641 lines
87 KiB
XML
641 lines
87 KiB
XML
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<mods:title id="E59ADFC8A40F7EB0121BC7F4B3D05A7F">A revision of the Aphodiini genus Cnemargulus Semenov, 1903 (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<taxonomicName id="82F23DD6FFFCFF9CFF27FD87FD56547C" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[192,763,558,582]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="krulikovskyi">
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<emphasis id="77869A47FFFCFF9CFF27FD87FD86547C" bold="true" box="[192,555,558,582]" italics="true" pageId="5" pageNumber="4">Cnemargulus krulikovskyi</emphasis>
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<bibRefCitation id="21633BA4FFFCFF9CFDD5FD87FD56547C" author="Semenov, A. P." box="[562,763,558,582]" pageId="5" pageNumber="4" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
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<paragraph id="454D4655FFFCFF9CFF27FDE6FEE9545D" blockId="5.[192,763,558,615]" box="[192,324,591,615]" pageId="5" pageNumber="4">
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(
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<figureCitation id="DDC95AD0FFFCFF9CFF2EFDE6FE96545D" box="[201,315,591,615]" captionStart="Figures 7–13" captionStartId="10.[192,287,1762,1784]" captionTargetBox="[324,1278,218,1720]" captionTargetId="figure@10.[263,1370,205,1728]" captionTargetPageId="10" captionText="Figures 7–13. Cnemargulus krulikoƲskyi Semenov, 1903. 7) Habitus of male [lectotype]. 8) Habitus of female [paralectotype, Turkmenistan, Mary prov., Ýolöten]. 9) Habitus of male [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 10) Aedeagus, dorsal view [lectotype]. 11) Aedeagus, dorsal view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 12) Aedeagus, lateral view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 13) Epipharynx [lectotype]. Photos by Augusto Degiovanni, drawings by Ivo Gudenzi." figureDoi="http://doi.org/10.5281/zenodo.3670459" httpUri="https://zenodo.org/record/3670459/files/figure.png" pageId="5" pageNumber="4">Fig. 7–13</figureCitation>
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<emphasis id="77869A47FFFCFF9CFF27FDD7FE7254A9" box="[192,479,638,659]" italics="true" pageId="5" pageNumber="4">Cnemargulus krulikovskii</emphasis>
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<bibRefCitation id="21633BA4FFFCFF9CFE02FDD7FD1154AE" author="Semenov, A. P." box="[485,700,638,660]" pageId="5" pageNumber="4" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov 1903: 354</bibRefCitation>
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<bibRefCitation id="21633BA4FFFCFF9CFD21FDD7FC3C54A9" author="Schmidt, A." box="[710,913,638,660]" pageId="5" pageNumber="4" pagination="1 - 155" refId="ref6019" refString="Schmidt, A. 1910 a. Coleoptera Lamellicornia Aphodiinae. P. 1 - 155. In: Wytsman P.: Genera Insectorum. 110. Tervuren." type="book chapter" year="1910">Schmidt 1910a: 13</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFC5AFDD7FB8454A9" box="[957,1065,638,659]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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<bibRefCitation id="21633BA4FFFCFF9CFBDAFDD7FB5754A9" author="Schmidt, A." box="[1085,1274,638,660]" pageId="5" pageNumber="4" pagination="1 - 111" refId="ref6049" refString="Schmidt, A. 1910 b. Aphodiinae. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Berlin 20: 1 - 111." type="journal article" year="1910">Schmidt 1910b: 4</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFAC1FDD7FA3F54A9" box="[1318,1426,638,659]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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<emphasis id="77869A47FFFCFF9CFE5AFD32FD6D548A" box="[445,704,667,688]" italics="true" pageId="5" pageNumber="4">Cnemisus krulikovskyii</emphasis>
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);
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<treatmentCitation id="C4536044FFFCFF9CFD31FD32FC1C548B" author="Winkler, A." box="[726,945,667,689]" page="1059" pageId="5" pageNumber="4" year="1929">
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<bibRefCitation id="21633BA4FFFCFF9CFD31FD32FC1C548B" author="Winkler, A." box="[726,945,667,689]" pageId="5" pageNumber="4" refId="ref6213" refString="Winkler, A. 1929. Catalogus Coleopterorum Regionis Palaearcticae (1924 - 1932). Scarabaeidae. p. 1009 - 1136. A. Winkler; Wien 9. 1698 p." type="book" year="1929">Winkler 1929: 1059</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFC39FD32FBE7548A" box="[990,1098,667,688]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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);
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<treatmentCitation id="C4536044FFFCFF9CFBB8FD32FAED548B" author="Balthasar, V." box="[1119,1344,667,689]" page="474" pageId="5" pageNumber="4" year="1964">
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<bibRefCitation id="21633BA4FFFCFF9CFBB8FD32FAED548B" author="Balthasar, V." box="[1119,1344,667,689]" pageId="5" pageNumber="4" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar 1964: 474</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFA8AFD32FEAC54F4" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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(
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<emphasis id="77869A47FFFCFF9CFEF7FD11FE0854F7" box="[272,421,696,717]" italics="true" pageId="5" pageNumber="4">Cnemargulus</emphasis>
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)
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<emphasis id="77869A47FFFCFF9CFE53FD11FD9554F7" box="[436,568,696,717]" italics="true" pageId="5" pageNumber="4">krulikovskii</emphasis>
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<bibRefCitation id="21633BA4FFFCFF9CFDA8FD10FC7F54F4" author="Medvedev, S. I. & L. M. Nikritin" box="[591,978,697,719]" pageId="5" pageNumber="4" pagination="1881 - 1882" refId="ref5862" refString="Medvedev, S. I., and L. M. Nikritin. 1971. [Once more about the independence of the genus Cnemargulus Sem. (Coleoptera, Scarabaeidae) and its difference from the genus Cnemisus Motsch.] Zoologicheskiy Zhurnal 50: 1881 - 1882." type="journal article" year="1971">Medvedev and Nikritin 1971: 1882</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFBE7FD11FA8D54F7" box="[1024,1312,696,717]" italics="true" pageId="5" pageNumber="4">Cnemargulus krulikovskii</emphasis>
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);
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<treatmentCitation id="C4536044FFFCFF9CFAD0FD10FE8C54D1" author="Nikolajev, G. V." page="40" pageId="5" pageNumber="4" year="1979">
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<bibRefCitation id="21633BA4FFFCFF9CFAD0FD10FE8C54D1" author="Nikolajev, G. V." pageId="5" pageNumber="4" pagination="39 - 41" refId="ref5913" refString="Nikolajev, G. V. 1979. Novyi vid plastinchatousykh triby Aphodiini (Coleoptera, Scarabaeidae, Aphodiinae) iz srednej Azii i ego sistematicheskoe polozhenie. Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR 88: 39 - 41." type="journal article" year="1979">Nikolajev 1979: 40</bibRefCitation>
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<emphasis id="77869A47FFFCFF9CFEB9FD7FFE6A54D1" box="[350,455,726,747]" italics="true" pageId="5" pageNumber="4">Aphodius</emphasis>
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)
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<emphasis id="77869A47FFFCFF9CFE3EFD7FFDE254D1" box="[473,591,726,747]" italics="true" pageId="5" pageNumber="4">Mendidius</emphasis>
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)
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<emphasis id="77869A47FFFCFF9CFD85FD7FFD4B54D1" box="[610,742,726,747]" italics="true" pageId="5" pageNumber="4">krulikovskii</emphasis>
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);
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<treatmentCitation id="C4536044FFFCFF9CFD18FD7FFC6B54D1" author="Pittino, R." box="[767,966,726,748]" page="307" pageId="5" pageNumber="4" year="1984">
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(as
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<taxonomicName id="82F23DD6FFFCFF9CFC1DFD7FFAB054D1" authorityName="Semenov" authorityYear="1903" box="[1018,1309,726,747]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="krulikovskii">
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||
<emphasis id="77869A47FFFCFF9CFC1DFD7FFAB054D1" box="[1018,1309,726,747]" italics="true" pageId="5" pageNumber="4">Cnemargulus krulikovskii</emphasis>
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||
</taxonomicName>
|
||
);
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||
<treatmentCitation id="C4536044FFFCFF9CFAD0FD7FFE845533" author="Nikolajev, G. V." page="121" pageId="5" pageNumber="4" year="1987">
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</treatmentCitation>
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(as
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<emphasis id="77869A47FFFCFF9CFEB2FD5AFE125532" box="[341,447,755,776]" italics="true" pageId="5" pageNumber="4">Aphodius</emphasis>
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||
(
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||
<emphasis id="77869A47FFFCFF9CFE29FD5AFDE95532" box="[462,580,755,776]" italics="true" pageId="5" pageNumber="4">Mendidius</emphasis>
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||
)
|
||
<emphasis id="77869A47FFFCFF9CFDB5FD5AFD7B5532" box="[594,726,755,776]" italics="true" pageId="5" pageNumber="4">krulikovskii</emphasis>
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</taxonomicName>
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);
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<treatmentCitation id="C4536044FFFCFF9CFD0CFD5DFC665533" author="Dellacasa, M." box="[747,971,756,778]" page="401" pageId="5" pageNumber="4" year="1988">
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<bibRefCitation id="21633BA4FFFCFF9CFD0CFD5DFC665533" author="Dellacasa, M." box="[747,971,756,778]" pageId="5" pageNumber="4" pagination="1 - 455" refId="ref5585" refString="Dellacasa, M. 1988. Contribution to a world-wide catalogue of Aegialiidae Aphodiidae Aulonocnemidae Termitotrogidae. Memorie della Societa entomologica italiana 66 [1987]: 1 - 455" type="journal article" year="1988">Dellacasa 1988: 401</bibRefCitation>
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;
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<treatmentCitation id="C4536044FFFCFF9CFC32FD5DFB055533" author="Dellacasa, G." box="[981,1192,756,778]" page="22" pageId="5" pageNumber="4" year="1990">
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<bibRefCitation id="21633BA4FFFCFF9CFC32FD5DFB055533" author="Dellacasa, G." box="[981,1192,756,778]" pageId="5" pageNumber="4" pagination="19 - 30" refId="ref5414" refString="Dellacasa, G. 1990. The genera Apsteiniella, Cnemargulus, Cnemisus and their type species (Scarabaeoidea Aphodiidae). Frustula entomologica n. s. 13 (26): 19 - 30." type="journal article" year="1990">Dellacasa 1990: 22</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C4536044FFFCFF9CFB54FD5DFF44551C" author="Dellacasa, G. & P. Bordat & M. Dellacasa" page="119" pageId="5" pageNumber="4" year="2001">
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<bibRefCitation id="21633BA4FFFCFF9CFB54FD5DFF44551C" author="Dellacasa, G. & P. Bordat & M. Dellacasa" pageId="5" pageNumber="4" pagination="1 - 482" refId="ref5485" refString="Dellacasa, G., P. Bordat, and M. Dellacasa. 2001. A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera Aphodiidae). Memorie della Societa entomologica italiana 79 [2000]: 1 - 482." type="journal article" year="2001">Dellacasa et al. 2001: 119</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C4536044FFFCFF9CFF13FCB8FDD4551C" author="Dellacasa, M. & G. Dellacasa" box="[244,633,785,807]" page="140" pageId="5" pageNumber="4" year="2006">
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<bibRefCitation id="21633BA4FFFCFF9CFF13FCB8FDD4551C" author="Dellacasa, M. & G. Dellacasa" box="[244,633,785,807]" pageId="5" pageNumber="4" pagination="105 - 143" refId="ref5615" refString="Dellacasa, M., and G. Dellacasa. 2006. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 105 - 143. In Lobl, I. & Smetana, A. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books; Stenstrup. 690 p." type="book chapter" year="2006">Dellacasa and Dellacasa 2006: 140</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C4536044FFFCFF9CFD63FCB8FC90551D" author="Krajcik, M." box="[644,829,785,807]" page="75" pageId="5" pageNumber="4" year="2012">
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<bibRefCitation id="21633BA4FFFCFF9CFD63FCB8FC90551D" author="Krajcik, M." box="[644,829,785,807]" pageId="5" pageNumber="4" pagination="1 - 278" refId="ref5840" refString="Krajcik, M. 2012. Checklist of the World Scarabaeoidea. Animma X. Supplementum 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 75</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="C4536044FFFCFF9CFCAFFCB8FBC8551C" author="Dellacasa, M. & G. Dellacasa & D. Kral & A. Bezdek" box="[840,1125,785,807]" page="121" pageId="5" pageNumber="4" year="2016">
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<bibRefCitation id="21633BA4FFFCFF9CFCAFFCB8FBC8551C" author="Dellacasa, M. & G. Dellacasa & D. Kral & A. Bezdek" box="[840,1125,785,807]" pageId="5" pageNumber="4" pagination="98 - 155" refId="ref5681" refString="Dellacasa, M., G. Dellacasa, D. Kral, and A. Bezdek. 2016. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 98 - 155. In Lobl, I. & Lobl, D. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill; Leiden & Boston. 983 p." type="book chapter" year="2016">Dellacasa et al. 2016: 121</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="454D4655FFFCFF9CFF27FC87FDA455ED" blockId="5.[192,1441,638,983]" pageId="5" pageNumber="4">
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<treatmentCitation id="C4536044FFFCFF9CFF27FC87FDD6557E" author="Semenov, A. P." box="[192,635,814,836]" page="355" pageId="5" pageNumber="4" year="1903">
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<taxonomicName id="82F23DD6FFFCFF9CFF27FC87FDD6557E" authority="Semenov, 1903: 355" authorityName="Semenov" authorityPageNumber="355" authorityYear="1903" box="[192,635,814,836]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="pusio">
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<emphasis id="77869A47FFFCFF9CFF27FC87FE35557E" box="[192,408,814,836]" italics="true" pageId="5" pageNumber="4">Cnemargulus pusio</emphasis>
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<bibRefCitation id="21633BA4FFFCFF9CFE47FC86FDD6557E" author="Semenov, A. P." box="[416,635,814,836]" pageId="5" pageNumber="4" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903: 355</bibRefCitation>
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</taxonomicName>
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||
[type locality: “Prov. Transcaspica: Merv [
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<collectingCountry id="3DE506C5FFFCFF9CFBB0FC87FB50557E" box="[1111,1277,814,836]" name="Turkmenistan" pageId="5" pageNumber="4">Turkmenistan</collectingCountry>
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,
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<collectingRegion id="873688B7FFFCFF9CFAEFFC86FA38557E" box="[1288,1429,814,836]" country="Turkmenistan" name="Mary Welayaty" pageId="5" pageNumber="4">Mary region</collectingRegion>
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]; type material:
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<specimenCount id="53F48DDCFFFCFF9CFE8DFCE5FE48555B" box="[362,485,844,865]" pageId="5" pageNumber="4" type="generic" typeStatus="syntypes">3 syntypes</specimenCount>
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, 2 ♂♂ and
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||
<specimenCount id="53F48DDCFFFCFF9CFD8FFCE5FD3C555B" box="[616,657,844,865]" pageId="5" pageNumber="4" type="female">1 ♀</specimenCount>
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, in ZIN (examined)] (
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<emphasis id="77869A47FFFCFF9CFC6DFCE5FBE9555B" bold="true" box="[906,1092,844,865]" pageId="5" pageNumber="4">new synonymy</emphasis>
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)];
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<treatmentCitation id="C4536044FFFCFF9CFB84FCE5FA96555B" author="Schmidt, A." box="[1123,1339,844,866]" page="13" pageId="5" pageNumber="4" year="1910">
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<bibRefCitation id="21633BA4FFFCFF9CFB84FCE5FA96555B" author="Schmidt, A." box="[1123,1339,844,866]" pageId="5" pageNumber="4" pagination="1 - 155" refId="ref6019" refString="Schmidt, A. 1910 a. Coleoptera Lamellicornia Aphodiinae. P. 1 - 155. In: Wytsman P.: Genera Insectorum. 110. Tervuren." type="book chapter" year="1910">Schmidt, 1910a: 13</bibRefCitation>
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(as
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|
||
<emphasis id="77869A47FFFCFF9CFA8AFCE5FEAC5545" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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</taxonomicName>
|
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);
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<treatmentCitation id="C4536044FFFCFF9CFEFFFCC0FE4D5544" author="Schmidt, A." box="[280,480,873,895]" page="4" pageId="5" pageNumber="4" year="1910">
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<bibRefCitation id="21633BA4FFFCFF9CFEFFFCC0FE4D5544" author="Schmidt, A." box="[280,480,873,895]" pageId="5" pageNumber="4" pagination="1 - 111" refId="ref6049" refString="Schmidt, A. 1910 b. Aphodiinae. Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Berlin 20: 1 - 111." type="journal article" year="1910">Schmidt, 1910b: 4</bibRefCitation>
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</treatmentCitation>
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(as
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<taxonomicName id="82F23DD6FFFCFF9CFDF6FCC0FDD05544" baseAuthorityName="Balthasar" baseAuthorityYear="1964" box="[529,637,873,894]" class="Insecta" family="Aphodiidae" genus="Cnemisus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="77869A47FFFCFF9CFDF6FCC0FDD05544" box="[529,637,873,894]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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</taxonomicName>
|
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);
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<treatmentCitation id="C4536044FFFCFF9CFD74FCC0FCCB5544" author="Schmidt, A." box="[659,870,873,895]" page="344" pageId="5" pageNumber="4" year="1922">
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<bibRefCitation id="21633BA4FFFCFF9CFD74FCC0FCCB5544" author="Schmidt, A." box="[659,870,873,895]" pageId="5" pageNumber="4" refId="ref6079" refString="Schmidt, A. 1922. Coleoptera Aphodiinae. Das Tierreich. Vol. 45. W. de Gruyter and Co.; Berlin & Leipzig. 614 p." type="book" year="1922">Schmidt 1922: 344</bibRefCitation>
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</treatmentCitation>
|
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(as
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<taxonomicName id="82F23DD6FFFCFF9CFC71FCC0FBAF5544" baseAuthorityName="Balthasar" baseAuthorityYear="1964" box="[918,1026,873,894]" class="Insecta" family="Aphodiidae" genus="Cnemisus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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||
<emphasis id="77869A47FFFCFF9CFC71FCC0FBAF5544" box="[918,1026,873,894]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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</taxonomicName>
|
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);
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<treatmentCitation id="C4536044FFFCFF9CFBFFFCC0FB5B5545" author="Winkler, A." box="[1048,1270,873,895]" page="1059" pageId="5" pageNumber="4" year="1929">
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<bibRefCitation id="21633BA4FFFCFF9CFBFFFCC0FB5B5545" author="Winkler, A." box="[1048,1270,873,895]" pageId="5" pageNumber="4" refId="ref6213" refString="Winkler, A. 1929. Catalogus Coleopterorum Regionis Palaearcticae (1924 - 1932). Scarabaeidae. p. 1009 - 1136. A. Winkler; Wien 9. 1698 p." type="book" year="1929">Winkler 1929: 1059</bibRefCitation>
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</treatmentCitation>
|
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(as
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<taxonomicName id="82F23DD6FFFCFF9CFAC1FCC0FA3F5544" baseAuthorityName="Balthasar" baseAuthorityYear="1964" box="[1318,1426,873,894]" class="Insecta" family="Aphodiidae" genus="Cnemisus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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||
<emphasis id="77869A47FFFCFF9CFAC1FCC0FA3F5544" box="[1318,1426,873,894]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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</taxonomicName>
|
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);
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<treatmentCitation id="C4536044FFFCFF9CFF27FC2EFE3255A6" author="Balthasar, V." box="[192,415,903,925]" page="474" pageId="5" pageNumber="4" year="1964">
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<bibRefCitation id="21633BA4FFFCFF9CFF27FC2EFE3255A6" author="Balthasar, V." box="[192,415,903,925]" pageId="5" pageNumber="4" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar 1964: 474</bibRefCitation>
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||
</treatmentCitation>
|
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(as
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||
<taxonomicName id="82F23DD6FFFCFF9CFE2BFC2EFD9555A6" baseAuthorityName="Balthasar" baseAuthorityYear="1964" box="[460,568,903,924]" class="Insecta" family="Aphodiidae" genus="Cnemisus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="77869A47FFFCFF9CFE2BFC2EFD9555A6" box="[460,568,903,924]" italics="true" pageId="5" pageNumber="4">Cnemisus</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="82F23DD6FFFCFF9CFDA1FC2FFD7655A1" authorityName="Semenov" authorityYear="1903" box="[582,731,902,923]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="77869A47FFFCFF9CFDA1FC2FFD7655A1" box="[582,731,902,923]" italics="true" pageId="5" pageNumber="4">Cnemargulus</emphasis>
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</taxonomicName>
|
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));
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||
<treatmentCitation id="C4536044FFFCFF9CFD10FC2EFBD555A6" author="Medvedev, S. I. & L. M. Nikritin" box="[759,1144,903,925]" page="1882" pageId="5" pageNumber="4" year="1971">
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<bibRefCitation id="21633BA4FFFCFF9CFD10FC2EFBD555A6" author="Medvedev, S. I. & L. M. Nikritin" box="[759,1144,903,925]" pageId="5" pageNumber="4" pagination="1881 - 1882" refId="ref5862" refString="Medvedev, S. I., and L. M. Nikritin. 1971. [Once more about the independence of the genus Cnemargulus Sem. (Coleoptera, Scarabaeidae) and its difference from the genus Cnemisus Motsch.] Zoologicheskiy Zhurnal 50: 1881 - 1882." type="journal article" year="1971">Medvedev and Nikritin 1971: 1882</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="C4536044FFFCFF9CFB64FC2EFAE955A6" author="Pittino, R." box="[1155,1348,903,925]" page="307" pageId="5" pageNumber="4" year="1984">
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<bibRefCitation id="21633BA4FFFCFF9CFB64FC2EFAE955A6" author="Pittino, R." box="[1155,1348,903,925]" pageId="5" pageNumber="4" pagination="267 - 359" refId="ref5988" refString="Pittino, R. 1984. Insects of Saudi Arabia. Coleoptera Scarabaeoidea: a revision of the family Aphodiidae. Fauna of Saudi Arabia 6: 267 - 359." type="journal article" year="1984">Pittino 1984: 307</bibRefCitation>
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||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="C4536044FFFCFF9CFAA9FC2EFEFC5583" author="Nikolajev, G. V." page="121" pageId="5" pageNumber="4" year="1987">
|
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<bibRefCitation id="21633BA4FFFCFF9CFAA9FC2EFEFC5583" author="Nikolajev, G. V." pageId="5" pageNumber="4" refId="ref5954" refString="Nikolajev, G. V. 1987. Plastinchatousye zhuki (Coleoptera, Scarabaeoidea) Kazakhstana i Srednej Azii. Instituta Zoologii, Akademiya Nauk Kazakhskoj SSR; Alma Ata. 232 p." type="book" year="1987">Nikolajev 1987: 121</bibRefCitation>
|
||
</treatmentCitation>
|
||
(as
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||
<taxonomicName id="82F23DD6FFFCFF9CFE98FC0DFE445583" authorityName="Illiger" authorityYear="1789" box="[383,489,932,953]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="77869A47FFFCFF9CFE98FC0DFE445583" box="[383,489,932,953]" italics="true" pageId="5" pageNumber="4">Aphodius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="82F23DD6FFFCFF9CFE1FFC0DFDC35583" authorityName="Harold" authorityYear="1868" box="[504,622,932,953]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Mendidius">
|
||
<emphasis id="77869A47FFFCFF9CFE1FFC0DFDC35583" box="[504,622,932,953]" italics="true" pageId="5" pageNumber="4">Mendidius</emphasis>
|
||
</taxonomicName>
|
||
));
|
||
<treatmentCitation id="C4536044FFFCFF9CFD6CFC0DFCC15583" author="Dellacasa, M." box="[651,876,932,954]" page="401" pageId="5" pageNumber="4" year="1988">
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||
<bibRefCitation id="21633BA4FFFCFF9CFD6CFC0DFCC15583" author="Dellacasa, M." box="[651,876,932,954]" pageId="5" pageNumber="4" pagination="1 - 455" refId="ref5585" refString="Dellacasa, M. 1988. Contribution to a world-wide catalogue of Aegialiidae Aphodiidae Aulonocnemidae Termitotrogidae. Memorie della Societa entomologica italiana 66 [1987]: 1 - 455" type="journal article" year="1988">Dellacasa 1988: 401</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="C4536044FFFCFF9CFC9FFC0DFB535583" author="Dellacasa, M. & G. Dellacasa" box="[888,1278,932,954]" page="140" pageId="5" pageNumber="4" year="2006">
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<bibRefCitation id="21633BA4FFFCFF9CFC9FFC0DFB535583" author="Dellacasa, M. & G. Dellacasa" box="[888,1278,932,954]" pageId="5" pageNumber="4" pagination="105 - 143" refId="ref5615" refString="Dellacasa, M., and G. Dellacasa. 2006. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 105 - 143. In Lobl, I. & Smetana, A. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books; Stenstrup. 690 p." type="book chapter" year="2006">Dellacasa and Dellacasa 2006: 140</bibRefCitation>
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</treatmentCitation>
|
||
;
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<treatmentCitation id="C4536044FFFCFF9CFAEDFC0DFF7055ED" author="Krajcik, M." page="75" pageId="5" pageNumber="4" year="2012">
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<bibRefCitation id="21633BA4FFFCFF9CFAEDFC0DFF7055ED" author="Krajcik, M." pageId="5" pageNumber="4" pagination="1 - 278" refId="ref5840" refString="Krajcik, M. 2012. Checklist of the World Scarabaeoidea. Animma X. Supplementum 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 75</bibRefCitation>
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</treatmentCitation>
|
||
;
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<treatmentCitation id="C4536044FFFCFF9CFF0FFC68FDA955ED" author="Dellacasa, M. & G. Dellacasa & D. Kral & A. Bezdek" box="[232,516,961,983]" page="121" pageId="5" pageNumber="4" year="2016">
|
||
<bibRefCitation id="21633BA4FFFCFF9CFF0FFC68FDA955ED" author="Dellacasa, M. & G. Dellacasa & D. Kral & A. Bezdek" box="[232,516,961,983]" pageId="5" pageNumber="4" pagination="98 - 155" refId="ref5681" refString="Dellacasa, M., G. Dellacasa, D. Kral, and A. Bezdek. 2016. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 98 - 155. In Lobl, I. & Lobl, D. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill; Leiden & Boston. 983 p." type="book chapter" year="2016">Dellacasa et al. 2016: 121</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DE815DEFFFCFF9CFF27FC59FC3F5314" pageId="5" pageNumber="4" type="materials_examined">
|
||
<paragraph id="454D4655FFFCFF9CFF27FC59FE2F5213" blockId="5.[192,1440,1008,1066]" pageId="5" pageNumber="4">
|
||
<emphasis id="77869A47FFFCFF9CFF27FC59FED35232" bold="true" box="[192,382,1008,1032]" pageId="5" pageNumber="4">
|
||
<typeStatus id="9A49F8F7FFFCFF9CFF27FC59FEA95232" box="[192,260,1008,1032]" pageId="5" pageNumber="4">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
“Prov. Transcaspica: Imam-baba haud procul a Merv” [
|
||
<collectingCountry id="3DE506C5FFFCFF9CFBA6FC58FB5B5233" box="[1089,1270,1009,1033]" name="Turkmenistan" pageId="5" pageNumber="4">Turkmenistan</collectingCountry>
|
||
, Imam Baba,
|
||
<collectingRegion id="873688B7FFFCFF9CFF27FBBBFEDA5213" box="[192,375,1041,1066]" country="Turkmenistan" name="Mary Welayaty" pageId="5" pageNumber="4">Mary province</collectingRegion>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF27FBEBFA0D5260" blockId="5.[192,1440,1086,1114]" box="[192,1440,1086,1114]" pageId="5" pageNumber="4">
|
||
<emphasis id="77869A47FFFCFF9CFF27FBEBFE295263" bold="true" box="[192,388,1089,1114]" pageId="5" pageNumber="4">Type material.</emphasis>
|
||
<typeStatus id="9A49F8F7FFFCFF9CFE6EFBEBFE525260" box="[393,511,1090,1114]" pageId="5" pageNumber="4" type="lectotype">Lectotype</typeStatus>
|
||
♂, designated by
|
||
<bibRefCitation id="21633BA4FFFCFF9CFD28FBEBFC2B5260" author="Dellacasa, G." box="[719,902,1090,1114]" pageId="5" pageNumber="4" pagination="19 - 30" refId="ref5414" refString="Dellacasa, G. 1990. The genera Apsteiniella, Cnemargulus, Cnemisus and their type species (Scarabaeoidea Aphodiidae). Frustula entomologica n. s. 13 (26): 19 - 30." type="journal article" year="1990">Dellacasa 1990</bibRefCitation>
|
||
, and
|
||
<specimenCount id="53F48DDCFFFCFF9CFC25FBEBFB185263" box="[962,1205,1090,1114]" pageId="5" pageNumber="4" type="female" typeStatus="paralectotypes">2 paralectotypes ♀♀</specimenCount>
|
||
, in
|
||
<collectionCode id="23E3DE90FFFCFF9CFB3AFBEBFAA05263" box="[1245,1293,1090,1113]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="4" type="Museum">ZIN</collectionCode>
|
||
(examined).
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF27FBDBFC3F5314" blockId="5.[192,1440,1136,1326]" pageId="5" pageNumber="4">
|
||
<emphasis id="77869A47FFFCFF9CFF27FBDBFE2652B0" bold="true" box="[192,395,1138,1162]" pageId="5" pageNumber="4">Type labelling.</emphasis>
|
||
1
|
||
<superScript id="B287EB1DFFFCFF9CFE47FBD9FE005244" attach="none" box="[416,429,1136,1150]" fontSize="6" pageId="5" pageNumber="4">st</superScript>
|
||
,
|
||
<emphasis id="77869A47FFFCFF9CFE5DFBDAFE5952B0" box="[442,500,1139,1162]" italics="true" pageId="5" pageNumber="4">recto</emphasis>
|
||
, white with violet frame, printed in violet: “Transcaspien / C. O. Ahnger.” and handwritten in black: [in Cyrillic, transliterated] “Imam baba. / 3.XI.99.”,
|
||
<emphasis id="77869A47FFFCFF9CFB86FB3AFB325290" box="[1121,1183,1171,1194]" italics="true" pageId="5" pageNumber="4">verso</emphasis>
|
||
, white, handwritten in black: [in Cyrillic, transliterated] “peski, v kornjach”; 2
|
||
<superScript id="B287EB1DFFFCFF9CFC61FB18FC355285" attach="none" box="[902,920,1201,1215]" fontSize="6" pageId="5" pageNumber="4">nd</superScript>
|
||
, white, handwritten and printed in black: “
|
||
<taxonomicName id="82F23DD6FFFCFF9CFF2CFB7DFEDD52D6" authorityName="Semenov" authorityYear="1903" box="[203,368,1236,1260]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Cnemargulus</taxonomicName>
|
||
/ Krulikovskii m. / ♂. Typ. X.03 / A. Semenow det.”; 3
|
||
<superScript id="B287EB1DFFFCFF9CFBF2FB78FB8852E5" attach="none" box="[1045,1061,1233,1247]" fontSize="6" pageId="5" pageNumber="4">rd</superScript>
|
||
, white, printed in black: “Zoological Institute /
|
||
<collectingRegion id="873688B7FFFCFF9CFE7DFB5CFDE65337" box="[410,587,1269,1293]" country="Russia" name="Saint Petersburg" pageId="5" pageNumber="4">St. Petersburg</collectingRegion>
|
||
/ INS_COL_0001326”; 4
|
||
<superScript id="B287EB1DFFFCFF9CFC9FFB5BFC2A533A" attach="none" box="[888,903,1266,1280]" fontSize="6" pageId="5" pageNumber="4">th</superScript>
|
||
, red, handwritten in black: “
|
||
<taxonomicName id="82F23DD6FFFCFF9CFB0CFB5CFA3D5337" authorityName="Semenov" authorityYear="1903" box="[1259,1424,1269,1293]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">Cnemargulus</taxonomicName>
|
||
/
|
||
<taxonomicName id="82F23DD6FFFCFF9CFF27FABCFE325314" authority="Sem." authorityName="Semenov" authorityYear="1903" box="[192,415,1301,1326]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="krulikovskyi">krulikovskyi Sem.</taxonomicName>
|
||
/
|
||
<typeStatus id="9A49F8F7FFFCFF9CFE53FABFFD965314" box="[436,571,1302,1326]" pageId="5" pageNumber="4" type="lectotype">Lectotypus</typeStatus>
|
||
♂ / G. Dellacasa des. 1990”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DE815DEFFFCFF9FFF27FAEFFDF3543F" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="diagnosis">
|
||
<paragraph id="454D4655FFFCFF9CFF27FAEFFD7D5345" blockId="5.[192,1440,1350,1929]" pageId="5" pageNumber="4">
|
||
<emphasis id="77869A47FFFCFF9CFF27FAEFFE795364" bold="true" box="[192,468,1350,1374]" pageId="5" pageNumber="4">Diagnostic features.</emphasis>
|
||
Length from 3.2 to 5.0 mm. Colour light testaceous, head, pronotum and elytral suture pale brown; pubescence pale yellow.
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF0FFA2EFB3B5078" blockId="5.[192,1440,1350,1929]" pageId="5" pageNumber="4">Head shiny, convex, epistome distinctly granulate, frontal suture distinct, with slightly raised coarse tubercle at middle and one elongate transverse tubercle each side, and with a fourth, more or less slightly raised tubercle, at middle between frontal suture and anterior edge of clypeus, sometimes closer to the latter; vertex coarsely punctate; genae strongly auriculate, and elongately ciliate, protruding from eyes; clypeus quadridentate anteriorly, inner teeth longer than outer ones, each tooth strongly curved upward, edge between outer teeth and genae more or less straight, never clearly dentate.</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF0FF9E2FD3C509E" blockId="5.[192,1440,1350,1929]" pageId="5" pageNumber="4">Pronotum glossy, transverse, strongly convex, round laterally, punctate, punctures separated by two to three diameters on disc, completely margined, even if anterior margin sometimes barely perceptible, lateral margin with sparse long hairs.</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF0FF904FD4550FE" blockId="5.[192,1440,1350,1929]" box="[232,744,1709,1733]" pageId="5" pageNumber="4">Scutellum triangular, distinctly punctate.</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF0FF964FB9A517D" blockId="5.[192,1440,1350,1929]" pageId="5" pageNumber="4">Elytra elongate-oval, convex, shiny, moderately dilate apically in females, barely more straight in males; striae fine, distinctly impressed, evidently punctate, punctures slightly crenating sides, interstriae nearly flat or only barely convex on disc, sparsely superficially punctate; sutural interstria strongly narrowed apically; humeral apex of epipleural carina visible from above.</paragraph>
|
||
<paragraph id="454D4655FFFCFF9CFF0FF8F9FBA3515D" blockId="5.[192,1440,1350,1929]" box="[232,1038,1872,1896]" pageId="5" pageNumber="4">Metasternum almost flat, smooth, shiny, finely sparsely punctate.</paragraph>
|
||
<paragraph id="454D4655FFFCFF9FFF0FF8D9FA845709" blockId="5.[192,1440,1350,1929]" lastBlockId="6.[192,1440,217,436]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">Foretibiae externally with three strong distal teeth, and with three very small but always distinct proximal teeth; meso- and metatibiae barely widened apically, apical setae quite long, almost unequal, mesotibiae with a single extremely long erected seta, about twice as long as other setae; upper spur of metatibiae strong, flattened, slightly curved, almost as long as the first two tarsal segments.</paragraph>
|
||
<paragraph id="454D4655FFFFFF9FFF0FFE92FCAC5749" blockId="6.[192,1440,217,436]" pageId="6" pageNumber="5">
|
||
Aedeagus (
|
||
<figureCitation id="DDC95AD0FFFFFF9FFE95FE92FE5A5769" box="[370,503,315,339]" captionStart="Figures 7–13" captionStartId="10.[192,287,1762,1784]" captionTargetBox="[324,1278,218,1720]" captionTargetId="figure@10.[263,1370,205,1728]" captionTargetPageId="10" captionText="Figures 7–13. Cnemargulus krulikoƲskyi Semenov, 1903. 7) Habitus of male [lectotype]. 8) Habitus of female [paralectotype, Turkmenistan, Mary prov., Ýolöten]. 9) Habitus of male [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 10) Aedeagus, dorsal view [lectotype]. 11) Aedeagus, dorsal view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 12) Aedeagus, lateral view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 13) Epipharynx [lectotype]. Photos by Augusto Degiovanni, drawings by Ivo Gudenzi." figureDoi="http://doi.org/10.5281/zenodo.3670459" httpUri="https://zenodo.org/record/3670459/files/figure.png" pageId="6" pageNumber="5">Fig. 10–12</figureCitation>
|
||
) elongate, with paramera round and moderately widened apically and, in lateral view, distinctly acuminate toward apex.
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFFFF9FFF0FFED5FD2D578E" blockId="6.[192,1440,217,436]" pageId="6" pageNumber="5">
|
||
Epipharynx (
|
||
<figureCitation id="DDC95AD0FFFFFF9FFE6EFED5FE7157AE" box="[393,476,380,404]" captionStart="Figures 7–13" captionStartId="10.[192,287,1762,1784]" captionTargetBox="[324,1278,218,1720]" captionTargetId="figure@10.[263,1370,205,1728]" captionTargetPageId="10" captionText="Figures 7–13. Cnemargulus krulikoƲskyi Semenov, 1903. 7) Habitus of male [lectotype]. 8) Habitus of female [paralectotype, Turkmenistan, Mary prov., Ýolöten]. 9) Habitus of male [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 10) Aedeagus, dorsal view [lectotype]. 11) Aedeagus, dorsal view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 12) Aedeagus, lateral view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 13) Epipharynx [lectotype]. Photos by Augusto Degiovanni, drawings by Ivo Gudenzi." figureDoi="http://doi.org/10.5281/zenodo.3670459" httpUri="https://zenodo.org/record/3670459/files/figure.png" pageId="6" pageNumber="5">Fig. 13</figureCitation>
|
||
) transverse, round at sides; epitorma oval; corypha with two stout apical celtes as long as, or little more, acropariae.
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFFFF9FFF27FE65FDF3543F" blockId="6.[192,1440,460,517]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FE65FEF357DE" bold="true" box="[192,350,460,484]" pageId="6" pageNumber="5">Variability.</emphasis>
|
||
Clypeal teeth can be more or less developed and more or less acuminate apically and more or less separated from each other.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DE815DEFFFFFF9FFF27FDB4FEA6546C" pageId="6" pageNumber="5" type="distribution">
|
||
<paragraph id="454D4655FFFFFF9FFF27FDB4FEA6546C" blockId="6.[192,1440,541,598]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FDB4FEDE540F" bold="true" box="[192,371,541,565]" pageId="6" pageNumber="5">Distribution.</emphasis>
|
||
<collectingCountry id="3DE506C5FFFFFF9FFE9AFDB7FD80540C" box="[381,557,542,566]" name="Turkmenistan" pageId="6" pageNumber="5">Turkmenistan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="21633BA4FFFFFF9FFDA7FDB7FD54540C" author="Semenov, A. P." box="[576,761,542,566]" pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov 1903</bibRefCitation>
|
||
).
|
||
<collectingCountry id="3DE506C5FFFFFF9FFCF7FDB7FC30540C" box="[784,925,542,566]" name="Kyrgyzstan" pageId="6" pageNumber="5">Kyrgyzstan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="21633BA4FFFFFF9FFC57FDB7FBD9540C" author="Balthasar, V." box="[944,1140,542,566]" pageId="6" pageNumber="5" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar 1964</bibRefCitation>
|
||
).
|
||
<collectingCountry id="3DE506C5FFFFFF9FFB6CFDB7FABA540C" box="[1163,1303,542,566]" name="Uzbekistan" pageId="6" pageNumber="5">Uzbekistan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="21633BA4FFFFFF9FFACEFDB7FF52546C" author="Nikolajev, G. V." pageId="6" pageNumber="5" refId="ref5954" refString="Nikolajev, G. V. 1987. Plastinchatousye zhuki (Coleoptera, Scarabaeoidea) Kazakhstana i Srednej Azii. Instituta Zoologii, Akademiya Nauk Kazakhskoj SSR; Alma Ata. 232 p." type="book" year="1987">Nikolajev 1987</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DE815DEFFFFFF9FFF27FDC7FBB85503" pageId="6" pageNumber="5" type="materials_examined">
|
||
<paragraph id="454D4655FFFFFF9FFF27FDC7FCEF54D2" blockId="6.[192,1440,619,744]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FDC7FE7B54BC" bold="true" box="[192,470,622,646]" pageId="6" pageNumber="5">Material examined.</emphasis>
|
||
<materialsCitation id="F59A4C08FFFFFF9FFE06FDC6FBFA54BD" ID-GBIF-Occurrence="2565890913" box="[481,1111,623,647]" collectionCode="ZIN" country="Turkmenistan" location="Yoloten" pageId="6" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Mary Welayaty" typeStatus="lectotype">
|
||
<collectingCountry id="3DE506C5FFFFFF9FFE06FDC6FD6154BD" box="[481,716,623,647]" name="Turkmenistan" pageId="6" pageNumber="5">TURKMENISTAN</collectingCountry>
|
||
: “Imam baba”(
|
||
<typeStatus id="9A49F8F7FFFFFF9FFC68FDC6FBA954BD" box="[911,1028,623,647]" pageId="6" pageNumber="5" type="lectotype">lectotype</typeStatus>
|
||
,
|
||
<collectionCode id="23E3DE90FFFFFF9FFBF3FDC6FBE754BD" box="[1044,1098,623,647]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="5" type="Museum">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="F59A4C08FFFFFF9FFB86FDC6FB31549D" ID-GBIF-Occurrence="2565890906" collectionCode="ZIN" collectorName="Ahnger" country="Turkmenistan" isTailEnumeration="true" location="Yoloten" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="2" stateProvince="Mary Welayaty" typeStatus="paralectotype">
|
||
“Jolatan” [
|
||
<location id="402D108EFFFFFF9FFB0DFDC2FAE754BC" LSID="urn:lsid:plazi:treatment:CD5BF743FFFCFF9EFF27FD87FA0D542C:402D108EFFFFFF9FFB0DFDC2FAE754BC" box="[1258,1354,619,646]" country="Turkmenistan" name="Yoloten" pageId="6" pageNumber="5" stateProvince="Mary Welayaty">Ýolöten</location>
|
||
,
|
||
<collectingRegion id="873688B7FFFFFF9FFABAFDC6FE80549D" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary province</collectingRegion>
|
||
],
|
||
<date id="314C6095FFFFFF9FFEA6FD26FECC549D" box="[321,353,655,679]" pageId="6" pageNumber="5" value="1899-04-24">24</date>
|
||
.iv.1899 and
|
||
<date id="314C6095FFFFFF9FFE18FD26FDD3549D" box="[511,638,655,679]" pageId="6" pageNumber="5" value="1899-04-26">26.iv.1899</date>
|
||
, C.
|
||
<collectorName id="E8072383FFFFFF9FFD57FD26FCA6549D" box="[688,779,655,679]" pageId="6" pageNumber="5">Ahnger</collectorName>
|
||
leg.
|
||
<specimenCount id="53F48DDCFFFFFF9FFCAFFD26FC2A549D" box="[840,903,655,679]" pageId="6" pageNumber="5" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<typeStatus id="9A49F8F7FFFFFF9FFC7FFD26FBE3549D" box="[920,1102,655,679]" pageId="6" pageNumber="5" type="paralectotype">paralectotypes</typeStatus>
|
||
,
|
||
<collectionCode id="23E3DE90FFFFFF9FFBBBFD26FB3D549D" box="[1116,1168,655,679]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="5" type="Museum">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="F59A4C08FFFFFF9FFB42FD26FBC054F2" ID-GBIF-Occurrence="2565890910" collectionCode="ZIN" collectorName="Ahnger" country="Turkmenistan" isTailEnumeration="true" location="Yoloten" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="2" stateProvince="Mary Welayaty" typeStatus="syntype">
|
||
“Merv” [
|
||
<collectingRegion id="873688B7FFFFFF9FFAEDFD39FAFD5492" box="[1290,1360,656,680]" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary</collectingRegion>
|
||
,
|
||
<collectingRegion id="873688B7FFFFFF9FFAB9FD39FE8054F2" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary province</collectingRegion>
|
||
],
|
||
<date id="314C6095FFFFFF9FFED9FD19FEF654F2" box="[318,347,688,712]" pageId="6" pageNumber="5" value="1900-03-18">18</date>
|
||
.iii.1900, C.
|
||
<collectorName id="E8072383FFFFFF9FFE0AFD19FDE554F2" box="[493,584,688,712]" pageId="6" pageNumber="5">Ahnger</collectorName>
|
||
leg. 2 ♂♂ and
|
||
<specimenCount id="53F48DDCFFFFFF9FFD10FD19FC8D54FD" box="[759,800,688,712]" pageId="6" pageNumber="5" type="female">1 ♀</specimenCount>
|
||
(
|
||
<typeStatus id="9A49F8F7FFFFFF9FFCC9FD19FC3754F2" box="[814,922,688,712]" pageId="6" pageNumber="5" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="82F23DD6FFFFFF9FFC5AFD19FB8F54F2" authorityName="Semenov" authorityYear="1903" box="[957,1058,688,712]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">C. pusio</taxonomicName>
|
||
,
|
||
<collectionCode id="23E3DE90FFFFFF9FFBCAFD19FBCC54F2" box="[1069,1121,688,712]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="5" type="Museum">ZIN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="F59A4C08FFFFFF9FFB94FD19FCEF54D2" ID-GBIF-Occurrence="2565890907" collectionCode="ZIN" collectorName="Ahnger" country="Turkmenistan" isTailEnumeration="true" location="Yoloten" pageId="6" pageNumber="5" specimenCount="1" specimenCount-female="2" stateProvince="Mary Welayaty" typeStatus="lectotype">
|
||
“Jolatan” [
|
||
<location id="402D108EFFFFFF9FFB14FD05FAFD54FD" LSID="urn:lsid:plazi:treatment:CD5BF743FFFCFF9EFF27FD87FA0D542C:402D108EFFFFFF9FFB14FD05FAFD54FD" box="[1267,1360,684,711]" country="Turkmenistan" name="Yoloten" pageId="6" pageNumber="5" stateProvince="Mary Welayaty">Ýolöten</location>
|
||
,
|
||
<collectingRegion id="873688B7FFFFFF9FFAB9FD19FE8054D2" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary province</collectingRegion>
|
||
],
|
||
<date id="314C6095FFFFFF9FFED8FD79FEF154D2" box="[319,348,720,744]" pageId="6" pageNumber="5" value="1927-03-15">15</date>
|
||
.iii.1927, V.
|
||
<collectorName id="E8072383FFFFFF9FFE17FD79FDDD54D2" box="[496,624,720,744]" pageId="6" pageNumber="5">Kiseritzky</collectorName>
|
||
leg. 2 exx. (
|
||
<collectionCode id="23E3DE90FFFFFF9FFCE7FD79FC9854D2" box="[768,821,720,744]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="5" type="Museum">ZIN</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFFFF9FFF27FCA9FBB85503" blockId="6.[192,1440,768,825]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FCA9FE645522" bold="true" box="[192,457,768,792]" pageId="6" pageNumber="5">Literature records.</emphasis>
|
||
<collectingCountry id="3DE506C5FFFFFF9FFE36FCA8FDCC5523" box="[465,609,769,793]" name="Kyrgyzstan" pageId="6" pageNumber="5">Kyrgyzstan</collectingCountry>
|
||
,
|
||
<location id="402D108EFFFFFF9FFD89FCA8FD465523" LSID="urn:lsid:plazi:treatment:CD5BF743FFFCFF9EFF27FD87FA0D542C:402D108EFFFFFF9FFD89FCA8FD465523" box="[622,747,769,793]" country="Kyrgyzstan" name="Sussamyr" pageId="6" pageNumber="5">Sussamyr</location>
|
||
,
|
||
<specimenCount id="53F48DDCFFFFFF9FFD1FFCA8FC835522" box="[760,814,769,793]" pageId="6" pageNumber="5" type="generic">1 ex</specimenCount>
|
||
. (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFCA5FCA9FBA95522" author="Balthasar, V." box="[834,1028,768,792]" pageId="6" pageNumber="5" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar 1964</bibRefCitation>
|
||
).
|
||
<collectingCountry id="3DE506C5FFFFFF9FFBFEFCA9FB055522" box="[1049,1192,768,792]" name="Uzbekistan" pageId="6" pageNumber="5">Uzbekistan</collectingCountry>
|
||
,
|
||
<location id="402D108EFFFFFF9FFB52FCA9FA315522" LSID="urn:lsid:plazi:treatment:CD5BF743FFFCFF9EFF27FD87FA0D542C:402D108EFFFFFF9FFB52FCA9FA315522" box="[1205,1436,768,792]" country="Uzbekistan" name="Central Kyzylkum" pageId="6" pageNumber="5">Central Kyzylkum</location>
|
||
,
|
||
<specimenCount id="53F48DDCFFFFFF9FFF27FC88FF585502" box="[192,245,801,825]" pageId="6" pageNumber="5" type="generic">1 ex</specimenCount>
|
||
. (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFEEFFC88FE685503" author="Nikolajev, G. V." box="[264,453,801,825]" pageId="6" pageNumber="5" refId="ref5954" refString="Nikolajev, G. V. 1987. Plastinchatousye zhuki (Coleoptera, Scarabaeoidea) Kazakhstana i Srednej Azii. Instituta Zoologii, Akademiya Nauk Kazakhskoj SSR; Alma Ata. 232 p." type="book" year="1987">Nikolajev 1987</bibRefCitation>
|
||
).
|
||
<collectingCountry id="3DE506C5FFFFFF9FFE3FFC88FD265503" box="[472,651,801,825]" name="Turkmenistan" pageId="6" pageNumber="5">Turkmenistan</collectingCountry>
|
||
,
|
||
<location id="402D108EFFFFFF9FFD70FC88FD5B5503" LSID="urn:lsid:plazi:treatment:CD5BF743FFFCFF9EFF27FD87FA0D542C:402D108EFFFFFF9FFD70FC88FD5B5503" box="[663,758,801,825]" country="Turkmenistan" name="Jolatan" pageId="6" pageNumber="5">Jolatan</location>
|
||
,
|
||
<specimenCount id="53F48DDCFFFFFF9FFCE5FC88FC9A5502" box="[770,823,801,825]" pageId="6" pageNumber="5" type="generic">1 ex</specimenCount>
|
||
. (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFCACFC88FBA55503" author="Dellacasa, G." box="[843,1032,801,825]" pageId="6" pageNumber="5" pagination="19 - 30" refId="ref5414" refString="Dellacasa, G. 1990. The genera Apsteiniella, Cnemargulus, Cnemisus and their type species (Scarabaeoidea Aphodiidae). Frustula entomologica n. s. 13 (26): 19 - 30." type="journal article" year="1990">Dellacasa 1990</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DE815DEFFFFFF9EFF27FCF8FA0D542C" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5" type="discussion">
|
||
<paragraph id="454D4655FFFFFF9FFF27FCF8FC93534A" blockId="6.[192,1440,849,1393]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FCF8FEF05553" bold="true" box="[192,349,849,873]" pageId="6" pageNumber="5">Discussion.</emphasis>
|
||
<taxonomicName id="82F23DD6FFFFFF9FFE84FCF8FD0B5553" authorityName="Semenov" authorityYear="1903" box="[355,678,849,873]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFE84FCF8FD0B5553" box="[355,678,849,873]" italics="true" pageId="6" pageNumber="5">Cnemargulus krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="82F23DD6FFFFFF9FFD04FCFBFCE85553" authorityName="Semenov" authorityYear="1903" box="[739,837,850,873]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFD04FCFBFCE85553" box="[739,837,850,873]" italics="true" pageId="6" pageNumber="5">C. pusio</emphasis>
|
||
</taxonomicName>
|
||
, described by
|
||
<bibRefCitation id="21633BA4FFFFFF9FFC14FCFBFB185550" author="Semenov, A. P." box="[1011,1205,850,874]" pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov (1903)</bibRefCitation>
|
||
in the same paper, have as their type localities Imam Baba for the former and
|
||
<collectingRegion id="873688B7FFFFFF9FFC9AFCDBFC1355B0" box="[893,958,882,906]" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary</collectingRegion>
|
||
for the latter. Imam Baba, if understood as the archaeological site of the Mausoleum of Imam Baba, near Sandykachi, is
|
||
<quantity id="820AEBB0FFFFFF9FFB6EFC3AFB795590" box="[1161,1236,914,939]" metricMagnitude="4" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="5" unit="km" value="78.0">78 km</quantity>
|
||
south of Ýolöten (the locality of the two
|
||
<typeStatus id="9A49F8F7FFFFFF9FFE36FC1AFD2955F1" box="[465,644,947,971]" pageId="6" pageNumber="5" type="paralectotype">paralectotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="82F23DD6FFFFFF9FFD42FC1AFCF055F0" authorityName="Semenov" authorityYear="1903" box="[677,861,946,970]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFD42FC1AFCF055F0" box="[677,861,946,970]" italics="true" pageId="6" pageNumber="5">C. krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
), and some
|
||
<quantity id="820AEBB0FFFFFF9FFC0CFC1AFBE955F1" box="[1003,1092,947,971]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="5" unit="km" value="120.0">120 km</quantity>
|
||
southeast of
|
||
<collectingRegion id="873688B7FFFFFF9FFB05FC1AFA8555F1" box="[1250,1320,947,971]" country="Turkmenistan" name="Mary Welayaty" pageId="6" pageNumber="5">Mary</collectingRegion>
|
||
.
|
||
<bibRefCitation id="21633BA4FFFFFF9FFAD6FC1AFEA155D1" author="Semenov, A. P." pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov (1903)</bibRefCitation>
|
||
supplied some characters for distinguishing the two taxa. He wrote that
|
||
<taxonomicName id="82F23DD6FFFFFF9FFB67FC7AFB4F55D1" authorityName="Semenov" authorityYear="1903" box="[1152,1250,979,1003]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFB67FC7AFB4F55D1" box="[1152,1250,979,1003]" italics="true" pageId="6" pageNumber="5">C. pusio</emphasis>
|
||
</taxonomicName>
|
||
is smaller than
|
||
<taxonomicName id="82F23DD6FFFFFF9FFF27FC5DFED75231" authorityName="Semenov" authorityYear="1903" box="[192,378,1011,1035]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFF27FC5DFED75231" box="[192,378,1011,1035]" italics="true" pageId="6" pageNumber="5">C. krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
, paler, has granules more spaced on the head, clypeal apical teeth more produced and less separated from each other, the distal ones more lateral, the space between the apical tooth and genal suture not crenulate, the frontal carina slightly trituberculate, the pronotum wider, anterior angles broader and elytral interstriae almost convex and shiny. In his dichotomous key,
|
||
<bibRefCitation id="21633BA4FFFFFF9FFB99FBFCFA9E5257" author="Schmidt, A." box="[1150,1331,1109,1133]" pageId="6" pageNumber="5" refId="ref6079" refString="Schmidt, A. 1922. Coleoptera Aphodiinae. Das Tierreich. Vol. 45. W. de Gruyter and Co.; Berlin & Leipzig. 614 p." type="book" year="1922">Schmidt (1922)</bibRefCitation>
|
||
reported only one character to distinguish the two taxa, namely the space between the clypeal apical tooth and the genal suture, which is crenulate in
|
||
<taxonomicName id="82F23DD6FFFFFF9FFD40FB3FFCCC5294" authorityName="Semenov" authorityYear="1903" box="[679,865,1174,1198]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFD40FB3FFCCC5294" box="[679,865,1174,1198]" italics="true" pageId="6" pageNumber="5">C. krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
and uncrenulate in
|
||
<taxonomicName id="82F23DD6FFFFFF9FFBBAFB3FFB6C5297" authorityName="Semenov" authorityYear="1903" box="[1117,1217,1174,1197]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFBBAFB3FFB6C5297" box="[1117,1217,1174,1197]" italics="true" pageId="6" pageNumber="5">C. pusio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="21633BA4FFFFFF9FFB36FB3FFA0D5294" author="Balthasar, V." box="[1233,1440,1174,1198]" pageId="6" pageNumber="5" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar (1964)</bibRefCitation>
|
||
reported this character as well, and also added the distinct frontal carina in
|
||
<taxonomicName id="82F23DD6FFFFFF9FFB93FB1EFA8352F4" authorityName="Semenov" authorityYear="1903" box="[1140,1326,1206,1230]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFB93FB1EFA8352F4" box="[1140,1326,1206,1230]" italics="true" pageId="6" pageNumber="5">C. krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
(weak in
|
||
<taxonomicName id="82F23DD6FFFFFF9FFF27FB7EFE8E52D4" authorityName="Semenov" authorityYear="1903" box="[192,291,1239,1262]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFF27FB7EFE8E52D4" box="[192,291,1239,1262]" italics="true" pageId="6" pageNumber="5">C. pusio</emphasis>
|
||
</taxonomicName>
|
||
), the less-shiny pronotum, and the elytral interstriae almost flat and impunctuate (pronotum shinier and elytral interstriae convex and clearly punctate). In my opinion, these characters fall within infraspecific variability, and can be observed, more or less evident, in all the specimens I have examined. For these reasons, and for paramera that present a remarkable uniformity (
|
||
<figureCitation id="DDC95AD0FFFFFF9FFBADFA91FB67536B" box="[1098,1226,1336,1361]" captionStart="Figures 7–13" captionStartId="10.[192,287,1762,1784]" captionTargetBox="[324,1278,218,1720]" captionTargetId="figure@10.[263,1370,205,1728]" captionTargetPageId="10" captionText="Figures 7–13. Cnemargulus krulikoƲskyi Semenov, 1903. 7) Habitus of male [lectotype]. 8) Habitus of female [paralectotype, Turkmenistan, Mary prov., Ýolöten]. 9) Habitus of male [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 10) Aedeagus, dorsal view [lectotype]. 11) Aedeagus, dorsal view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 12) Aedeagus, lateral view [syntype of C. pusio Semenov, 1903: Turkmenistan, Mary prov., Ýolöten]. 13) Epipharynx [lectotype]. Photos by Augusto Degiovanni, drawings by Ivo Gudenzi." figureDoi="http://doi.org/10.5281/zenodo.3670459" httpUri="https://zenodo.org/record/3670459/files/figure.png" pageId="6" pageNumber="5">Fig. 10–11</figureCitation>
|
||
), I herein propose the synonymy
|
||
<taxonomicName id="82F23DD6FFFFFF9FFE94FAF0FD1B534B" authorityName="Semenov" authorityYear="1903" box="[371,694,1369,1393]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFE94FAF0FD1B534B" box="[371,694,1369,1393]" italics="true" pageId="6" pageNumber="5">Cnemargulus krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="82F23DD6FFFFFF9FFD33FAF0FC9A534A" authorityName="Semenov" authorityYear="1903" box="[724,823,1369,1392]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFD33FAF0FC9A534A" box="[724,823,1369,1392]" italics="true" pageId="6" pageNumber="5">C. pusio</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFFFF9FFF27FA20FE2A50BF" blockId="6.[192,1440,1417,1929]" pageId="6" pageNumber="5">
|
||
<emphasis id="77869A47FFFFFF9FFF27FA20FDBF539B" bold="true" box="[192,530,1417,1441]" pageId="6" pageNumber="5">Nomenclatural remarks.</emphasis>
|
||
Articles 24.2.1 and 24.2.2 of the International Code of Zoological Nomenclature (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFEC2FA03FE1D53F8" author="ICZN & International Commission on Zoological Nomenclature" box="[293,432,1450,1474]" pageId="6" pageNumber="5" refId="ref5761" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences (Fourth Edition). International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), from now on called the Code, stated that when the precedence between names or nomenclatural acts cannot be objectively determined, the precedence is fixed by the action of the first author – termed “First Reviser” - citing in a published work those names or acts and selecting from them. The “First Reviser” fixes the precedence of a name or an act if those names or acts are published on the same date in the same work. As “First Reviser”, I herein fix the precedence of
|
||
<taxonomicName id="82F23DD6FFFFFF9FFB1CF982FDB8505E" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFB1CF982FEFA505E" italics="true" pageId="6" pageNumber="5">Cnemargulus krulikovskyi</emphasis>
|
||
<bibRefCitation id="21633BA4FFFFFF9FFEBAF9E5FDB8505E" author="Semenov, A. P." box="[349,533,1612,1636]" pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
upon
|
||
<taxonomicName id="82F23DD6FFFFFF9FFDB9F9E5FBA1505E" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[606,1036,1612,1636]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pusio">
|
||
<emphasis id="77869A47FFFFFF9FFDB9F9E5FCE6505E" box="[606,843,1612,1636]" italics="true" pageId="6" pageNumber="5">Cnemargulus pusio</emphasis>
|
||
<bibRefCitation id="21633BA4FFFFFF9FFCB5F9E5FBA1505E" author="Semenov, A. P." box="[850,1036,1612,1636]" pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is therefore considered its junior synonym.
|
||
</paragraph>
|
||
<paragraph id="454D4655FFFFFF9EFF0FF924FA0D542C" blockId="6.[192,1440,1417,1929]" lastBlockId="7.[192,1441,221,534]" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">
|
||
<bibRefCitation id="21633BA4FFFFFF9FFF0FF924FE07509F" author="Semenov, A. P." box="[232,426,1677,1701]" pageId="6" pageNumber="5" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov (1903)</bibRefCitation>
|
||
described the species as
|
||
<taxonomicName id="82F23DD6FFFFFF9FFD3BF924FBB7509F" authorityName="Semenov" authorityYear="1903" box="[732,1050,1677,1701]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskii">
|
||
<emphasis id="77869A47FFFFFF9FFD3BF924FBB7509F" box="[732,1050,1677,1701]" italics="true" pageId="6" pageNumber="5">Cnemargulus krulikovskii</emphasis>
|
||
</taxonomicName>
|
||
, and in a footnote declared that it was dedicated to L. C. Krulikovsky [Leonid Konstantinovitsch Krulikowski (
|
||
<date id="314C6095FFFFFF9FFB6AF907FAA250FC" box="[1165,1295,1710,1734]" pageId="6" pageNumber="5" value="1864" valueMax="1920">1864-1920</date>
|
||
), a Russian lepidopterologist (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFE7CF967FD2550DC" author="Schmitt, M. & H. Hubner & R. Gaedike" box="[411,648,1742,1766]" pageId="6" pageNumber="5" pagination="1 - 192" refId="ref6110" refString="Schmitt, M., H. Hubner, and R. Gaedike. 1998. Nomina auctorum - Aflosung von Abkurzungen taxonomischer Autoren-Namen. Nova Supplementa Entomologica 11: 1 - 192." type="journal article" year="1998">Schmitt et al. 1998</bibRefCitation>
|
||
)]. Therefore,
|
||
<taxonomicName id="82F23DD6FFFFFF9FFCCEF967FC1650DC" authorityName="Semenov" authorityYear="1903" box="[809,955,1742,1766]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskii">
|
||
<emphasis id="77869A47FFFFFF9FFCCEF967FC1650DC" box="[809,955,1742,1766]" italics="true" pageId="6" pageNumber="5">krulikovskii</emphasis>
|
||
</taxonomicName>
|
||
is the correct original spelling, and it is not possible to apply article 32.5 of the Code (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFD09F946FCD4513D" author="ICZN & International Commission on Zoological Nomenclature" box="[750,889,1775,1799]" pageId="6" pageNumber="5" refId="ref5761" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences (Fourth Edition). International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
). The first author who used
|
||
<taxonomicName id="82F23DD6FFFFFF9FFB35F947FAC4513C" authorityName="Semenov" authorityYear="1903" box="[1234,1385,1774,1798]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFFFF9FFB35F947FAC4513C" box="[1234,1385,1774,1798]" italics="true" pageId="6" pageNumber="5">krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
was
|
||
<bibRefCitation id="21633BA4FFFFFF9FFF27F8A6FED3511D" author="Schmidt, A." box="[192,382,1807,1831]" pageId="6" pageNumber="5" pagination="1 - 155" refId="ref6019" refString="Schmidt, A. 1910 a. Coleoptera Lamellicornia Aphodiinae. P. 1 - 155. In: Wytsman P.: Genera Insectorum. 110. Tervuren." type="book chapter" year="1910">Schmidt (1910a</bibRefCitation>
|
||
, b), but he did not demonstrate an intentional change in the original spelling of the name. Without an explicit statement of intention, the name should be considered as a subsequent incorrect spelling, and it should not enter into homonymy and cannot be used as a substitute name. Nevertheless, according to article 33.3.1 of the Code (
|
||
<bibRefCitation id="21633BA4FFFFFF9FFD70F8D8FC8C51B3" author="ICZN & International Commission on Zoological Nomenclature" box="[663,801,1905,1929]" pageId="6" pageNumber="5" refId="ref5761" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences (Fourth Edition). International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), if the incorrect subsequent spelling is in prevailing usage and attributed to the publication of the original spelling, it is preserved and deemed to be the correct original spelling. In the glossary of the Code (
|
||
<bibRefCitation id="21633BA4FFFEFF9EFCB7FF54FC76572F" author="ICZN & International Commission on Zoological Nomenclature" box="[848,987,253,277]" pageId="7" pageNumber="6" refId="ref5761" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences (Fourth Edition). International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), it is explained that a name is to be considered of prevailing usage when its usage is adopted by at least a substantial majority of the most recent authors concerned with the relevant taxon. In our case, the name
|
||
<taxonomicName id="82F23DD6FFFEFF9EFBAFFE94FA0D576C" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[1096,1440,317,342]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFEFF9EFBAFFE94FB72576F" box="[1096,1247,317,341]" italics="true" pageId="7" pageNumber="6">krulikovskyi</emphasis>
|
||
<bibRefCitation id="21633BA4FFFEFF9EFB00FE97FA0D576C" author="Semenov, A. P." box="[1255,1440,318,342]" pageId="7" pageNumber="6" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
has been used nine times (
|
||
<bibRefCitation id="21633BA4FFFEFF9EFE19FEF7FD15574C" author="Schmidt, A." box="[510,696,350,374]" pageId="7" pageNumber="6" pagination="1 - 155" refId="ref6019" refString="Schmidt, A. 1910 a. Coleoptera Lamellicornia Aphodiinae. P. 1 - 155. In: Wytsman P.: Genera Insectorum. 110. Tervuren." type="book chapter" year="1910">Schmidt 1910a</bibRefCitation>
|
||
, b;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFD39FEF7FC2B574C" author="Winkler, A." box="[734,902,350,374]" pageId="7" pageNumber="6" refId="ref6213" refString="Winkler, A. 1929. Catalogus Coleopterorum Regionis Palaearcticae (1924 - 1932). Scarabaeidae. p. 1009 - 1136. A. Winkler; Wien 9. 1698 p." type="book" year="1929">Winkler 1929</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFC76FEF7FBE1574C" author="Dellacasa, M." box="[913,1100,350,374]" pageId="7" pageNumber="6" pagination="1 - 455" refId="ref5585" refString="Dellacasa, M. 1988. Contribution to a world-wide catalogue of Aegialiidae Aphodiidae Aulonocnemidae Termitotrogidae. Memorie della Societa entomologica italiana 66 [1987]: 1 - 455" type="journal article" year="1988">Dellacasa 1988</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="21633BA4FFFEFF9EFBB1FEF7FB39574C" author="Dellacasa, G." box="[1110,1172,350,374]" pageId="7" pageNumber="6" pagination="19 - 30" refId="ref5414" refString="Dellacasa, G. 1990. The genera Apsteiniella, Cnemargulus, Cnemisus and their type species (Scarabaeoidea Aphodiidae). Frustula entomologica n. s. 13 (26): 19 - 30." type="journal article" year="1990">1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFB79FEF7FA36574C" author="Dellacasa, G. & P. Bordat & M. Dellacasa" box="[1182,1435,350,374]" pageId="7" pageNumber="6" pagination="1 - 482" refId="ref5485" refString="Dellacasa, G., P. Bordat, and M. Dellacasa. 2001. A revisional essay of world genus-group taxa of Aphodiinae (Coleoptera Aphodiidae). Memorie della Societa entomologica italiana 79 [2000]: 1 - 482." type="journal article" year="2001">Dellacasa et al. 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="21633BA4FFFEFF9EFF27FED7FF5357AC" author="Dellacasa, M. & G. Dellacasa & D. Kral & A. Bezdek" box="[192,254,382,406]" pageId="7" pageNumber="6" pagination="98 - 155" refId="ref5681" refString="Dellacasa, M., G. Dellacasa, D. Kral, and A. Bezdek. 2016. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 98 - 155. In Lobl, I. & Lobl, D. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill; Leiden & Boston. 983 p." type="book chapter" year="2016">2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFEEFFED7FDDF57AC" author="Dellacasa, M. & G. Dellacasa" box="[264,626,382,406]" pageId="7" pageNumber="6" pagination="105 - 143" refId="ref5615" refString="Dellacasa, M., and G. Dellacasa. 2006. Scarabaeidae, subfamily Aphodiinae, tribe Aphodiini, p. 105 - 143. In Lobl, I. & Smetana, A. (Eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books; Stenstrup. 690 p." type="book chapter" year="2006">Dellacasa and Dellacasa 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFD9BFED7FCB757AC" author="Krajcik, M." box="[636,794,382,406]" pageId="7" pageNumber="6" pagination="1 - 278" refId="ref5840" refString="Krajcik, M. 2012. Checklist of the World Scarabaeoidea. Animma X. Supplementum 5: 1 - 278." type="journal article" year="2012">Krajcik 2012</bibRefCitation>
|
||
), the name
|
||
<taxonomicName id="82F23DD6FFFEFF9EFC43FED4FB5F57AC" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[932,1266,381,406]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="krulikovskii">
|
||
<emphasis id="77869A47FFFEFF9EFC43FED4FB9B57AF" box="[932,1078,381,405]" italics="true" pageId="7" pageNumber="6">krulikovskii</emphasis>
|
||
<bibRefCitation id="21633BA4FFFEFF9EFBDCFED7FB5F57AC" author="Semenov, A. P." box="[1083,1266,382,406]" pageId="7" pageNumber="6" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
has been used five times (
|
||
<bibRefCitation id="21633BA4FFFEFF9EFEAFFE37FDA4578C" author="Balthasar, V." box="[328,521,414,438]" pageId="7" pageNumber="6" refId="ref5378" refString="Balthasar, V. 1964. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 3. Aphodiidae. Tschechoslowakischen Akademie der Wissenschaften; Prague. 652 p." type="book" year="1964">Balthasar 1964</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFDF2FE37FCD7578C" author="Medvedev, S. I. & L. M. Nikritin" box="[533,890,414,438]" pageId="7" pageNumber="6" pagination="1881 - 1882" refId="ref5862" refString="Medvedev, S. I., and L. M. Nikritin. 1971. [Once more about the independence of the genus Cnemargulus Sem. (Coleoptera, Scarabaeidae) and its difference from the genus Cnemisus Motsch.] Zoologicheskiy Zhurnal 50: 1881 - 1882." type="journal article" year="1971">Medvedev and Nikritin 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFC60FE37FBEF578C" author="Nikolajev, G. V." box="[903,1090,414,438]" pageId="7" pageNumber="6" pagination="39 - 41" refId="ref5913" refString="Nikolajev, G. V. 1979. Novyi vid plastinchatousykh triby Aphodiini (Coleoptera, Scarabaeidae, Aphodiinae) iz srednej Azii i ego sistematicheskoe polozhenie. Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR 88: 39 - 41." type="journal article" year="1979">Nikolajev 1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="21633BA4FFFEFF9EFBA8FE37FB20578C" author="Nikolajev, G. V." box="[1103,1165,414,438]" pageId="7" pageNumber="6" refId="ref5954" refString="Nikolajev, G. V. 1987. Plastinchatousye zhuki (Coleoptera, Scarabaeoidea) Kazakhstana i Srednej Azii. Instituta Zoologii, Akademiya Nauk Kazakhskoj SSR; Alma Ata. 232 p." type="book" year="1987">1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="21633BA4FFFEFF9EFB7EFE37FA9F578C" author="Pittino, R." box="[1177,1330,414,438]" pageId="7" pageNumber="6" pagination="267 - 359" refId="ref5988" refString="Pittino, R. 1984. Insects of Saudi Arabia. Coleoptera Scarabaeoidea: a revision of the family Aphodiidae. Fauna of Saudi Arabia 6: 267 - 359." type="journal article" year="1984">Pittino 1984</bibRefCitation>
|
||
) and the name
|
||
<taxonomicName id="82F23DD6FFFEFF9EFEEDFE17FDC557EC" authority="Semenov, 1903" authorityName="Semenov" authorityYear="1903" box="[266,616,446,470]" class="Insecta" family="Aphodiidae" genus="Cnemisus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="krulikovskyii">
|
||
<emphasis id="77869A47FFFEFF9EFEEDFE17FE0457EC" box="[266,425,446,470]" italics="true" pageId="7" pageNumber="6">krulikovskyii</emphasis>
|
||
<bibRefCitation id="21633BA4FFFEFF9EFE57FE17FDC557EC" author="Semenov, A. P." box="[432,616,446,470]" pageId="7" pageNumber="6" pagination="353 - 357" refId="ref6144" refString="Semenov, A. P. 1903. Coleoptera nova faunae turanicae. II. Russkoe Entomologicheskoe Obozrenie 3: 353 - 357." type="journal article" year="1903">Semenov, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
only once (
|
||
<bibRefCitation id="21633BA4FFFEFF9EFD09FE17FC3757EC" author="Schmidt, A." box="[750,922,446,470]" pageId="7" pageNumber="6" refId="ref6079" refString="Schmidt, A. 1922. Coleoptera Aphodiinae. Das Tierreich. Vol. 45. W. de Gruyter and Co.; Berlin & Leipzig. 614 p." type="book" year="1922">Schmidt 1922</bibRefCitation>
|
||
). In addition, the more recent authors that dealt with the subject have used the spelling
|
||
<taxonomicName id="82F23DD6FFFEFF9EFD00FE77FCD357CC" authorityName="Semenov" authorityYear="1903" box="[743,894,478,502]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFEFF9EFD00FE77FCD357CC" box="[743,894,478,502]" italics="true" pageId="7" pageNumber="6">krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
. For that reasons, under article 33.3.1 of the Code (
|
||
<bibRefCitation id="21633BA4FFFEFF9EFEEEFE57FE3D542C" author="ICZN & International Commission on Zoological Nomenclature" box="[265,400,510,534]" pageId="7" pageNumber="6" refId="ref5761" refString="ICZN [International Commission on Zoological Nomenclature]. 1999. International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences (Fourth Edition). International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
), the spelling
|
||
<taxonomicName id="82F23DD6FFFEFF9EFDD3FE57FD64542C" authorityName="Semenov" authorityYear="1903" box="[564,713,510,534]" class="Insecta" family="Aphodiidae" genus="Cnemargulus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="krulikovskyi">
|
||
<emphasis id="77869A47FFFEFF9EFDD3FE57FD64542C" box="[564,713,510,534]" italics="true" pageId="7" pageNumber="6">krulikovskyi</emphasis>
|
||
</taxonomicName>
|
||
is deemed to be the correct original spelling and is preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |