222 lines
21 KiB
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222 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.341.5816" ID-GBIF-Dataset="1485e168-157a-47d4-b9fa-2a21400a28ff" ID-PMC="PMC3800810" ID-Pensoft-Pub="1313-2970-341-77" ID-PubMed="24146599" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-341-77" ModsDocOrigin="ZooKeys 341" ModsDocTitle="Sea anemones (Cnidaria, Anthozoa, Actiniaria) from coral reefs in the southern Gulf of Mexico" checkinTime="1451246812539" checkinUser="pensoft" docAuthor="Gonzalez-Munoz, Ricardo, Simoes, Nuno, Tello-Musi, Jose Luis & Rodriguez, Estefania" docDate="2013" docId="317908D7142B9A1F1ED4ED77A136CF78" docLanguage="en" docName="ZooKeys 341: 77-106" docOrigin="ZooKeys 341" docSource="http://dx.doi.org/10.3897/zookeys.341.5816" docTitle="Isoaulactinia stelloides McMurrich 1889" docType="treatment" docVersion="3" lastPageNumber="85" masterDocId="FFEBA2745B17421B007FDB25C72D5A0A" masterDocTitle="Sea anemones (Cnidaria, Anthozoa, Actiniaria) from coral reefs in the southern Gulf of Mexico" masterLastPageNumber="106" masterPageNumber="77" pageNumber="84" updateTime="1668156630225" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Sea anemones (Cnidaria, Anthozoa, Actiniaria) from coral reefs in the southern Gulf of Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gonzalez-Munoz, Ricardo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Simoes, Nuno</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tello-Musi, Jose Luis</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodriguez, Estefania</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>341</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>106</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.341.5816</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.341.5816</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-341-77</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152048732" LSID="urn:lsid:plazi:treatment:317908D7142B9A1F1ED4ED77A136CF78" httpUri="http://treatment.plazi.org/id/317908D7142B9A1F1ED4ED77A136CF78" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
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<subSubSection pageId="7" pageNumber="84" type="nomenclature">
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName LSID="http://species-id.net/wiki/Isoaulactinia_stelloides" authority="McMurrich, 1889" authorityName="McMurrich" authorityYear="1889" class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides (McMurrich, 1889)</taxonomicName>
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Figure 6, Table 2
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="84" type="reference_group">
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Aulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aulactinia stelloides" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stelloides">Aulactinia stelloides</taxonomicName>
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<bibRefCitation author="McMurrich, JP" journalOrPublisher="W. I. Journal of Morphology" pageId="17" pageNumber="94" pagination="1 - 80" title="The Actiniaria of the Bahama Islands." url="10.1002/jmor.1050030102" volume="3" year="1889">McMurrich 1889</bibRefCitation>
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: 28-31.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Aulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aulactinia stella" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stella">Aulactinia stella</taxonomicName>
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:
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<bibRefCitation author="Duerden, JE" journalOrPublisher="Annals and Magazine of Natural History" pageId="16" pageNumber="93" pagination="1 - 15" title="The actiniarian family Aliciidae." url="10.1080/00222939708680594" volume="20" year="1897">Duerden 1897</bibRefCitation>
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: 454-455.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Bunodella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bunodella stelloides" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stelloides">Bunodella stelloides</taxonomicName>
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:
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<bibRefCitation author="Verrill, AE" journalOrPublisher="American Journal of Science and Arts" pageId="18" pageNumber="95" pagination="41 - 50" title="Descriptions of imperfectly known and new actinians, with critical notes on other species, II." url="10.2475/ajs.s4-7.37.41" volume="7" year="1899">Verrill 1899</bibRefCitation>
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: 43-44.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Bunodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bunodes stella" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stella">Bunodes stella</taxonomicName>
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:
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<bibRefCitation author="Duerden, JE" journalOrPublisher="Journal of the Institute of Jamaica" pageId="16" pageNumber="93" pagination="449 - 465" title="The Actiniaria around Jamaica." volume="2" year="1898">Duerden 1898</bibRefCitation>
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: 455.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Bunodactis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bunodactis stelloides" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stelloides">Bunodactis stelloides</taxonomicName>
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:
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<bibRefCitation author="Verrill, AE" journalOrPublisher="Transactions of the Connecticut Academy of Arts and Sciences" pageId="18" pageNumber="95" pagination="551 - 572" title="Additions to the Anthozoa and Hydrozoa of the Bermudas. Anthozoa." volume="10" year="1900">Verrill 1900</bibRefCitation>
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: 556.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Anthopleura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthopleura catenulata" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="catenulata">Anthopleura catenulata</taxonomicName>
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:
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<bibRefCitation pageId="7" pageNumber="84">Cairns, den Hartog and Arneson 1986</bibRefCitation>
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: 177-178; Pl. 51.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Anthopleura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthopleura carneola" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="carneola">Anthopleura carneola</taxonomicName>
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:
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<bibRefCitation pageId="7" pageNumber="84">Cairns, den Hartog and Arneson 1986</bibRefCitation>
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: 177-178; Pl. 51.
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</paragraph>
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<paragraph pageId="7" pageNumber="84">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="7" pageNumber="84" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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:
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<bibRefCitation pageId="7" pageNumber="84">
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<normalizedToken originalValue="Belém">Belem</normalizedToken>
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, Herrera-Moreno and Schlenz 1996
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</bibRefCitation>
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: 77-88.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="84" type="material examined">
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<paragraph pageId="7" pageNumber="84">Material examined.</paragraph>
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<paragraph pageId="7" pageNumber="84">
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La Gallega reef (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="19.222223">19°13'20"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-96.1275">96°07'39"W</geoCoordinate>
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; six specimens).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="85" type="diagnosis">
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<paragraph pageId="8" pageNumber="85">
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<pageBreakToken pageId="8" pageNumber="85" start="start">Diagnosis</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="8" pageNumber="85">
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Fully expanded oral disc and tentacles to 24-38 mm in diameter. Oral disc smooth, slightly wider than column, 9-11 mm in diameter, light- or olive-green, sometimes with small white stripes near tentacles bases (Figure 6A, B). Tentacles hexamerously arranged in four cycles (about 48 in number), simple, smooth, moderately long (9-14 mm in length), conical, tapering distally, inner ones longer than outer ones, contractile, olive-green with white bands along entire length (Figure 6A, B). Deep fosse (Figure 6G). Twenty-four endocoelic marginal projections (Figure 6C, G) with basitrichs and macrobasic p-mastigophores. Column cylindrical, 8-12 in diameter and 13-22 mm in height, with approximately 48 longitudinal rows of verrucae along entire column, but more conspicuous distally (Figure 6C). Pedal disc well-developed, 9-16 mm in diameter (Figure 6C). Column, verrucae, and pedal disc light-brown or beige (Figure 6C). Mesenteries hexamerously arranged in three cycles (24 pairs in specimens examined): all cycles perfect; same number of mesenteries distally and proximally. First and second cycles fertile (except directives); hermaphroditic (?), only oocytes observed in specimens examined (Figure 6E). Developing polyps in coelenteron (Figure 6F). Two pairs of directives each attached to a well-developed siphonoglyph (Figure 6D). Retractor muscles strong and restricted; parietobasilar muscles well-developed with relatively long and thick free mesogleal pennon (Figure 6E). Basilar muscles well-developed (Figure 6H). Marginal sphincter muscle endodermal, strong and circumscribed, palmate (Figure 6G). Longitudinal muscles of tentacles ectodermal (Figure 6I). Zooxanthellae present. Cnidom: basitrichs, microbasic b-mastigophores, macrobasic and microbasic p-mastigophores, and spirocysts (Figure 6
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<normalizedToken originalValue="J–Y">J-Y</normalizedToken>
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; see Table 2).
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</paragraph>
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<caption pageId="8" pageNumber="85">
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<paragraph pageId="8" pageNumber="85">
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Figure 6.
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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. A Live specimen in natural habitat B Oral view C Lateral view D Detail of directives showing a siphonoglyph E Cross section through proximal column F Detail of brooded juvenile G Longitudinal section through margin showing marginal sphincter muscle and marginal projection H Longitudinal section though base showing basilar muscles I Cross section through tentacle
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<normalizedToken originalValue="J–Y">J-Y</normalizedToken>
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Cnidae.- marginal projection: J small basitrich K macrobasic p-mastigophore; actinopharynx: M basitrich N macrobasic p-mastigophore O microbasic p-mastigophore P long, curved basitrich; column: Q small basitrich R macrobasic p-mastigophore; filament: S small basitrich T basitrich U macrobasic p-mastigophore V microbasic b-mastigophore W microbasic p-mastigophore; tentacle: X basitrich Y spirocyst. Abbreviations.- b: brooded juvenile, bm: basilar muscle, c: column, d: directives, e: epidermis, fo: fosse, lm: longitudinal muscle, m: mesoglea, mp: marginal projection, o: oocyst, od: oral disc, pd: pedal disc, pm: parietobasilar muscle, rm: retractor muscle, s: siphonoglyph, sp: sphincter, t: tentacle, vr: verrucae, z: zooxanthellae. Scale bars:
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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: 10 mm;
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<normalizedToken originalValue="D–I">D-I</normalizedToken>
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: 200
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<normalizedToken originalValue="μm">μm</normalizedToken>
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;
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<normalizedToken originalValue="J–Y">J-Y</normalizedToken>
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: 25
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<normalizedToken originalValue="μm">μm</normalizedToken>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="85" type="natural history">
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<paragraph pageId="8" pageNumber="85">Natural history.</paragraph>
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<paragraph pageId="8" pageNumber="85">
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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inhabits shallow waters in the lagoon reef zone, at 1-2 m, near
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Actinostella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Actinostella flosculifera" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="flosculifera">Actinostella flosculifera</taxonomicName>
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,
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<taxonomicName class="Anthozoa" family="Stichodactylidae" genus="Stichodactyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stichodactyla helianthus" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="helianthus">Stichodactyla helianthus</taxonomicName>
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, and the zoanthid
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<taxonomicName class="Anthozoa" family="Sphenopidae" genus="Palythoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palythoa caribaeorum" order="Zoantharia" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="caribaeorum">Palythoa caribaeorum</taxonomicName>
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(Duchassaing & Michelotti, 1860). It lives with the column burrowed in the sand but the pedal disc attached to rocks and coral rubble. Although we only observed developing oocytes in the two specimens histologically examined,
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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has been reported as a simultaneous hermaphroditic, internally brooding, often with developing polyps in the coelenteron (
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<bibRefCitation author="Belem, MJ" journalOrPublisher="Biociencias" pageId="15" pageNumber="92" pagination="77 - 88" title="On Isoaulactinia stelloides (McMurrich, 1889), n. gen., n. comb. (Cnidaria; Actiniaria; Actiniidae)." volume="4" year="1996">
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<normalizedToken originalValue="Belém">Belem</normalizedToken>
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et al. 1996
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</bibRefCitation>
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,
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<bibRefCitation author="Daly, M" journalOrPublisher="Bulletin of Marine Sciences" pageId="16" pageNumber="93" pagination="401 - 421" title="Taxonomy, circumscription, and usage in Anthopleura (Cnidaria: Anthozoa: Actiniaria) from the Gulf of Mexico and the Caribbean." volume="74" year="2004">Daly and den Hartog 2004</bibRefCitation>
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); the latter have also been observed in the present study (Figure 6F).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="85" type="distribution">
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<paragraph pageId="8" pageNumber="85">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="85">
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Western Atlantic, from Bermuda to Barbados, and along the Caribbean Sea (
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<bibRefCitation author="Belem, MJ" journalOrPublisher="Biociencias" pageId="15" pageNumber="92" pagination="77 - 88" title="On Isoaulactinia stelloides (McMurrich, 1889), n. gen., n. comb. (Cnidaria; Actiniaria; Actiniidae)." volume="4" year="1996">
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<normalizedToken originalValue="Belém">Belem</normalizedToken>
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et al. 1996
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</bibRefCitation>
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,
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<bibRefCitation author="Daly, M" journalOrPublisher="Bulletin of Marine Sciences" pageId="16" pageNumber="93" pagination="401 - 421" title="Taxonomy, circumscription, and usage in Anthopleura (Cnidaria: Anthozoa: Actiniaria) from the Gulf of Mexico and the Caribbean." volume="74" year="2004">Daly and den Hartog 2004</bibRefCitation>
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). This is the first record for the coast of Mexico; found in the VRS (see Table 1).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="85" type="remarks">
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<paragraph pageId="8" pageNumber="85">Remarks.</paragraph>
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<paragraph pageId="8" pageNumber="85">
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Currently
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="genus">Isoaulactinia</taxonomicName>
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has two valid species (
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<bibRefCitation author="Daly, M" journalOrPublisher="Pacific Science" pageId="16" pageNumber="93" pagination="377 - 390" title="Anatomy and taxonomy of three species of sea anemones (Cnidaria: Anthozoa: Actiniidae) from the Gulf of California, including Isoaulactinia hespervolita n. sp." url="10.1353/psc.2004.0030" volume="58" year="2004">Daly 2004</bibRefCitation>
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,
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<bibRefCitation author="Fautin, DG" journalOrPublisher="Texas University Press, College Station, Texas" pageId="16" pageNumber="93" title="Hexacorallians of the World." url="http://geoportal.kgs.ku.edu/hexacoral/anemone2/index.cfm" year="2013">Fautin 2013</bibRefCitation>
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):
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia hespervolita" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="hespervolita">Isoaulactinia hespervolita</taxonomicName>
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Daly, 2004, and
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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. According to
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<bibRefCitation author="Daly, M" journalOrPublisher="Pacific Science" pageId="16" pageNumber="93" pagination="377 - 390" title="Anatomy and taxonomy of three species of sea anemones (Cnidaria: Anthozoa: Actiniidae) from the Gulf of California, including Isoaulactinia hespervolita n. sp." url="10.1353/psc.2004.0030" volume="58" year="2004">Daly (2004)</bibRefCitation>
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia hespervolita" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="hespervolita">Isoaulactinia hespervolita</taxonomicName>
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differs from
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
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in having an unmarked oral disc and tentacles, spinose holotrichs in the column and being gonochoric rather than hermaphroditic. In addition,
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia hespervolita" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="hespervolita">Isoaulactinia hespervolita</taxonomicName>
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has a reddish-orange to greenish-brown column, oral disc and tentacles; approximately 80 tentacles arranged in up to five cycles, and macrobasic p-mastigophores only in the column and tentacles (
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<bibRefCitation author="Daly, M" journalOrPublisher="Pacific Science" pageId="16" pageNumber="93" pagination="377 - 390" title="Anatomy and taxonomy of three species of sea anemones (Cnidaria: Anthozoa: Actiniidae) from the Gulf of California, including Isoaulactinia hespervolita n. sp." url="10.1353/psc.2004.0030" volume="58" year="2004">Daly 2004</bibRefCitation>
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). We found additional microbasic b-mastigophores in the actinopharynx of
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<taxonomicName class="Anthozoa" family="Actiniidae" genus="Isoaulactinia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isoaulactinia stelloides" order="Actiniaria" pageId="8" pageNumber="85" phylum="Cnidaria" rank="species" species="stelloides">Isoaulactinia stelloides</taxonomicName>
|
||
but they were not abundant (Table 2). This category of nematocyst has not been previously reported in the actinopharynx of either of the species (
|
||
<bibRefCitation author="Belem, MJ" journalOrPublisher="Biociencias" pageId="15" pageNumber="92" pagination="77 - 88" title="On Isoaulactinia stelloides (McMurrich, 1889), n. gen., n. comb. (Cnidaria; Actiniaria; Actiniidae)." volume="4" year="1996">
|
||
<normalizedToken originalValue="Belém">Belem</normalizedToken>
|
||
et al. 1996
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Daly, M" journalOrPublisher="Bulletin of Marine Sciences" pageId="16" pageNumber="93" pagination="401 - 421" title="Taxonomy, circumscription, and usage in Anthopleura (Cnidaria: Anthozoa: Actiniaria) from the Gulf of Mexico and the Caribbean." volume="74" year="2004">Daly and den Hartog 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |