307 lines
61 KiB
XML
307 lines
61 KiB
XML
<document id="459DAD83064636CCDF1D516134FDE2E5" ID-CLB-Dataset="5866" ID-DOI="10.5852/ejt.2021.757.1415" ID-GBIF-Dataset="9950dbcc-5627-479c-b9e7-abe6b8a0ddd1" ID-ISSN="2118-9773" ID-Zenodo-Dep="5047683" ID-ZooBank="F95D404B-0A7C-429F-BC4F-AAEDF9DC870C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625079024571" checkinUser="felipe" docAuthor="Bordera, Santiago & Broad, Gavin R." docDate="2021" docId="03858793FFD0FFA3DA58FA7D7D68F85B" docLanguage="en" docName="ejt-757_80-101.pdf" docOrigin="European Journal of Taxonomy 757" docSource="http://dx.doi.org/10.5852/ejt.2021.757.1415" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Handaoia Seyrig 1952" docType="key" docVersion="10" lastPageNumber="82" masterDocId="FFBCFFEBFFD1FFA1D83CFFBE781FFF84" masterDocTitle="Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World" masterLastPageNumber="101" masterPageNumber="80" pageNumber="81" updateTime="1699021642505" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="22F991AB81B1BB57A59AD253E54C2077">Six new species of Handaoia Seyrig, 1952 (Hymenoptera, Ichneumonidae, Phygadeuontinae): the first to be described from the New World</mods:title>
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<mods:roleTerm id="517342011C2D84C1E1C3570BC930ABBC">Author</mods:roleTerm>
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<mods:namePart id="73528E61A41230CB39131C34073E736F">Bordera, Santiago</mods:namePart>
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<mods:affiliation id="6EF94B591E0EF42658AF93CF6657AC93">Departamento de Ciencias Ambientales y Recursos Naturales, University of Alicante, San Vicente del Raspeig, E- 03690 Alicante, Spain.</mods:affiliation>
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<mods:nameIdentifier id="A83265A2B4BD388198CFB15B5A9C773B" type="email">s.bordera@ua.es</mods:nameIdentifier>
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<mods:namePart id="C8122E20D3C4EDCAECB48ABB9058357A">Broad, Gavin R.</mods:namePart>
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<mods:affiliation id="11CE04DC064D3490E3D1A99FE232DD31">Department of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<mods:nameIdentifier id="AA4DFCD5898144F20013F0EE0BB82108" type="email">g.broad@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="BFF395EF36516A14B99FD393B88A55A5">European Journal of Taxonomy</mods:title>
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<mods:part id="39C060B4F85FFE967B3157DF06003751">
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<mods:date id="C8E7EB497DFA4C8A060CA926E88E9544">2021</mods:date>
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<mods:number id="C6F7229E5D4AD65D90BE19BE0DA6EB31">2021-06-30</mods:number>
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<treatment id="03858793FFD0FFA3DA58FA7D7D68F85B" ID-DOI="http://doi.org/10.5281/zenodo.5515757" ID-GBIF-Taxon="183231741" ID-Zenodo-Dep="5515757" LSID="urn:lsid:plazi:treatment:03858793FFD0FFA3DA58FA7D7D68F85B" httpUri="http://treatment.plazi.org/id/03858793FFD0FFA3DA58FA7D7D68F85B" lastPageId="2" lastPageNumber="82" pageId="1" pageNumber="81">
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<subSubSection id="C336650EFFD0FFA0DA58FA7D7BD0FA5A" box="[612,975,1474,1502]" pageId="1" pageNumber="81" type="nomenclature">
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<paragraph id="8B933685FFD0FFA0DA58FA7D7BD0FA5A" blockId="1.[612,975,1474,1502]" box="[612,975,1474,1502]" pageId="1" pageNumber="81">
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<heading id="D0DB81E9FFD0FFA0DA58FA7D7BD0FA5A" box="[612,975,1474,1502]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="81" reason="2">
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<emphasis id="B958EA97FFD0FFA0DA58FA7D7AAFFA59" box="[612,688,1475,1501]" italics="true" pageId="1" pageNumber="81">Genus</emphasis>
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<taxonomicName id="4C2C4D06FFD0FFA0DA84FA7C7BD0FA5A" ID-CoL="4T7S" ID-ENA="494752" authority="Seyrig, 1952" authorityName="Seyrig" authorityYear="1952" box="[696,975,1474,1502]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EA97FFD0FFA0DA84FA7C7B2DFA59" bold="true" box="[696,818,1474,1501]" italics="true" pageId="1" pageNumber="81">Handaoia</emphasis>
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<bibRefCitation id="EFBD4B74FFD0FFA0DB05FA7D7BD0FA5A" author="Seyrig A." box="[825,975,1475,1502]" pageId="1" pageNumber="81" pagination="1 - 213" refId="ref10102" refString="Seyrig A. 1952. Les Ichneumonides de Madagascar. IV Ichneumonidae Cryptinae. Memoires de l'Academie malgache. Fascicule XIX: 1 - 213." type="book chapter" year="1952">Seyrig, 1952</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<treatmentCitationGroup id="AB3C11ABFFD0FFA0D881F9BA7D4FF99B" box="[189,1360,1540,1567]" pageId="1" pageNumber="81">
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<subSubSection id="C336650EFFD0FFA0D881F9BA79E2F99A" box="[189,509,1540,1567]" pageId="1" pageNumber="81" type="reference_group">
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<paragraph id="8B933685FFD0FFA0D881F9BA79E2F99A" blockId="1.[189,1360,1540,1567]" box="[189,509,1540,1567]" pageId="1" pageNumber="81">
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<treatmentCitation id="0A8D1094FFD0FFA0D881F9BA79E2F99A" author="Seyrig A." box="[189,509,1540,1567]" page="26" pageId="1" pageNumber="81" year="1952">
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<taxonomicName id="4C2C4D06FFD0FFA0D881F9BA79E2F99A" ID-CoL="4T7S" authority="Seyrig, 1952: 26" authorityName="Seyrig" authorityPageNumber="26" authorityYear="1952" box="[189,509,1540,1567]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EA97FFD0FFA0D881F9BA792CF99A" box="[189,307,1540,1566]" italics="true" pageId="1" pageNumber="81">Handaoia</emphasis>
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<bibRefCitation id="EFBD4B74FFD0FFA0D907F9BA79E2F99A" author="Seyrig A." box="[315,509,1540,1567]" pageId="1" pageNumber="81" pagination="1 - 213" refId="ref10102" refString="Seyrig A. 1952. Les Ichneumonides de Madagascar. IV Ichneumonidae Cryptinae. Memoires de l'Academie malgache. Fascicule XIX: 1 - 213." type="book chapter" year="1952">Seyrig, 1952: 26</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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<subSubSection id="C336650EFFD0FFA0DA38F9BB7D4FF99B" box="[516,1360,1540,1567]" pageId="1" pageNumber="81" type="type_taxon">
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<paragraph id="8B933685FFD0FFA0DA38F9BB7D4FF99B" blockId="1.[189,1360,1540,1567]" box="[516,1360,1540,1567]" pageId="1" pageNumber="81">
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(type species:
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<taxonomicName id="4C2C4D06FFD0FFA0DA97F9BA7C3BF99B" ID-CoL="3JJNZ" authority="Seyrig, 1952" authorityName="Seyrig" authorityYear="1952" box="[683,1060,1540,1567]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa">
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<emphasis id="B958EA97FFD0FFA0DA97F9BA7B9BF99A" box="[683,900,1540,1566]" italics="true" pageId="1" pageNumber="81">Handaoia spinosa</emphasis>
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<bibRefCitation id="EFBD4B74FFD0FFA0DBB0F9BA7C3BF99B" author="Seyrig A." box="[908,1060,1540,1567]" pageId="1" pageNumber="81" pagination="1 - 213" refId="ref10102" refString="Seyrig A. 1952. Les Ichneumonides de Madagascar. IV Ichneumonidae Cryptinae. Memoires de l'Academie malgache. Fascicule XIX: 1 - 213." type="book chapter" year="1952">Seyrig, 1952</bibRefCitation>
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</taxonomicName>
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, by original designation).
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</paragraph>
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</treatmentCitationGroup>
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<subSubSection id="C336650EFFD0FFA3D881F9F87BCCFE57" lastPageId="2" lastPageNumber="82" pageId="1" pageNumber="81" type="diagnosis">
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<paragraph id="8B933685FFD0FFA0D881F9F87928F9E4" blockId="1.[189,311,1606,1632]" box="[189,311,1606,1632]" pageId="1" pageNumber="81">
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<heading id="D0DB81E9FFD0FFA0D881F9F87928F9E4" bold="true" box="[189,311,1606,1632]" fontSize="11" level="3" pageId="1" pageNumber="81" reason="3">
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<emphasis id="B958EA97FFD0FFA0D881F9F87928F9E4" bold="true" box="[189,311,1606,1632]" pageId="1" pageNumber="81">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B933685FFD0FFA3D881F9CA7BCCFE57" blockId="1.[189,1400,1652,1996]" lastBlockId="2.[188,1398,265,468]" lastPageId="2" lastPageNumber="82" pageId="1" pageNumber="81">
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<taxonomicName id="4C2C4D06FFD0FFA0D881F9CA792CF90A" authorityName="Seyrig" authorityYear="1952" box="[189,307,1652,1678]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EA97FFD0FFA0D881F9CA792CF90A" box="[189,307,1652,1678]" italics="true" pageId="1" pageNumber="81">Handaoia</emphasis>
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</taxonomicName>
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can be recognized and separated from other
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<taxonomicName id="4C2C4D06FFD0FFA0DB63F9CA7C36F90A" authorityName="Forster" authorityYear="1869" box="[863,1065,1652,1678]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="subFamily" subFamily="Phygadeuontinae">Phygadeuontinae</taxonomicName>
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genera by the combination of the distally expanded and ventrally flattened antennal flagellum, complete posterior transverse carina of the mesosternum, isolated ʻpit’ (episternal scrobe) in the mesopleuron, and a single bulla in fore wing vein 2
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<emphasis id="B958EA97FFD0FFA0D9BFF96179A1F97C" box="[387,446,1759,1784]" italics="true" pageId="1" pageNumber="81">m-cu</emphasis>
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. The following more complete diagnosis is modified from
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<bibRefCitation id="EFBD4B74FFD0FFA0DCBDF9617D29F97C" author="Townes H." box="[1153,1334,1758,1785]" pageId="1" pageNumber="81" pagination="1 - 537" refId="ref10198" refString="Townes H. 1970. Genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute 12: 1 - 537." type="journal article" year="1970">Townes (1970)</bibRefCitation>
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, who included
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<taxonomicName id="4C2C4D06FFD0FFA0D914F8BF7981F89F" authorityName="Seyrig" authorityYear="1952" box="[296,414,1793,1819]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="81" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EA97FFD0FFA0D914F8BF7981F89F" box="[296,414,1793,1819]" italics="true" pageId="1" pageNumber="81">Handaoia</emphasis>
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</taxonomicName>
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in a key to the genera of ‘Chiroticina’. Townes’ definition applies equally well to the Afrotropical species originally included as well as to the Neotropical and Palaearctic species: flagellum of female antenna fusiform (expanded beyond the middle and flattened ventrally) with a median white ring; genal carina reaching base of mandible; mandibular teeth subequal; maxillary palp long, reaching beyond centre of mesosternum; pronotum with two dorsal pits separated by weak longitudinal carina, usually flanked laterally by 3–4 longitudinal carinae; epomia absent; median lobe of mesoscutum without median longitudinal groove; scutellum without lateral carinae; mesopleural impression (episternal scrobe) ventral to speculum as an isolated pit, some distance from mesopleural furrow; posterior transverse carina of mesosternum complete; fore wing with areolet pentagonal, its outer side open (vein 3
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<emphasis id="B958EA97FFD3FFA3D8F0FECD791DFE08" box="[204,258,371,396]" italics="true" pageId="2" pageNumber="82">rs-m</emphasis>
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missing), vein 2
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<emphasis id="B958EA97FFD3FFA3D9FEFECD79E2FE08" box="[450,509,371,396]" italics="true" pageId="2" pageNumber="82">m-cu</emphasis>
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with one wide bulla; area superomedia always separated from area petiolaris and usually confluent with area basalis; laterotergites of metasomal tergites 2 and 3 not separated or only weakly separated from tergite. Small species, body length
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<quantity id="4CD49B60FFD3FFA3DB56FE077BD0FE57" box="[874,975,441,468]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="82" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C336650EFFD3FFA3D881FE457AC8FD91" box="[189,727,506,533]" pageId="2" pageNumber="82" type="discussion">
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<paragraph id="8B933685FFD3FFA3D881FE457AC8FD91" blockId="2.[189,727,506,533]" box="[189,727,506,533]" pageId="2" pageNumber="82">
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<heading id="D0DB81E9FFD3FFA3D881FE457AC8FD91" bold="true" box="[189,727,506,533]" fontSize="11" level="3" pageId="2" pageNumber="82" reason="3">
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<emphasis id="B958EA97FFD3FFA3D881FE457AC8FD91" bold="true" box="[189,727,506,533]" pageId="2" pageNumber="82">
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Key to the New World species of
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<taxonomicName id="4C2C4D06FFD3FFA3DA61FE447AC8FD91" authorityName="Seyrig" authorityYear="1952" box="[605,727,506,533]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B958EA97FFD3FFA3DA61FE447AC8FD91" bold="true" box="[605,727,506,533]" italics="true" pageId="2" pageNumber="82">Handaoia</emphasis>
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</taxonomicName>
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||
</emphasis>
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||
</heading>
|
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C336650EFFD3FFA3D881FD977D68F85B" pageId="2" pageNumber="82" type="key">
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<key id="FFBD61B4FFD3FFA3D881FD977D68F85B" pageId="2" pageNumber="82">
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<keyStep id="30D82E20FFD3FFA3D881FD977D68FD29" pageId="2" pageNumber="82">
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<paragraph id="8B933685FFD3FFA3D881FD977D68FDE2" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
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<keyLead id="30DD95B0FFD3FFA3D881FD977D68FDE2" pageId="2" pageNumber="82">
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1. Apophysis of propodeum high, parallel sided or subtriangular, with blunt tip (
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<figureCitation id="13172A00FFD3FFA3DC91FD977D13FDC7" box="[1197,1292,553,579]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="2" pageNumber="82">Figs 2F</figureCitation>
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,
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<figureCitation id="13172A00FFD3FFA3DD28FD977D24FDC7" box="[1300,1339,553,579]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">4G</figureCitation>
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,
|
||
<figureCitation id="13172A00FFD3FFA3DD77FD977D6DFDC7" box="[1355,1394,553,579]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">7H</figureCitation>
|
||
, arrows) .............................................................................................................................................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD3FFA3D8FDFDCE7D68FD29" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D8FDFDCE7D68FD29" pageId="2" pageNumber="82">
|
||
– Apophysis of propodeum as a low transverse rounded crest, or slightly pointed (
|
||
<figureCitation id="13172A00FFD3FFA3DC85FDD17D02FD0D" box="[1209,1309,623,649]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Figs 3G</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3DD11FDD17D58FD0E" box="[1325,1351,623,650]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[221,1366,267,1709]" captionTargetId="figure-11@12.[221,1366,267,1712]" captionTargetPageId="12" captionText="Fig. 5. Morphology of Handaoia plaumanni sp. nov. A. Habitus, lateral view, holotype, ♀ (NHMUK010636386). B. Head, dorsal view, paratype, ♀ (NHMUK010636395). C. Head, front view, paratype, ♀ (NHMUK010636382). D. Mesoscutum, dorsal view, paratype, ♀ (NHMUK010636395). E. Head and mesosoma, lateral view, paratype, ♀ (NHMUK010636395). F. Metasoma, lateral view, paratype, ♀ (NHMUK010636382). G. Metasoma, dorsal view, paratype, ♀ (NHMUK010636382). H. Metapleuron, paratype, ♀ (NHMUK010636395). I. Propodeum, paratype, ♀ (NHMUK010636382) (arrow: propodeal apophysis). J. Tergite I, dorsal view, paratype, ♀ (NHMUK010636382). Scale bars: A = 0.5 mm; B–G = 0.2 mm; H–J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047695" httpUri="https://zenodo.org/record/5047695/files/figure.png" pageId="2" pageNumber="82">5I</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3DD6BFDD17D6EFD0D" box="[1367,1393,623,649]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">8I</figureCitation>
|
||
, arrows) .............................................................................................................................................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D82E20FFD3FFA3D881FD737D68FC13" pageId="2" pageNumber="82">
|
||
<paragraph id="8B933685FFD3FFA3D881FD737D68FCA9" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D881FD737D68FCA9" pageId="2" pageNumber="82">
|
||
2. Area externa of propodeum confluent with area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3DBF0FD727C30FD63" box="[972,1071,716,743]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1A</figureCitation>
|
||
). Juxtacoxal carina absent (
|
||
<figureCitation id="13172A00FFD3FFA3D8CFFD4E7950FC8E" box="[243,335,752,778]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="2" pageNumber="82">Fig. 2C</figureCitation>
|
||
). First and second flagellomeres about 5.6 and 6.5 × as long as wide respectively. Hind coxa tinged with white (
|
||
<figureCitation id="13172A00FFD3FFA3D9F4FCAD7A3BFCA9" box="[456,548,787,813]" captionStart="Fig" captionStartId="5.[189,232,1891,1917]" captionTargetBox="[189,1398,557,1849]" captionTargetId="figure-66@5.[189,1398,557,1853]" captionTargetPageId="5" captionText="Fig. 2. Morphology of Handaoia cuscoensis Bordera sp. nov., holotype, ♀ (AEIC). A. Habitus, lateral view. B. Head, front view. C. Metapleuron. D. Head and mesosoma, lateral view. E. Head and mesoscutum, dorsal view. F. Propodeum (arrow: propodeal apophysis). G. Metasoma, dorsal view. Scale bars: A = 1 mm; B, D–E, G = 0.2 mm; C, F = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047687" httpUri="https://zenodo.org/record/5047687/files/figure.png" pageId="2" pageNumber="82">Fig. 2A</figureCitation>
|
||
) ..............................................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DC3BFCAD7D0AFCA9" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[1031,1301,787,813]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="cuscoensis" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DC3BFCAD7CB0FCA9" bold="true" box="[1031,1199,787,813]" italics="true" pageId="2" pageNumber="82">H. cuscoensis</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD20FCAD7D68FCA9" box="[1308,1399,787,813]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD3FFA3D8FDFC897D68FC13" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D8FDFC897D68FC13" pageId="2" pageNumber="82">
|
||
– Area externa of propodeum separated from area dentipara by strong carina (
|
||
<figureCitation id="13172A00FFD3FFA3DC5AFC887CFBFCD5" box="[1126,1252,822,849]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1C, E</figureCitation>
|
||
). Juxtacoxal carina present, usually strong (
|
||
<figureCitation id="13172A00FFD3FFA3DA6CFCE77AB4FCF7" box="[592,683,857,883]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">Figs 4F</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3DA8AFCE77AC1FCF7" box="[694,734,857,883]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">7G</figureCitation>
|
||
). First and second flagellomeres about 4–5 and 3.2–5.4 × as long as wide, respectively. Hind coxa entirely dark brown (
|
||
<figureCitation id="13172A00FFD3FFA3DB82FCC27C3FFC13" box="[958,1056,892,919]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3DC10FCC37C4DFC13" box="[1068,1106,893,919]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">7A</figureCitation>
|
||
) .................................. 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D82E20FFD3FFA3D881FC007D68FABC" pageId="2" pageNumber="82">
|
||
<paragraph id="8B933685FFD3FFA3D881FC007D68FB0C" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D881FC007D68FB0C" pageId="2" pageNumber="82">
|
||
3. Fore wing with well-defined transverse dark brown band (
|
||
<figureCitation id="13172A00FFD3FFA3DB80FC007C00FC5C" box="[956,1055,958,984]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">Fig. 4A</figureCitation>
|
||
). Inner side of area externa 4.7 × as long as inner side of area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3DB3BFC5F7B7AFC7F" box="[775,869,993,1019]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1C</figureCitation>
|
||
). Juxtacoxal carina strong, irregular, joining submetapleural carina at its mid-length (
|
||
<figureCitation id="13172A00FFD3FFA3DAE4FBBA7B29FB9A" box="[728,822,1028,1054]" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">Fig. 4F</figureCitation>
|
||
). Tergite I about 1.8× as long as its maximum width. First and second flagellomeres about 5 and 5.4× as long as wide respectively. Flagellum dark brown (except white band), with first and second flagellomeres light brown (
|
||
<figureCitation id="13172A00FFD3FFA3DD39FBF578E1FB0C" captionStart="Fig" captionStartId="9.[189,232,1891,1917]" captionTargetBox="[189,1398,390,1851]" captionTargetId="figure-16@9.[189,1398,390,1853]" captionTargetPageId="9" captionText="Fig. 4. Morphology of Handaoia mercedensis Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Metasoma, dorsal view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A = 1 mm; B–E = 0.2 mm; F–G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047693" httpUri="https://zenodo.org/record/5047693/files/figure.png" pageId="2" pageNumber="82">Fig. 4A– B</figureCitation>
|
||
) ....................................................................................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DBCFFBD07D0AFB0C" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[1011,1301,1133,1160]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="mercedensis" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DBCFFBD07CB0FB0C" bold="true" box="[1011,1199,1133,1160]" italics="true" pageId="2" pageNumber="82">H. mercedensis</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD20FBD07D68FB0C" box="[1308,1399,1134,1160]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD3FFA3D8FDFB2C7D68FABC" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D8FDFB2C7D68FABC" pageId="2" pageNumber="82">
|
||
– Fore wing evenly infuscate (
|
||
<figureCitation id="13172A00FFD3FFA3DA02FB2F7A80FB2F" box="[574,671,1169,1195]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">Fig. 7A</figureCitation>
|
||
). Inner side of area externa 7.0× as long as inner side of area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3D956FB0A79D9FB4A" box="[362,454,1204,1230]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1E</figureCitation>
|
||
). Juxtacoxal carina weak but complete, joining anterior part of submetapleural carina (
|
||
<figureCitation id="13172A00FFD3FFA3D978FB6979BFFB76" box="[324,416,1239,1266]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">Fig. 7G</figureCitation>
|
||
). Tergite I about 2.4× as long as its maximum width. First and second flagellomeres 4.1 and 3.3× as long as wide respectively. Flagellum dark brown (except white band), with only first flagellomere light brown (
|
||
<figureCitation id="13172A00FFD3FFA3DA20FAA07A67FABC" box="[540,632,1310,1336]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,322,1851]" captionTargetId="figure-17@15.[189,1398,322,1855]" captionTargetPageId="15" captionText="Fig. 7. Morphology of Handaoia ruizcancinoi Bordera sp. nov., holotype, ♀ (CEUA). A. Habitus, lateral view. B. Head, front view. C. Head and mososoma, lateral view. D. Head and mososoma, dorsal view. E. Tergite I, dorsal view. F. Metasoma, dorsal view. G. Metapleuron (arrow: juxtacoxal carina). H. Propodeum (arrow: propodeal apophysis). Scale bars: A = 0.5 mm; B–F = 0.2 mm; G–H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047699" httpUri="https://zenodo.org/record/5047699/files/figure.png" pageId="2" pageNumber="82">Fig. 7A</figureCitation>
|
||
) ................................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DBCDFAA07D09FABC" authority="Bordera" authorityName="Bordera" authorityYear="2021" box="[1009,1302,1310,1336]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="ruizcancinoi" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DBCDFAA07CAFFABC" bold="true" box="[1009,1200,1310,1336]" italics="true" pageId="2" pageNumber="82">H. ruizcancinoi</emphasis>
|
||
Bordera
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD21FAA07D68FABC" box="[1309,1399,1310,1336]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D82E20FFD3FFA3D881FAE67D68F9A7" pageId="2" pageNumber="82">
|
||
<paragraph id="8B933685FFD3FFA3D881FAE67D68FA3D" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D881FAE67D68FA3D" pageId="2" pageNumber="82">
|
||
4. Area externa of propodeum confluent with area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3DBF0FAE67C30FAF6" box="[972,1071,1368,1394]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1D</figureCitation>
|
||
). Juxtacoxal carina absent (
|
||
<figureCitation id="13172A00FFD3FFA3D8CFFAC5794BFA12" box="[243,340,1403,1430]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[221,1366,267,1709]" captionTargetId="figure-11@12.[221,1366,267,1712]" captionTargetPageId="12" captionText="Fig. 5. Morphology of Handaoia plaumanni sp. nov. A. Habitus, lateral view, holotype, ♀ (NHMUK010636386). B. Head, dorsal view, paratype, ♀ (NHMUK010636395). C. Head, front view, paratype, ♀ (NHMUK010636382). D. Mesoscutum, dorsal view, paratype, ♀ (NHMUK010636395). E. Head and mesosoma, lateral view, paratype, ♀ (NHMUK010636395). F. Metasoma, lateral view, paratype, ♀ (NHMUK010636382). G. Metasoma, dorsal view, paratype, ♀ (NHMUK010636382). H. Metapleuron, paratype, ♀ (NHMUK010636395). I. Propodeum, paratype, ♀ (NHMUK010636382) (arrow: propodeal apophysis). J. Tergite I, dorsal view, paratype, ♀ (NHMUK010636382). Scale bars: A = 0.5 mm; B–G = 0.2 mm; H–J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047695" httpUri="https://zenodo.org/record/5047695/files/figure.png" pageId="2" pageNumber="82">Fig. 5H</figureCitation>
|
||
). Sternaulus very short and weak, reaching at most 0.3× the length of the mesopleuron (
|
||
<figureCitation id="13172A00FFD3FFA3D8CFFA207951FA3D" box="[243,334,1438,1465]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[221,1366,267,1709]" captionTargetId="figure-11@12.[221,1366,267,1712]" captionTargetPageId="12" captionText="Fig. 5. Morphology of Handaoia plaumanni sp. nov. A. Habitus, lateral view, holotype, ♀ (NHMUK010636386). B. Head, dorsal view, paratype, ♀ (NHMUK010636395). C. Head, front view, paratype, ♀ (NHMUK010636382). D. Mesoscutum, dorsal view, paratype, ♀ (NHMUK010636395). E. Head and mesosoma, lateral view, paratype, ♀ (NHMUK010636395). F. Metasoma, lateral view, paratype, ♀ (NHMUK010636382). G. Metasoma, dorsal view, paratype, ♀ (NHMUK010636382). H. Metapleuron, paratype, ♀ (NHMUK010636395). I. Propodeum, paratype, ♀ (NHMUK010636382) (arrow: propodeal apophysis). J. Tergite I, dorsal view, paratype, ♀ (NHMUK010636382). Scale bars: A = 0.5 mm; B–G = 0.2 mm; H–J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047695" httpUri="https://zenodo.org/record/5047695/files/figure.png" pageId="2" pageNumber="82">Fig. 5E</figureCitation>
|
||
). Head coarsely granulate (
|
||
<figureCitation id="13172A00FFD3FFA3DAB2FA207B14FA3D" box="[654,779,1438,1465]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[221,1366,267,1709]" captionTargetId="figure-11@12.[221,1366,267,1712]" captionTargetPageId="12" captionText="Fig. 5. Morphology of Handaoia plaumanni sp. nov. A. Habitus, lateral view, holotype, ♀ (NHMUK010636386). B. Head, dorsal view, paratype, ♀ (NHMUK010636395). C. Head, front view, paratype, ♀ (NHMUK010636382). D. Mesoscutum, dorsal view, paratype, ♀ (NHMUK010636395). E. Head and mesosoma, lateral view, paratype, ♀ (NHMUK010636395). F. Metasoma, lateral view, paratype, ♀ (NHMUK010636382). G. Metasoma, dorsal view, paratype, ♀ (NHMUK010636382). H. Metapleuron, paratype, ♀ (NHMUK010636395). I. Propodeum, paratype, ♀ (NHMUK010636382) (arrow: propodeal apophysis). J. Tergite I, dorsal view, paratype, ♀ (NHMUK010636382). Scale bars: A = 0.5 mm; B–G = 0.2 mm; H–J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047695" httpUri="https://zenodo.org/record/5047695/files/figure.png" pageId="2" pageNumber="82">Fig. 5B–C</figureCitation>
|
||
) .............................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DC55FA207D0BFA3D" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[1129,1300,1438,1465]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="plaumanni" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DC55FA207D0BFA3D" bold="true" box="[1129,1300,1438,1465]" italics="true" pageId="2" pageNumber="82">H. plaumanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD20FA217D68FA3D" box="[1308,1399,1439,1465]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD3FFA3D8FDFA7C7D68F9A7" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D8FDFA7C7D68F9A7" pageId="2" pageNumber="82">
|
||
– Area externa of propodeum separated from area dentipara by a strong carina (
|
||
<figureCitation id="13172A00FFD3FFA3DC50FA7F7CFAFA58" box="[1132,1253,1473,1500]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1B, F</figureCitation>
|
||
). Juxtacoxal carina present (
|
||
<figureCitation id="13172A00FFD3FFA3D99AFA5A7A18FA7B" box="[422,519,1508,1535]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3DA24FA5B7A21FA7B" box="[536,574,1509,1535]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">8H</figureCitation>
|
||
, arrows). Sternaulus deep, reaching at least 0.5 × the length of the mesopleuron (
|
||
<figureCitation id="13172A00FFD3FFA3D9ADF9B679ECF9A6" box="[401,499,1544,1570]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="13172A00FFD3FFA3D9C3F9B67A38F9A6" box="[511,551,1544,1570]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">8G</figureCitation>
|
||
). Head finely granulate .......................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D82E20FFD3FFA3D881F9FF7D68F85B" pageId="2" pageNumber="82">
|
||
<paragraph id="8B933685FFD3FFA3D881F9FF7D68F8AB" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D881F9FF7D68F8AB" pageId="2" pageNumber="82">
|
||
5. Inner side of area externa 3.0–3.4 × as long as inner side of area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3DC50F9FC7CD7F9D8" box="[1132,1224,1602,1628]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1B</figureCitation>
|
||
); inner side of area dentipara (section of the lateromedian longitudinal carina) 0.4–0.5 × the length of the outer side (section of the lateral longitudinal carina) (
|
||
<figureCitation id="13172A00FFD3FFA3DB1DF9367B62F926" box="[801,893,1672,1698]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1B</figureCitation>
|
||
). Lateromedian longitudinal carina of first metasomal tergite weak, reaching at most 0.6× the length of the tergite, or sometimes inconspicuous (
|
||
<figureCitation id="13172A00FFD3FFA3D8CFF9707950F96D" box="[243,335,1742,1769]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1B</figureCitation>
|
||
). Malar space 1.15–1.25× as long as basal mandibular width (
|
||
<figureCitation id="13172A00FFD3FFA3DC0BF9707C8BF96D" box="[1079,1172,1742,1769]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Fig. 3E</figureCitation>
|
||
). Tergite II smooth and shiny (
|
||
<figureCitation id="13172A00FFD3FFA3D950F94C79D8F888" box="[364,455,1778,1804]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Fig. 3B</figureCitation>
|
||
). Clypeus 2.1–2.2 × as wide as high (
|
||
<figureCitation id="13172A00FFD3FFA3DB40F94C7BCAF888" box="[892,981,1778,1804]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,333,1851]" captionTargetId="figure-16@7.[189,1398,317,1854]" captionTargetPageId="7" captionText="Fig. 3. Morphology of Handaoia fritzi sp. nov., holotype, ♀ (NHMUK010636396). A. Habitus, lateral view. B. Habitus, dorsal view. C. Head and mesoscutum, dorsal view. D. Head and mesosoma, lateral view. E. Head, front view. F. Metapleuron (arrow: juxtacoxal carina). G. Propodeum (arrow: propodeal apophysis). Scale bars: A–D = 0.5 mm; E–F = 0.2 mm; G = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047691" httpUri="https://zenodo.org/record/5047691/files/figure.png" pageId="2" pageNumber="82">Fig. 3E</figureCitation>
|
||
) ..................................................... ...................................................................................................................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DC89F8AB7D0AF8AA" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[1205,1301,1812,1839]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DC89F8AB7D0AF8AA" bold="true" box="[1205,1301,1812,1839]" italics="true" pageId="2" pageNumber="82">H. fritzi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD20F8AB7D68F8AB" box="[1308,1399,1813,1839]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B933685FFD3FFA3D8FDF8877D68F85B" blockId="2.[189,1399,553,2015]" pageId="2" pageNumber="82">
|
||
<keyLead id="30DD95B0FFD3FFA3D8FDF8877D68F85B" pageId="2" pageNumber="82">
|
||
– Inner side of area externa about 8.0× as long as inner side of area dentipara (
|
||
<figureCitation id="13172A00FFD3FFA3DCB2F8867CF6F8D6" box="[1166,1257,1848,1874]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1F</figureCitation>
|
||
); inner side of area dentipara 0.25 × the length of the outer side (
|
||
<figureCitation id="13172A00FFD3FFA3DB4FF8E57BD0F8F1" box="[883,975,1883,1909]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1F</figureCitation>
|
||
). Lateromedian longitudinal carina of tergite I strong, reaching 0.8 × the length of the tergite (
|
||
<figureCitation id="13172A00FFD3FFA3DBF8F8C07C3FF81D" box="[964,1056,1918,1945]" captionStart="Fig" captionStartId="4.[189,232,1891,1917]" captionTargetBox="[189,1398,400,1853]" captionTargetId="figure-46@4.[189,1398,400,1853]" captionTargetPageId="4" captionText="Fig. 1. Propodeum and tergite I of Handaoia spp., ♀♀. A. H. cuscoensis Bordera sp. nov., holotype (AEIC).B. H. fritzi sp. nov., holotype (NHMUK).C. H. mercedensisBordera sp. nov., holotype (CEUA). D. H. plaumanni sp. nov., holotype (NHMUK). E. H. ruizcancinoi Bordera sp. nov., holotype (CEUA). F. H. urceus sp. nov., holotype (NHMUK). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5047685" httpUri="https://zenodo.org/record/5047685/files/figure.png" pageId="2" pageNumber="82">Fig. 1F</figureCitation>
|
||
). Malar space 1.4 × as long as mandible base (
|
||
<figureCitation id="13172A00FFD3FFA3D9F1F81C7A32F838" box="[461,557,1954,1980]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">Fig. 8B</figureCitation>
|
||
). Tergite II finely granulate (
|
||
<figureCitation id="13172A00FFD3FFA3DBB3F81C7BEEF838" box="[911,1009,1954,1980]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">Fig. 8D</figureCitation>
|
||
). Clypeus 1.6× as wide as high (
|
||
<figureCitation id="13172A00FFD3FFA3D8CFF87B7952F85B" box="[243,333,1989,2015]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[189,1398,271,1816]" captionTargetId="figure-12@18.[189,1398,271,1818]" captionTargetPageId="18" captionText="Fig. 8. Morphology of Handaoia urceus sp. nov., holotype, ♀ (NMHUK). A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Metasoma, dorsal view. E. Head and mesosoma, dorsal view. F. Metasoma, lateral view. G. Head and mesosoma, dorsolateral view. H. Metapleuron (arrow: juxtacoxal carina). I. Propodeum (arrow: propodeal apophysis). Scale bars: A, D–G = 0.5 mm; B–C, H–I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5047701" httpUri="https://zenodo.org/record/5047701/files/figure.png" pageId="2" pageNumber="82">Fig. 8B</figureCitation>
|
||
) .................................................................................................................
|
||
<taxonomicName id="4C2C4D06FFD3FFA3DCA0F87B7D0BF85B" authority="Bordera & Broad, 2021" authorityName="Bordera & Broad" authorityYear="2021" box="[1180,1300,1989,2015]" class="Insecta" family="Ichneumonidae" genus="Handaoia" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="82" phylum="Arthropoda" rank="species" species="urceus" status="sp. nov.">
|
||
<emphasis id="B958EA97FFD3FFA3DCA0F87B7D0BF85B" bold="true" box="[1180,1300,1989,2015]" italics="true" pageId="2" pageNumber="82">H. urceus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26B57ECFFD3FFA3DD20F87B7D68F85B" box="[1308,1399,1989,2015]" pageId="2" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |