226 lines
16 KiB
XML
226 lines
16 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6213086" ID-GBIF-Dataset="e04117e5-9306-489d-a0a3-9dbd97774f74" ID-GBIF-Taxon="119613332" ID-ISSN="1175-5326" ID-Zenodo-Dep="6213086" checkinTime="1461057042134" checkinUser="plazi" docAuthor="Hong, Yong & James, Samuel W." docDate="2009" docId="03858791FFE1FFFFFF67FD65FD71F3A1" docLanguage="en" docName="zt02059p045.pdf" docOrigin="Zootaxa 2059" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@46f0bf0d" docStyleName="zootaxa.2007.monograph" docTitle="Pheretima gorasi Hong & James, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FFBCFFE9FFEBFFF5FFF0FF8AFFEBF41B" masterDocTitle="New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines" masterLastPageNumber="45" masterPageNumber="33" pageId="10" pageNumber="43" updateTime="1645486389336" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New earthworms of the Pheretima urceolata species group (Oligochaeta: Megascolecidae) from southern Luzon, Philippines</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hong, Yong</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>James, Samuel W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="volume">
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<mods:number>2059</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.186796</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e04117e5-9306-489d-a0a3-9dbd97774f74</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">186796</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6213086" ID-GBIF-Taxon="119613332" ID-Zenodo-Dep="6213086" LSID="urn:lsid:plazi:treatment:03858791FFE1FFFFFF67FD65FD71F3A1" httpUri="http://treatment.plazi.org/id/03858791FFE1FFFFFF67FD65FD71F3A1" lastPageNumber="43" pageId="10" pageNumber="43">
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<subSubSection pageId="10" pageNumber="43" type="nomenclature">
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<paragraph blockId="10.[151,459,751,812]" box="[151,459,751,777]" pageId="10" pageNumber="43">
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<heading bold="true" box="[151,459,751,777]" fontSize="11" level="1" pageId="10" pageNumber="43" reason="1">
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<emphasis bold="true" box="[151,459,751,777]" pageId="10" pageNumber="43">
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<taxonomicName box="[151,359,751,777]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="43" phylum="Annelida" rank="species" species="gorasi" status="sp. nov.">
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<emphasis bold="true" box="[151,359,751,777]" italics="true" pageId="10" pageNumber="43">Pheretima gorasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[366,459,751,777]" pageId="10" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="10.[151,459,751,812]" box="[151,294,786,812]" pageId="10" pageNumber="43">
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(
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<figureCitation box="[159,227,786,812]" captionStart="FIGURE 3" captionStartId="9.[151,255,1706,1730]" captionTargetBox="[243,1321,208,1666]" captionTargetId="figure@9.[237,1329,194,1682]" captionTargetPageId="9" captionText="FIGURE 3. Pheretima buhiensis sp. nov. A: ventral view; B: spermatheca. Pheretima gorasi sp. nov. C: ventral view; D: spermatheca. Scale bars = 2.5 mm (ventral views), 2 mm (spermathecae)." httpUri="https://zenodo.org/record/186799/files/figure.png" pageId="10" pageNumber="43">
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Fig.
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<number box="[214,227,786,812]" pageId="10" pageNumber="43" value="3.0">3</number>
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</figureCitation>
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C–D)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="materials_examined">
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[151,267,866,892]" pageId="10" pageNumber="43">Material.</emphasis>
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<typeStatus box="[278,392,866,892]" pageId="10" pageNumber="43">Holotype</typeStatus>
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: One clitellate (NMA 004321):
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<collectingCountry box="[787,921,866,892]" name="Philippines" pageId="10" pageNumber="43">Philippines</collectingCountry>
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, Camarines Sur, Mt. Isarog (
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<geoCoordinate box="[1279,1431,866,892]" direction="north" orientation="latitude" pageId="10" pageNumber="43" precision="9" value="13.661667">
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<number box="[1279,1309,866,892]" pageId="10" pageNumber="43" value="13.0">13</number>
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°
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<number box="[1332,1404,866,892]" pageId="10" pageNumber="43" value="39.7">39.70</number>
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’N
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</geoCoordinate>
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,
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<geoCoordinate box="[151,311,906,932]" direction="east" orientation="longitude" pageId="10" pageNumber="43" precision="9" value="123.367836">
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<number box="[151,195,906,932]" pageId="10" pageNumber="43" value="123.0">123</number>
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°
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<number box="[214,287,906,932]" pageId="10" pageNumber="43" value="22.07">22.07</number>
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’E
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</geoCoordinate>
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),
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<quantity box="[332,423,906,932]" metricMagnitude="3" metricUnit="m" metricValue="1.7449999999999999" pageId="10" pageNumber="43" unit="m" value="1745.0">1745 m</quantity>
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, soil,
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<date box="[491,650,906,932]" pageId="10" pageNumber="43" value="2001-05-16">
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<number box="[491,520,906,932]" pageId="10" pageNumber="43" value="16.0">16</number>
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May
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<number box="[589,650,906,932]" pageId="10" pageNumber="43" value="2001.0">2001</number>
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</date>
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, S. W. James coll.
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<number box="[869,884,906,932]" pageId="10" pageNumber="43" value="2.0">2</number>
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<typeStatus box="[891,1008,906,932]" pageId="10" pageNumber="43">paratypes</typeStatus>
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:
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<number box="[1018,1032,906,932]" pageId="10" pageNumber="43" value="1.0">1</number>
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clitellate (NMA 004326),
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<number box="[1345,1360,906,932]" pageId="10" pageNumber="43" value="1.0">1</number>
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clitellate (
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<collectionCode box="[208,321,946,972]" pageId="10" pageNumber="43">KUNHM</collectionCode>
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003015): Same data as for
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<typeStatus box="[643,749,946,972]" pageId="10" pageNumber="43">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[197,1045,986,1012]" pageId="10" pageNumber="43" type="etymology">
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<paragraph blockId="10.[151,1437,866,1978]" box="[197,1045,986,1012]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[197,338,986,1012]" pageId="10" pageNumber="43">Etymology.</emphasis>
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The species name is an anagram of Isarog, the
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<typeStatus box="[894,945,986,1012]" pageId="10" pageNumber="43">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="diagnosis">
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[197,325,1026,1052]" pageId="10" pageNumber="43">Diagnosis.</emphasis>
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<taxonomicName box="[333,444,1026,1051]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="43" phylum="Annelida" rank="species" species="urceolata">
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<emphasis box="[333,444,1026,1051]" italics="true" pageId="10" pageNumber="43">urceolata</emphasis>
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</taxonomicName>
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-group without setal gaps, closely-spaced genital pores, septum
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<number box="[1197,1210,1026,1052]" pageId="10" pageNumber="43" value="8.0">8</number>
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/
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<number box="[1220,1234,1026,1052]" pageId="10" pageNumber="43" value="9.0">9</number>
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present, and two bursal pads.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="description">
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[197,349,1106,1132]" pageId="10" pageNumber="43">Description.</emphasis>
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Brown dorsal pigment darkest at head and tail. Dimensions
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<quantity box="[1062,1218,1106,1132]" metricMagnitude="-1" metricUnit="m" metricValue="1.66" metricValueMax="1.77" metricValueMin="1.55" pageId="10" pageNumber="43" unit="mm" value="166.0" valueMax="177.0" valueMin="155.0">155–177 mm</quantity>
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by
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<number box="[1263,1299,1106,1132]" pageId="10" pageNumber="43" value="6.7">6.7</number>
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–7.0 at segment X, 7.0 mm at XXX,
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<number box="[467,503,1146,1172]" pageId="10" pageNumber="43" value="6.2">6.2</number>
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–7.0 mm at clitellum, segments
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<number box="[888,992,1146,1172]" pageId="10" pageNumber="43" value="116.5" valueMax="118.0" valueMin="115.0">115–118</number>
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; body circular in cross-section. Setae numbering
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<number box="[286,359,1186,1212]" pageId="10" pageNumber="43" value="28.0" valueMax="32.0" valueMin="24.0">24–32</number>
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at VII,
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<number box="[453,526,1186,1212]" pageId="10" pageNumber="43" value="33.0" valueMax="41.0" valueMin="25.0">25–41</number>
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at XX; 0 between male pores, size uniform within segments, but setae II–IX larger; setal formula AA:AB:YZ:ZZ=
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<number box="[601,618,1226,1252]" pageId="10" pageNumber="43" value="2.0">2</number>
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:
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<number box="[623,640,1226,1252]" pageId="10" pageNumber="43" value="2.0">2</number>
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:
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<number box="[644,662,1226,1252]" pageId="10" pageNumber="43" value="4.0">4</number>
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:
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<number box="[667,685,1226,1252]" pageId="10" pageNumber="43" value="4.0">4</number>
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at XIII. Clitellum annular XIV–XVI, setae invisible externally.
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</paragraph>
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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First dorsal pores
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<number box="[409,440,1269,1295]" pageId="10" pageNumber="43" value="12.0">12</number>
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/
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<number box="[450,479,1269,1295]" pageId="10" pageNumber="43" value="13.0">13</number>
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. One pair of spermathecal pores in
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<number box="[897,910,1269,1295]" pageId="10" pageNumber="43" value="5.0">5</number>
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/
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<number box="[920,934,1269,1295]" pageId="10" pageNumber="43" value="6.0">6</number>
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between
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<number box="[1047,1062,1269,1295]" pageId="10" pageNumber="43" value="2.0">2</number>
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nd–
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<number box="[1095,1108,1269,1295]" pageId="10" pageNumber="43" value="3.0">3</number>
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rd setal lines, grooved protuberance on anterior side of each pore below level of pore lips,
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<number box="[912,1030,1309,1335]" pageId="10" pageNumber="43" value="0.065" valueMax="0.07" valueMin="0.06">0.06–0.07</number>
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circumference apart,
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<quantity box="[1292,1436,1309,1335]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="10" pageNumber="43" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
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between spermathecal pores;
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<quantity box="[498,640,1352,1378]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="10" pageNumber="43" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
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openings of copulatory bursa paired in XVIII at
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<number box="[1218,1232,1352,1378]" pageId="10" pageNumber="43" value="6.0">6</number>
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th setal lines,
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<number box="[1385,1436,1352,1378]" pageId="10" pageNumber="43" value="0.1">0.10</number>
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circumference apart, distance between openings 2.0–
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<quantity box="[773,865,1392,1418]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="43" unit="mm" value="2.3">2.3 mm</quantity>
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. Genital markings lacking.
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</paragraph>
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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<collectingCountry box="[197,262,1432,1458]" name="Spain" pageId="10" pageNumber="43">Septa</collectingCountry>
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<number box="[273,287,1432,1458]" pageId="10" pageNumber="43" value="5.0">5</number>
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/6, 6/
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<number box="[351,365,1432,1458]" pageId="10" pageNumber="43" value="7.0">7</number>
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thick,
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<number box="[452,466,1432,1458]" pageId="10" pageNumber="43" value="7.0">7</number>
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/
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<number box="[476,489,1432,1458]" pageId="10" pageNumber="43" value="8.0">8</number>
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thin,
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<number box="[563,576,1432,1458]" pageId="10" pageNumber="43" value="8.0">8</number>
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/
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<number box="[586,600,1432,1458]" pageId="10" pageNumber="43" value="9.0">9</number>
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very thin,
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<number box="[736,750,1432,1458]" pageId="10" pageNumber="43" value="9.0">9</number>
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/
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<number box="[760,788,1432,1458]" pageId="10" pageNumber="43" value="10.0">10</number>
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absent,
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<date box="[891,1038,1432,1458]" pageId="10" pageNumber="43" value="1914-10-11" valueMax="1914-10-13" valueMin="1914-10-11">
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<number box="[891,920,1432,1458]" pageId="10" pageNumber="43" value="10.0">10</number>
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/
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<number box="[929,999,1432,1458]" pageId="10" pageNumber="43" value="12.0" valueMax="13.0" valueMin="11.0">11–13</number>
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/
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<number box="[1009,1038,1432,1458]" pageId="10" pageNumber="43" value="14.0">14</number>
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</date>
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thinly muscled. Gizzard in VIII, intestine begins in XVI, small paired lymph glands from XXVII along dorsal vessel; intestinal caeca simple originating in XXVII, extending anteriorly about to XXIII, a finger-shaped sac; typhlosole medium from XXVI,
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<number box="[238,267,1552,1578]" pageId="10" pageNumber="43" value="38.0">38</number>
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intestinal wall blood vessels. Hearts in XI–XIII esophageal; X absent; IX left side only, VIII to gizzard, VII to ventral vessel.
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</paragraph>
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">Ovaries and funnels in XIII, spermathecae one pair in VI with nephridia on ducts, spermathecae smooth heart-shaped ampulla chamber; duct much shorter than ampulla, diverticulum chamber match-shaped, iridescent, with stalk shorter than ampulla. Male sexual system holandric, testes and funnels in ventrally paired sacs in X–XI. Seminal vesicles two pairs in XI–XII with long apical dorsal lobes, prostates in XVIII–XIX, with stout muscular ducts, entering anterior half of bilobate copulatory bursae without stalked glands; penis oval protrusion from the roof, lateral end free greater height posterior section of copulatory bursa mainly glandular, with very large folded pad extending from roof to floor, small pad anterior, medial to penis on roof.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="discussion">
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<paragraph blockId="10.[151,1437,866,1978]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[197,318,1912,1938]" pageId="10" pageNumber="43">Remarks.</emphasis>
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<taxonomicName box="[329,535,1912,1937]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="43" phylum="Annelida" rank="species" species="gorasi" status="sp. nov.">
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<emphasis box="[329,535,1912,1937]" italics="true" pageId="10" pageNumber="43">Pheretima gorasi</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[545,641,1912,1938]" pageId="10" pageNumber="43">
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<taxonomicNameLabel box="[545,641,1912,1938]" pageId="10" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
is distinguished from its species group members according to the remarks under
|
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<taxonomicName box="[326,566,1952,1977]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="43" phylum="Annelida" rank="species" species="buhiensis" status="sp. nov.">
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<emphasis box="[326,566,1952,1977]" italics="true" pageId="10" pageNumber="43">Pheretima buhiensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[573,666,1952,1978]" pageId="10" pageNumber="43">
|
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<taxonomicNameLabel box="[573,666,1952,1978]" pageId="10" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |