3" box="[375,1212,300,327]" pageId="6" pageNumber="86">
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urn:lsid:zoobank.org:act:
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<uuid id="FF8A0C50FFD7FFA7DAA4FE927CA3FEC3" box="[664,1212,300,327]" pageId="6" pageNumber="86">3FE16228-86F1-4A8C-8FA0-227999E65A4F</uuid>
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</uri>
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</paragraph>
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<paragraph id="8B933685FFD7FFA7DAF9FEF17B70FEED" blockId="6.[375,1212,264,361]" box="[709,879,335,361]" pageId="6" pageNumber="86">
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<figureCitation id="13172A00FFD7FFA7DAF9FEF17B3DFEED" box="[709,802,335,361]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="6" pageNumber="86">Figs 1B</figureCitation>
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,
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<figureCitation id="13172A00FFD7FFA7DB13FEF17B20FEED" box="[815,831,335,361]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">3</figureCitation>
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,
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<figureCitation id="13172A00FFD7FFA7DB70FEF17B70FEED" box="[844,879,335,361]" captionStart="Fig" captionStartId="19.[186,229,1963,1989]" captionTargetBox="[189,1398,1195,1923]" captionTargetId="figure-309@19.[189,1398,1195,1923]" captionTargetPageId="19" captionText="Fig. 9. Distribution of New World species of Handaoia Seyrig, 1952. A. North and Central America. B. South America." figureDoi="http://doi.org/10.5281/zenodo.5047703" httpUri="https://zenodo.org/record/5047703/files/figure.png" pageId="6" pageNumber="86">9B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD7FFA7D881FE2E792FFD29" pageId="6" pageNumber="86" type="diagnosis">
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<paragraph id="8B933685FFD7FFA7D881FE2E7928FE2E" blockId="6.[189,311,400,426]" box="[189,311,400,426]" pageId="6" pageNumber="86">
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<heading id="D0DB81E9FFD7FFA7D881FE2E7928FE2E" bold="true" box="[189,311,400,426]" fontSize="11" level="3" pageId="6" pageNumber="86" reason="3">
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<emphasis id="B958EA97FFD7FFA7D881FE2E7928FE2E" bold="true" box="[189,311,400,426]" pageId="6" pageNumber="86">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD7FFA7D881FE01792FFD29" blockId="6.[189,1401,447,685]" pageId="6" pageNumber="86">
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<taxonomicName id="4C2C4D06FFD7FFA7D881FE017969FE5D" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[189,374,447,473]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="86" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
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<emphasis id="B958EA97FFD7FFA7D881FE017969FE5D" box="[189,374,447,473]" italics="true" pageId="6" pageNumber="86">Handaoia fritzi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57ECFFD7FFA7D9BDFE7E79C0FE5E" box="[385,479,448,474]" pageId="6" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from all other New World species by the combination of the following characters: apophysis of propodeum as a low transverse rounded or slightly pointed triangular crest (
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<figureCitation id="13172A00FFD7FFA7D9B9FDB879FBFDA4" box="[389,484,518,544]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3G</figureCitation>
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, arrow); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3–0.4 (
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<figureCitation id="13172A00FFD7FFA7D9B7FD9779FBFDC7" box="[395,484,553,579]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3F</figureCitation>
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, arrow); area externa of propodeum separated from area dentipara by strong carina (
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<figureCitation id="13172A00FFD7FFA7D924FDF2796DFDE2" box="[280,370,588,614]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="6" pageNumber="86">Fig. 1B</figureCitation>
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); inner side of area externa 3.0–3.4 × as long as inner side of area dentipara (
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<figureCitation id="13172A00FFD7FFA7DD36FDF27D79FDE2" box="[1290,1382,588,614]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="6" pageNumber="86">Fig. 1B</figureCitation>
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); inner side of area dentipara 0.46–0.5 × the length of outer side (
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<figureCitation id="13172A00FFD7FFA7DB80FDD17C07FD0D" box="[956,1048,623,649]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="6" pageNumber="86">Fig. 1B</figureCitation>
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); tergite II smooth and shiny (
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<figureCitation id="13172A00FFD7FFA7D8FAFD2C793DFD29" box="[198,290,658,685]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD7FFA7D881FD6A7BB2FCC4" pageId="6" pageNumber="86" type="etymology">
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<paragraph id="8B933685FFD7FFA7D881FD6A795AFD6A" blockId="6.[189,325,724,750]" box="[189,325,724,750]" pageId="6" pageNumber="86">
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<heading id="D0DB81E9FFD7FFA7D881FD6A795AFD6A" bold="true" box="[189,325,724,750]" fontSize="11" level="3" pageId="6" pageNumber="86" reason="3">
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<emphasis id="B958EA97FFD7FFA7D881FD6A795AFD6A" bold="true" box="[189,325,724,750]" pageId="6" pageNumber="86">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD7FFA7D881FCBC7BB2FCC4" blockId="6.[189,1399,770,832]" pageId="6" pageNumber="86">
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This species is named in honour of Dr Fritz Plaumann, illustrious botanist and entomologist based in
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<collectingCountry id="F33B7615FFD7FFA7D883FC987915FCC4" box="[191,266,806,832]" name="Brazil" pageId="6" pageNumber="86">Brazil</collectingCountry>
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, who collected the
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<typeStatus id="54978827FFD7FFA7D9D7FC987A01FCC4" box="[491,542,806,832]" pageId="6" pageNumber="86">type</typeStatus>
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material. A noun in genitive case.
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</paragraph>
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</subSubSection>
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<subSubSection id="C336650EFFD7FFA7D881FCD97CBBFA82" pageId="6" pageNumber="86" type="materials_examined">
|
||
<paragraph id="8B933685FFD7FFA7D881FCD979B2FC05" blockId="6.[189,429,871,897]" box="[189,429,871,897]" pageId="6" pageNumber="86">
|
||
<heading id="D0DB81E9FFD7FFA7D881FCD979B2FC05" bold="true" box="[189,429,871,897]" fontSize="11" level="3" pageId="6" pageNumber="86" reason="3">
|
||
<emphasis id="B958EA97FFD7FFA7D881FCD979B2FC05" bold="true" box="[189,429,871,897]" pageId="6" pageNumber="86">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D881FC2879EEFC34" blockId="6.[189,497,918,944]" box="[189,497,918,944]" pageId="6" pageNumber="86">
|
||
Known from
|
||
<specimenCount id="9D2AFD0CFFD7FFA7D967FC2879F2FC34" box="[347,493,918,944]" pageId="6" pageNumber="86" type="female">five females</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D8D6FC697AB6FBB3" blockId="6.[188,1398,983,1080]" pageId="6" pageNumber="86">
|
||
<materialsCitation id="3B443CD8FFD7FFA7D8D6FC697AB6FBB3" ID-GBIF-Occurrence="3312836314" collectingDate="1937-08-26" collectionCode="NHMUK" collectorName="F. Plaumann & B. M. Type Hym" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="6" pageNumber="86" specimenCode="NHMUK010636381" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<emphasis id="B958EA97FFD7FFA7D8D6FC697942FC75" bold="true" box="[234,349,983,1009]" pageId="6" pageNumber="86">
|
||
<typeStatus id="54978827FFD7FFA7D8D6FC697942FC75" box="[234,349,983,1009]" pageId="6" pageNumber="86" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33B7615FFD7FFA7D881FC457939FB91" box="[189,294,1019,1045]" name="Brazil" pageId="6" pageNumber="86">BRAZIL</collectingCountry>
|
||
•
|
||
<specimenCount id="9D2AFD0CFFD7FFA7D900FC45794CFB91" box="[316,339,1019,1045]" pageId="6" pageNumber="86" type="female">♀</specimenCount>
|
||
;
|
||
<location id="8EF3605EFFD7FFA7D95CFC457A10FB90" LSID="urn:lsid:plazi:treatment:03858793FFD7FFA9DAB7FEB67926FB9E:8EF3605EFFD7FFA7D95CFC457A10FB90" box="[352,527,1018,1045]" country="Brazil" latitude="-27.183332" longLatPrecision="1238" longitude="-52.383335" name="Nova Teutonia" pageId="6" pageNumber="86">Nova Teutonia</location>
|
||
;
|
||
<geoCoordinate id="EE185042FFD7FFA7DA26FC447A64FB90" box="[538,635,1017,1045]" degrees="27" direction="south" minutes="11" orientation="latitude" pageId="6" pageNumber="86" precision="925" value="-27.183332">27º11′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE185042FFD7FFA7DABAFC447AEBFB91" box="[646,756,1017,1045]" degrees="52" direction="west" minutes="23" orientation="longitude" pageId="6" pageNumber="86" precision="925" value="-52.383335">52º23′ W</geoCoordinate>
|
||
;
|
||
<date id="FF921045FFD7FFA7DAC3FC447B81FB90" box="[767,926,1018,1045]" pageId="6" pageNumber="86" value="1937-08-26">
|
||
<collectingDate id="EFD6E9ADFFD7FFA7DAC3FC447B81FB90" box="[767,926,1018,1045]" pageId="6" pageNumber="86" value="1937-08-26">26 Aug. 1937</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D95353FFD7FFA7DB95FC457C25FB90" box="[937,1082,1018,1045]" pageId="6" pageNumber="86">F. Plaumann</collectorName>
|
||
leg.; B.M. 1937-748;
|
||
<collectorName id="26D95353FFD7FFA7DD06FC457927FBBC" pageId="6" pageNumber="86">B.M. Type Hym</collectorName>
|
||
3b.2871;
|
||
<specimenCode id="DB8A9EFEFFD7FFA7D992FBA37ABAFBB3" box="[430,677,1053,1079]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="86" type="Museum">NHMUK010636381</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D8D6FBE17CBBFA82" blockId="6.[189,1398,1119,1286]" pageId="6" pageNumber="86">
|
||
<materialsCitation id="3B443CD8FFD7FFA7D8D6FBE17D2BFB18" ID-GBIF-Occurrence="3312836306" collectingDate="1935-08" collectionCode="NHMUK" collectorName="F. Plaumann & B. M. Type Hym" country="Brazil" latitude="-27.183332" location="BRAZIL" longLatPrecision="1238" longitude="-52.383335" pageId="6" pageNumber="86" specimenCode="NHMUK010636388" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<emphasis id="B958EA97FFD7FFA7D8D6FBE17976FBFD" bold="true" box="[234,361,1119,1145]" pageId="6" pageNumber="86">
|
||
<typeStatus id="54978827FFD7FFA7D8D6FBE17976FBFD" box="[234,361,1119,1145]" pageId="6" pageNumber="86" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33B7615FFD7FFA7D881FB3C7938FB18" box="[189,295,1154,1180]" name="Brazil" pageId="6" pageNumber="86">BRAZIL</collectingCountry>
|
||
•
|
||
<specimenCount id="9D2AFD0CFFD7FFA7D904FB3C797DFB19" box="[312,354,1154,1181]" pageId="6" pageNumber="86" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<date id="FF921045FFD7FFA7DB32FB3C7B96FB19" box="[782,905,1154,1181]" pageId="6" pageNumber="86" value="1935-08">
|
||
<collectingDate id="EFD6E9ADFFD7FFA7DB32FB3C7B96FB19" box="[782,905,1154,1181]" pageId="6" pageNumber="86" value="1935-08">Aug.1935</collectingDate>
|
||
</date>
|
||
;B.M. 1937-47;
|
||
<specimenCode id="DB8A9EFEFFD7FFA7DC7DFB3C7D2BFB18" box="[1089,1332,1154,1180]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="86" type="Museum">NHMUK010636388</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B443CD8FFD7FFA7DD7AFB3C7CFAFB3B" ID-GBIF-Occurrence="3312836302" collectingDate="1938-05-28" collectionCode="NHMUK" collectorName="F. Plaumann & B. M. Type Hym" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="6" pageNumber="86" specimenCode="NHMUK010636387" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D2AFD0CFFD7FFA7DD7AFB3C7D70FB19" box="[1350,1391,1154,1181]" pageId="6" pageNumber="86" type="female">1♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="FF921045FFD7FFA7DABDFB1B7B3DFB3B" box="[641,802,1189,1216]" pageId="6" pageNumber="86" value="1938-05-28">
|
||
<collectingDate id="EFD6E9ADFFD7FFA7DABDFB1B7B3DFB3B" box="[641,802,1189,1216]" pageId="6" pageNumber="86" value="1938-05-28">28 May 1938</collectingDate>
|
||
</date>
|
||
; B.M. 1938-458;
|
||
<specimenCode id="DB8A9EFEFFD7FFA7DBCEFB1B7CFAFB3B" box="[1010,1253,1189,1215]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="86" type="Museum">NHMUK010636387</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B443CD8FFD7FFA7DCC2FB1B7CCDFB66" ID-GBIF-Occurrence="3312836307" collectingDate="1938-08-11" collectionCode="NHMUK" collectorName="F. Plaumann & B. M. Type Hym" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="6" pageNumber="86" specimenCode="NHMUK010636396" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D2AFD0CFFD7FFA7DCC2FB1B7D34FB44" box="[1278,1323,1189,1216]" pageId="6" pageNumber="86" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="FF921045FFD7FFA7DA69FB777B1FFB67" box="[597,768,1225,1251]" pageId="6" pageNumber="86" value="1938-08-11">
|
||
<collectingDate id="EFD6E9ADFFD7FFA7DA69FB777B1FFB67" box="[597,768,1225,1251]" pageId="6" pageNumber="86" value="1938-08-11">11 Aug. 1938</collectingDate>
|
||
</date>
|
||
; B.M. 1938-682;
|
||
<specimenCode id="DB8A9EFEFFD7FFA7DBE3FB767CCDFB66" box="[991,1234,1224,1250]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="86" type="Museum">NHMUK010636396</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B443CD8FFD7FFA7DCC9FB777CBEFA82" ID-GBIF-Occurrence="3312836309" collectingDate="1938-09-16" collectionCode="NHMUK" collectorName="F. Plaumann & B. M. Type Hym" country="Brazil" latitude="-27.183332" location="Nova Teutonia" longLatPrecision="1238" longitude="-52.383335" pageId="6" pageNumber="86" specimenCode="NHMUK010636383" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D2AFD0CFFD7FFA7DCC9FB777D38FB67" box="[1269,1319,1225,1251]" pageId="6" pageNumber="86" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<date id="FF921045FFD7FFA7DA01FB527AC5FA82" box="[573,730,1260,1286]" pageId="6" pageNumber="86" value="1938-09-16">
|
||
<collectingDate id="EFD6E9ADFFD7FFA7DA01FB527AC5FA82" box="[573,730,1260,1286]" pageId="6" pageNumber="86" value="1938-09-16">16 Sep. 1938</collectingDate>
|
||
</date>
|
||
; B.M. 1938-682;
|
||
<specimenCode id="DB8A9EFEFFD7FFA7DB96FB527CBEFA82" box="[938,1185,1260,1286]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="86" type="Museum">NHMUK010636383</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336650EFFD7FFA9D881FA937926FB9E" lastPageId="8" lastPageNumber="88" pageId="6" pageNumber="86" type="description">
|
||
<paragraph id="8B933685FFD7FFA7D881FA937950FAC3" blockId="6.[189,335,1325,1351]" box="[189,335,1325,1351]" pageId="6" pageNumber="86">
|
||
<heading id="D0DB81E9FFD7FFA7D881FA937950FAC3" bold="true" box="[189,335,1325,1351]" fontSize="11" level="3" pageId="6" pageNumber="86" reason="3">
|
||
<emphasis id="B958EA97FFD7FFA7D881FA937950FAC3" bold="true" box="[189,335,1325,1351]" pageId="6" pageNumber="86">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D8D6FAE27959FAF2" blockId="6.[189,1265,1372,1434]" box="[234,326,1372,1398]" pageId="6" pageNumber="86">
|
||
<emphasis id="B958EA97FFD7FFA7D8D6FAE27959FAF2" bold="true" box="[234,326,1372,1398]" pageId="6" pageNumber="86">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D881FAC17CEEFA1D" blockId="6.[189,1265,1372,1434]" box="[189,1265,1407,1434]" pageId="6" pageNumber="86">
|
||
MEASUREMENTS. Body length (without ovipositor)
|
||
<quantity id="4CD49B60FFD7FFA7DB3AFAC17B89FA1D" box="[774,918,1407,1433]" metricMagnitude="-3" metricUnit="m" metricValue="3.6999999999999997" metricValueMax="4.1" metricValueMin="3.3" pageId="6" pageNumber="86" unit="mm" value="3.6999999999999997" valueMax="4.1" valueMin="3.3">3.3–4.1 mm</quantity>
|
||
. Fore wing
|
||
<quantity id="4CD49B60FFD7FFA7DC1CFAC17CB1FA1D" box="[1056,1198,1407,1433]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.3" metricValueMin="2.7" pageId="6" pageNumber="86" unit="mm" value="3.0" valueMax="3.3" valueMin="2.7">2.7–3.3 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA7D881FA7F7C5EF893" blockId="6.[187,1399,1472,1816]" pageId="6" pageNumber="86">
|
||
HEAD. Finely granulate, matt, distinctly narrowed behind eyes, occiput very concave centrally in dorsal view (
|
||
<figureCitation id="13172A00FFD7FFA7D934FA5D7979FA7A" box="[264,358,1507,1534]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3C</figureCitation>
|
||
); gena 0.2–0.25 × length of eye in dorsal view; posterior ocellus separated from eye by 0.75–1.25 × its own maximum diameter; distance between posterior ocelli 0.75–1.0 × maximum ocellar diameter (
|
||
<figureCitation id="13172A00FFD7FFA7D908F994798EF9C0" box="[308,401,1578,1604]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3C</figureCitation>
|
||
); occipital carina gently V-shaped medially (
|
||
<figureCitation id="13172A00FFD7FFA7DB9EF9947BE1F9C0" box="[930,1022,1578,1604]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3B</figureCitation>
|
||
); malar space 1.15–1.25 × basal width of mandible; face slightly swollen medially; clypeus (
|
||
<figureCitation id="13172A00FFD7FFA7DBBBF9F37BFEF9E3" box="[903,993,1613,1639]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3E</figureCitation>
|
||
) weakly convex, faintly granulate dorsally, smooth and shiny ventrally, with scattered punctures, its apex almost straight, 2.1–2.2× as wide as high; mandible slightly tapered to apex, ventral tooth of mandible approximately same length as upper tooth; maxillary palp relatively short, reaching at most ⅔ length of mesosternum; flagellum widened in middle, tapered towards apex, flagellomeres 8–17 flattened ventrally; flagellum with 19–20 segments, length-thickness ratios: 1
|
||
<superScript id="7C599BCDFFD7FFA7D9D8F94379F0F88F" attach="left" box="[484,495,1789,1803]" fontSize="6" pageId="6" pageNumber="86">st</superScript>
|
||
segment = 4.74–5.0, 2
|
||
<superScript id="7C599BCDFFD7FFA7DAC0F9427B12F88E" attach="left" box="[764,781,1788,1802]" fontSize="6" pageId="6" pageNumber="86">nd</superScript>
|
||
= 5.0–5.28 and 10
|
||
<superScript id="7C599BCDFFD7FFA7DBD5F9427BE9F88E" attach="left" box="[1001,1014,1788,1802]" fontSize="6" pageId="6" pageNumber="86">th</superScript>
|
||
= 1.0.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD7FFA9D881F8817955FE57" blockId="6.[189,1399,1855,2022]" lastBlockId="8.[189,1399,265,467]" lastPageId="8" lastPageNumber="88" pageId="6" pageNumber="86">
|
||
MESOSOMA. Finely granulate and matt (
|
||
<figureCitation id="13172A00FFD7FFA7DA9AF8817B31F8DD" box="[678,814,1855,1881]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3B–D</figureCitation>
|
||
); mesoscutum convex, notauli weak anteriorly, reaching about 0.3 distance to scuto-scutellar groove, sometimes inconspicuous (
|
||
<figureCitation id="13172A00FFD7FFA7DCB0F8DC7CF3F8F8" box="[1164,1260,1890,1916]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3C</figureCitation>
|
||
); scutellum moderately convex, without lateral carinae; sternaulus moderately deep, reaching at least middle of mesopleuron (
|
||
<figureCitation id="13172A00FFD7FFA7D954F81679D5F846" box="[360,458,1960,1986]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3D</figureCitation>
|
||
); juxtacoxal carina strong, irregular, joining submetapleural carina at anterior 0.3–0.4 (
|
||
<figureCitation id="13172A00FFD7FFA7D91AF8757962F862" box="[294,381,1995,2022]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="6" pageNumber="86">Fig. 3F</figureCitation>
|
||
, arrow); propodeum (
|
||
<figureCitation id="13172A00FFD7FFA7DA42F8757AC7F862" box="[638,728,1995,2022]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="6" pageNumber="86">Fig. 1B</figureCitation>
|
||
) with strong and conspicuous carinae; area superomedia confluent with area basalis and separated from area petiolaris; inner side of area externa about 3.0– 4.0× as long as inner side of area dentipara; inner side of area dentipara 0.46–0.5× length of outer side; area spiracularis confluent with area lateralis; posterior transverse carina strong, forming a low and wide rounded or slightly subtriangular crest joining lateral longitudinal carina (
|
||
<figureCitation id="13172A00FFD9FFA9DC0FFECC7C8CFE09" box="[1075,1171,370,397]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3G</figureCitation>
|
||
, arrow). Length of hind femur 4.0–4.7 × its height. Hind wing with vein
|
||
<emphasis id="B958EA97FFD9FFA9DB0CFE297B7AFE34" box="[816,869,407,432]" italics="true" pageId="8" pageNumber="88">cu-a</emphasis>
|
||
0.4–0.6 × as long as abscissa of
|
||
<emphasis id="B958EA97FFD9FFA9DCD8FE287D12FE34" box="[1252,1293,406,432]" italics="true" pageId="8" pageNumber="88">CU</emphasis>
|
||
between
|
||
<emphasis id="B958EA97FFD9FFA9D881FE0478CAFE57" box="[189,213,442,467]" italics="true" pageId="8" pageNumber="88">M</emphasis>
|
||
and
|
||
<emphasis id="B958EA97FFD9FFA9D933FE04795BFE57" box="[271,324,442,467]" italics="true" pageId="8" pageNumber="88">cu-a</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF53660DFFD6FFA6D881F8DA7BFDF86C" ID-DOI="http://doi.org/10.5281/zenodo.5047691" ID-Zenodo-Dep="5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="7" pageNumber="87" startId="7.[189,232,1892,1918]" targetBox="[189,1398,333,1851]" targetPageId="7">
|
||
<paragraph id="8B933685FFD6FFA6D881F8DA7BFDF86C" blockId="7.[189,1398,1892,2024]" pageId="7" pageNumber="87">
|
||
<emphasis id="B958EA97FFD6FFA6D881F8DA7914F8FA" bold="true" box="[189,267,1892,1918]" pageId="7" pageNumber="87">Fig. 3.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C2C4D06FFD6FFA6D9F2F8DA7A9BF8FA" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[462,644,1892,1918]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="87" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD6FFA6D9F2F8DA7A9BF8FA" box="[462,644,1892,1918]" italics="true" pageId="7" pageNumber="87">Handaoia fritzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD6FFA6DAB0F8DA7AF7F8FA" box="[652,744,1892,1918]" pageId="7" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♀ (NHMUK010636396).
|
||
<emphasis id="B958EA97FFD6FFA6DCA3F8DA7CABF8FA" bold="true" box="[1183,1204,1892,1918]" pageId="7" pageNumber="87">A</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="B958EA97FFD6FFA6D938F8397907F825" bold="true" box="[260,280,1927,1953]" pageId="7" pageNumber="87">B</emphasis>
|
||
. Habitus, dorsal view.
|
||
<emphasis id="B958EA97FFD6FFA6DA17F8397A20F825" bold="true" box="[555,575,1927,1953]" pageId="7" pageNumber="87">C</emphasis>
|
||
. Head and mesoscutum, dorsal view.
|
||
<emphasis id="B958EA97FFD6FFA6DC38F8397C07F825" bold="true" box="[1028,1048,1927,1953]" pageId="7" pageNumber="87">D</emphasis>
|
||
. Head and mesosoma, lateral view.
|
||
<emphasis id="B958EA97FFD6FFA6D93EF8147909F840" bold="true" box="[258,278,1962,1988]" pageId="7" pageNumber="87">E</emphasis>
|
||
. Head, front view.
|
||
<emphasis id="B958EA97FFD6FFA6D9C9F8147A18F840" bold="true" box="[501,519,1962,1988]" pageId="7" pageNumber="87">F</emphasis>
|
||
. Metapleuron (arrow: juxtacoxal carina).
|
||
<emphasis id="B958EA97FFD6FFA6DBCCF8147C18F840" bold="true" box="[1008,1031,1962,1988]" pageId="7" pageNumber="87">G</emphasis>
|
||
. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B933685FFD9FFA9D881FE457D59FD25" blockId="8.[188,1399,506,674]" pageId="8" pageNumber="88">
|
||
METASOMA. Tergite I (
|
||
<figureCitation id="13172A00FFD9FFA9D9F9FE447A38FD91" box="[453,551,506,533]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="8" pageNumber="88">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD9FFA9DA04FE457A43FD91" box="[568,604,507,533]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">3B</figureCitation>
|
||
), finely granulate, matt, 1.8–2.0 × as long as its maximum width; postpetiole laterally and posteriorly tending to fine strigosity; lateromedian longitudinal carina weak, reaching at least 0.6 × length of tergite; lateral longitudinal carina weak, complete or absent posterior to spiracles; tergite II (
|
||
<figureCitation id="13172A00FFD9FFA9D99AFDDA7A1FFDFA" box="[422,512,612,638]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3B</figureCitation>
|
||
) smooth and shiny, glabrous; remaining tergites (
|
||
<figureCitation id="13172A00FFD9FFA9DC7CFDDA7C86FDFA" box="[1088,1177,612,638]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3B</figureCitation>
|
||
) smooth and shiny with inconspicuous dense setiferous punctures. Ovipositor sheath 0.45–0.49 × as long as hind tibia.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD9FFA9D881FD777C52FC32" blockId="8.[188,1399,713,950]" pageId="8" pageNumber="88">
|
||
COLOUR. Head (
|
||
<figureCitation id="13172A00FFD9FFA9D943FD777A1EFD67" box="[383,513,713,739]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3A–E</figureCitation>
|
||
) dark brown, clypeus and mandibles reddish brown; palpi light brown; antenna with scape, pedicel and flagellomeres 1–4 light brown, darker dorsally, remaining flagellomeres dark brown, flagellomeres 4–7 dorsally white (
|
||
<figureCitation id="13172A00FFD9FFA9DAD1FCB17B52FCAD" box="[749,845,783,809]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3A</figureCitation>
|
||
). Mesosoma (
|
||
<figureCitation id="13172A00FFD9FFA9DBC9FCB17C68FCAD" box="[1013,1143,783,809]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3A–D</figureCitation>
|
||
) mostly dark orange; mesoscutum with wide dark brown longitudinal bands on central and lateral lobes; dorsal anterior part of propodeum dark brown. Metasoma (
|
||
<figureCitation id="13172A00FFD9FFA9DABBFCEB7AFFFCF4" box="[647,736,853,880]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3B</figureCitation>
|
||
) mostly brown, tergite I sometimes centrally tinged dark brown. Legs (
|
||
<figureCitation id="13172A00FFD9FFA9D962FCC779A5FC17" box="[350,442,889,915]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3A</figureCitation>
|
||
) light brown, distal tarsomeres dark brown, fore and mid coxae, part of hind coxa and all trochanters and trochantelli lighter. Wings (
|
||
<figureCitation id="13172A00FFD9FFA9DB2CFC227B73FC32" box="[784,876,924,950]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="8" pageNumber="88">Fig. 3A</figureCitation>
|
||
) slightly infuscate.
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD9FFA9D8D6FC637935FC73" blockId="8.[189,313,989,1050]" box="[234,298,989,1015]" pageId="8" pageNumber="88">
|
||
<emphasis id="B958EA97FFD9FFA9D8D6FC637935FC73" bold="true" box="[234,298,989,1015]" pageId="8" pageNumber="88">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD9FFA9D881FBBE7926FB9E" blockId="8.[189,313,989,1050]" box="[189,313,1024,1050]" pageId="8" pageNumber="88">Unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |