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<document id="B647875BA0CA450B131342DEE763D4DE" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes &amp; Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC345A4D6BBF1EA2FAF7F84A" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys vicina Frey 1969" docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="50" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="288598A0FA901AB5C04E3FFBEDD51966">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="2CEB9A7C16D02B143ECEAC6D66A60F15">Cherman, Mariana Alejandra</mods:namePart>
<mods:affiliation id="CBC81C32C7D1CBB47A98A3AA1D9CAB41">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="E2C34994D4A9BE9602120F2BD4E6C459">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="76DE18C7EAC26F1217C8218C04ED5CB6">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
<mods:nameIdentifier id="0E1E76DDD2DEB0D7A2E2FFA80701CC0B" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C75F23E7643ADCD849A9826144BB6D0D">Mise, Kleber Makoto</mods:namePart>
<mods:nameIdentifier id="51A1D1228A8DF88C590A60A07F716D22" type="ORCID">0000-0003-4391-9589</mods:nameIdentifier>
<mods:affiliation id="2758371E4059FA55C8B827F930C92239">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
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<mods:namePart id="0385A4E500801706DAA9BF9477E8ED3B">Frisch, Johannes</mods:namePart>
<mods:affiliation id="0FEF5141D79FEF2FE3C200F2CE62E9CD">Museum für Naturkunde Berlin, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="37F37761735A27781BECB663A37361F5">Almeida, Lúcia Massutti De</mods:namePart>
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<mods:affiliation id="CE15C3C57EA1DC5F77DDCA8D71D42B6A">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; lalmeida @ ufpr. br; https: // orcid. org / 0000 - 0003 - 4277 - 711 X</mods:affiliation>
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<treatment id="CD475E66EC345A4D6BBF1EA2FAF7F84A" ID-DOI="http://doi.org/10.5281/zenodo.4590948" ID-GBIF-Taxon="180301822" ID-Zenodo-Dep="4590948" LSID="urn:lsid:plazi:treatment:CD475E66EC345A4D6BBF1EA2FAF7F84A" httpUri="http://treatment.plazi.org/id/CD475E66EC345A4D6BBF1EA2FAF7F84A" lastPageNumber="50" pageId="49" pageNumber="50">
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<heading id="1E19581CEC345A4D6BBF1EA2FE74FCA9" bold="true" box="[151,478,762,789]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
<taxonomicName id="82EE94F3EC345A4D6BBF1EA2FE74FCA9" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[151,478,762,789]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="779A3362EC345A4D6BBF1EA2FE74FCA9" bold="true" box="[151,478,762,789]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BBF1EA2FEFFFCA8" bold="true" box="[151,341,763,789]" italics="true" pageId="49" pageNumber="50">Liogenys vicina</emphasis>
<bibRefCitation id="217F9281EC345A4D6A741EA2FE74FCA9" author="Frey, G." box="[348,478,762,789]" pageId="49" pageNumber="50" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</emphasis>
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<figureCitation id="DDD5F3F5EC345A4D6BBF1F46FF5EFC84" box="[151,244,799,825]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Figs. 22</figureCitation>
;
<figureCitation id="DDD5F3F5EC345A4D6A281F46FEB5FC84" box="[256,287,799,825]" captionStart="FIGURE 26" captionStartId="54.[151,250,1457,1481]" captionTargetBox="[171,1416,194,1421]" captionTargetId="figure-21@54.[151,1435,181,1433]" captionTargetPageId="54" captionText="FIGURE 26. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561330" httpUri="https://zenodo.org/record/4561330/files/figure.png" pageId="49" pageNumber="50">26</figureCitation>
.
</paragraph>
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<subSubSection id="0DF4BCFBEC345A4D6BBF1F3DFA30FC27" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="4551EF70EC345A4D6BBF1F3DFAB5FCC6" blockId="49.[151,1434,867,922]" box="[151,1311,867,892]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="65FEC85EEC345A4D6BBF1F3DFAB5FCC6" box="[151,1311,867,892]" pageId="49" pageNumber="50">
<treatmentCitation id="C44FC961EC345A4D6BBF1F3DFE77FCC6" author="Frey, G." box="[151,477,867,892]" page="62" pageId="49" pageNumber="50" year="1969">
<taxonomicName id="82EE94F3EC345A4D6BBF1F3DFE77FCC6" authority="Frey, 1969: 62" authorityName="Frey" authorityPageNumber="62" authorityYear="1969" box="[151,477,867,892]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="779A3362EC345A4D6BBF1F3DFEEBFCC6" box="[151,321,868,891]" italics="true" pageId="49" pageNumber="50">Liogenys vicinus</emphasis>
<bibRefCitation id="217F9281EC345A4D6A6F1F3DFE77FCC6" author="Frey, G." box="[327,477,867,892]" pageId="49" pageNumber="50" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969: 62</bibRefCitation>
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, 51 (key);
<treatmentCitation id="C44FC961EC345A4D69611F3AFCABFCC6" author="Krajcik, M." box="[585,769,867,891]" page="145" pageId="49" pageNumber="50" year="2012">
<bibRefCitation id="217F9281EC345A4D69611F3AFCABFCC6" author="Krajcik, M." box="[585,769,867,891]" pageId="49" pageNumber="50" refId="ref36405" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp." type="book" year="2012">Krajčík 2012: 145</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="C44FC961EC345A4D68A81F3AFBC3FCC6" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[896,1129,867,891]" page="5" pageId="49" pageNumber="50" year="2017">
<bibRefCitation id="217F9281EC345A4D68A81F3AFBC3FCC6" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[896,1129,867,891]" pageId="49" pageNumber="50" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC345A4D68CB1F3DFBBAFCC6" box="[995,1040,867,891]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2017: 5
</bibRefCitation>
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(generic history).
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</paragraph>
<paragraph id="4551EF70EC345A4D6BBF1FDBFA30FC27" blockId="49.[151,1434,867,922]" box="[151,1434,898,922]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="65FEC85EEC345A4D6BBF1FDBFA30FC27" box="[151,1434,898,922]" pageId="49" pageNumber="50">
<treatmentCitation id="C44FC961EC345A4D6BBF1FDBFE45FC27" author="Evans, A. V." box="[151,495,898,922]" page="215" pageId="49" pageNumber="50" year="2003">
<taxonomicName id="82EE94F3EC345A4D6BBF1FDBFE45FC27" authority=": Evans 2003: 215" authorityName=": Evans" authorityPageNumber="215" authorityYear="2003" box="[151,495,898,922]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="779A3362EC345A4D6BBF1FDBFE9DFC24" box="[151,311,898,921]" italics="true" pageId="49" pageNumber="50">Liogenys vicina</emphasis>
:
<bibRefCitation id="217F9281EC345A4D6A6C1FDBFE45FC27" author="Evans, A. V." box="[324,495,898,922]" pageId="49" pageNumber="50" pagination="1 - 458" refId="ref35927" refString="Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 215</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(checklist);
<treatmentCitation id="C44FC961EC345A4D69451FDBFCDEFC27" author="Evans, A. V. &amp; Smith, A. B. T." box="[621,884,898,922]" page="183" pageId="49" pageNumber="50" year="2009">
<bibRefCitation id="217F9281EC345A4D69451FDBFCDEFC27" author="Evans, A. V. &amp; Smith, A. B. T." box="[621,884,898,922]" pageId="49" pageNumber="50" refId="ref35974" refString="Evans, A. V. &amp; Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)" type="url" year="2009">Evans &amp; Smith 2009: 183</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="C44FC961EC345A4D68DB1FDBFB42FC27" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[1011,1256,898,922]" page="20" pageId="49" pageNumber="50" year="2017">
<bibRefCitation id="217F9281EC345A4D68DB1FDBFB42FC27" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[1011,1256,898,922]" pageId="49" pageNumber="50" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC345A4D6F7E1FDAFB23FC27" box="[1110,1161,898,922]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
2017: 20
</bibRefCitation>
</treatmentCitation>
(natural history).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BBF1F91FF41FA0B" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="4551EF70EC345A4D6BBF1F91FD3DFB63" blockId="49.[151,1437,967,2039]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BBF1F91FEEBFC5F" bold="true" box="[151,321,968,994]" pageId="49" pageNumber="50">Type material</emphasis>
.
<taxonomicName id="82EE94F3EC345A4D6A7A1F90FDBBFC5F" authorityName="Frey" authorityYear="1969" box="[338,529,969,994]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="779A3362EC345A4D6A7A1F90FDBBFC5F" box="[338,529,969,994]" italics="true" pageId="49" pageNumber="50">Liogenys vicinus</emphasis>
</taxonomicName>
<materialsCitation id="F586E52DEC345A4D69331F91FD3EFB63" ID-GBIF-Occurrence="3047558477" collectingDate="1949-12" collectionCode="USNM, NHMB" collectorName="P. Pilcomayo &amp; A. Martinez" country="Taiwan" determinerName="G Frey &amp; G. Frey" location="One" municipality="One" pageId="49" pageNumber="50" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
male
<typeStatus id="9A5551D2EC345A4D69741F91FD14FC5F" box="[604,702,968,994]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
(
<collectionCode id="23FF77B5EC345A4D69F81F91FC82FC5F" box="[720,808,968,994]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="49" pageNumber="50" type="Museum">USNM</collectionCode>
): [white handwritten] “Formosa / Clorinda / 8.XI.950. Daguerre”, [red, typeset and handwritten] “Type No / 74010 / USNM”, [red typeset] “TYPUS”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC345A4D6A1E1849FE0BFB97" box="[310,417,1040,1066]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ vicinus / n.sp. / Type / det.
<determinerName id="B08D03B4EC345A4D69F61849FC87FB96" box="[734,813,1040,1067]" pageId="49" pageNumber="50">G Frey</determinerName>
, 1968”, [pink typeset] “Property / USNM”, [white typeset] “Loan from / USNMNH / 2065060”, genitalia mounted.
<typeStatus id="9A5551D2EC345A4D686D186CFC18FBF2" box="[837,946,1077,1103]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
(2):
<location id="4031B9ABEC345A4D68C1186DFBB2FBF3" LSID="urn:lsid:plazi:treatment:CD475E66EC345A4D6BBF1EA2FAF7F84A:4031B9ABEC345A4D68C1186DFBB2FBF3" box="[1001,1048,1076,1102]" country="Taiwan" municipality="One" name="One" pageId="49" pageNumber="50">One</location>
female (
<collectionCode id="23FF77B5EC345A4D6F52186DFB7BFBF3" box="[1146,1233,1076,1102]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="49" pageNumber="50">NHMB</collectionCode>
) [white handwritten] “
<collectingCountry id="3DF9AFE0EC345A4D6BFC1801FE93FBCF" box="[212,313,1112,1138]" name="Taiwan" pageId="49" pageNumber="50">Formosa</collectingCountry>
/
<collectorName id="E81B8AA6EC345A4D6A781800FE42FBCF" box="[336,488,1112,1139]" pageId="49" pageNumber="50">P. Pilcomayo</collectorName>
”, [white typeset] “
<specimenCount id="53E824F9EC345A4D69931801FD64FBCF" box="[699,718,1112,1138]" pageId="49" pageNumber="50" type="female">♀</specimenCount>
”, [red typeset] “
<typeStatus id="9A5551D2EC345A4D68AF1801FBB1FBCF" box="[903,1051,1112,1138]" pageId="49" pageNumber="50" type="paratype">PARATYPE</typeStatus>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC345A4D6B891825FEA6FB2B" box="[161,268,1148,1174]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ vicinus / n.sp. / det.
<determinerName id="B08D03B4EC345A4D692A1825FDF2FB2A" box="[514,600,1148,1175]" pageId="49" pageNumber="50">G. Frey</determinerName>
, 1968”.
<collectingMunicipality id="A535750AEC345A4D699D1825FD4EFB2B" box="[693,740,1148,1174]" pageId="49" pageNumber="50">One</collectingMunicipality>
male (
<collectionCode id="23FF77B5EC345A4D68071825FC2CFB2B" box="[815,902,1148,1174]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="49" pageNumber="50">NHMB</collectionCode>
) [white typeset] “
<collectingCountry id="3DF9AFE0EC345A4D6F641825FB15FB2B" box="[1100,1215,1148,1174]" name="Argentina" pageId="49" pageNumber="50">Argentina</collectingCountry>
/ Formosa / Cindad
<date id="3150C9B0EC345A4D6BBF18F9FF57FB07" box="[151,253,1184,1210]" pageId="49" pageNumber="50" value="1949-12">
<collectingDate id="21143058EC345A4D6BBF18F9FF57FB07" box="[151,253,1184,1210]" pageId="49" pageNumber="50" value="1949-12">XII.1949</collectingDate>
</date>
/
<collectorName id="E81B8AA6EC345A4D6A3C18F9FE37FB07" box="[276,413,1184,1210]" pageId="49" pageNumber="50">A. Martinez</collectorName>
leg.”, [red typeset] “
<typeStatus id="9A5551D2EC345A4D69A318F9FCB5FB07" box="[651,799,1184,1210]" pageId="49" pageNumber="50" type="paratype">PARATYPE</typeStatus>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC345A4D6F9F18F9FA88FB07" box="[1207,1314,1184,1210]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ vicinus / n.sp. / det.
<determinerName id="B08D03B4EC345A4D6A3A189DFEC2FB62" box="[274,360,1220,1247]" pageId="49" pageNumber="50">G. Frey</determinerName>
, 1968”, genitalia mounted
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC345A4D6BEF18B1FF41FA0B" blockId="49.[151,1437,967,2039]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF18B1FE03FABF" bold="true" box="[199,425,1256,1282]" pageId="49" pageNumber="50">Non-type material</emphasis>
(9).
<materialsCitation id="F586E52DEC345A4D6AC418B1FAF7FABE" ID-GBIF-Occurrence="3047558366" box="[492,1373,1256,1283]" collectionCode="CEDEVA" collectorName="Laguna Yema &amp; Centro" country="Argentina" location="Formosa" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Formosa">
<collectingCountry id="3DF9AFE0EC345A4D6AC418B1FD39FABF" box="[492,659,1256,1282]" name="Argentina" pageId="49" pageNumber="50">ARGENTINA</collectingCountry>
.
<collectingRegion id="872A2192EC345A4D698D18B0FCA1FABE" box="[677,779,1257,1283]" country="Argentina" name="Formosa" pageId="49" pageNumber="50">Formosa</collectingRegion>
:
<collectorName id="E81B8AA6EC345A4D683618B0FC6EFABE" box="[798,964,1257,1283]" pageId="49" pageNumber="50">Laguna Yema</collectorName>
:
<collectorName id="E81B8AA6EC345A4D68FD18B1FB8BFABF" box="[981,1057,1256,1282]" pageId="49" pageNumber="50">Centro</collectorName>
de Validación-(
<collectionCode id="23FF77B5EC345A4D6FCB18B1FAFFFABE" box="[1251,1365,1256,1283]" pageId="49" pageNumber="50">CEDEVA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="F586E52DEC345A4D6E5818B1FEA4FAF7" ID-GBIF-Occurrence="3047558326" collectingDate="2008-12-14" collectionCode="IADIZA" collectorName="Ocampo &amp; San Blas &amp; Campon" country="Argentina" elevation="163" latitude="-24.2675" longLatPrecision="21" longitude="-61.248333" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Formosa">
barrio CEDEVA,
<geoCoordinate id="20DA89B7EC345A4D6A161955FE6EFA9B" box="[318,452,1292,1318]" degrees="24" direction="south" minutes="16" orientation="latitude" pageId="49" pageNumber="50" precision="15" seconds="03" value="-24.2675">24º16’03’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC345A4D6AE61955FDF5FA9A" box="[462,607,1292,1319]" degrees="61" direction="west" minutes="14" orientation="longitude" pageId="49" pageNumber="50" precision="15" seconds="54" value="-61.248333">61º14’54’’W</geoCoordinate>
,
<quantity id="82164295EC345A4D69451955FD12FA9A" box="[621,696,1292,1319]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="49" pageNumber="50" unit="m" value="163.0">
<elevation id="CEC30843EC345A4D69451955FD12FA9A" box="[621,696,1292,1319]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="49" pageNumber="50" unit="m" value="163.0">163 m</elevation>
</quantity>
,
<date id="3150C9B0EC345A4D69EF1955FCF8FA9B" box="[711,850,1292,1318]" pageId="49" pageNumber="50" value="2008-12-14">
<collectingDate id="21143058EC345A4D69EF1955FCF8FA9B" box="[711,850,1292,1318]" pageId="49" pageNumber="50" value="2008-12-14">14.XII.2008</collectingDate>
</date>
, F.C.
<collectorName id="E81B8AA6EC345A4D68B01955FC50FA9B" box="[920,1018,1292,1318]" pageId="49" pageNumber="50">Ocampo</collectorName>
, G.
<collectorName id="E81B8AA6EC345A4D6F041955FB39FA9B" box="[1068,1171,1292,1318]" pageId="49" pageNumber="50">San Blas</collectorName>
, &amp; F.
<collectorName id="E81B8AA6EC345A4D6FF71955FAEBFA9B" box="[1247,1345,1292,1318]" pageId="49" pageNumber="50">Campon</collectorName>
,
<specimenCount id="53E824F9EC345A4D6E661955FA36FA9B" box="[1358,1436,1292,1318]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC345A4D6BB71969FEACFAF7" box="[159,262,1328,1354]" pageId="49" pageNumber="50">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC345A4D6A341968FD0AFAD3" ID-GBIF-Occurrence="3047558451" collectingDate="2008-12-11" collectionCode="IADIZA" country="Argentina" latitude="-24.35582" location="Ruta Provincial" longLatPrecision="1" longitude="-61.315342" municipality="Reserva de Biosfera Teuquito" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Formosa">
<collectingMunicipality id="A535750AEC345A4D6A341968FDD8FAF7" box="[284,626,1328,1355]" pageId="49" pageNumber="50">Reserva de Biósfera Teuquito</collectingMunicipality>
,
<location id="4031B9ABEC345A4D69A81968FC9BFAF7" LSID="urn:lsid:plazi:treatment:CD475E66EC345A4D6BBF1EA2FAF7F84A:4031B9ABEC345A4D69A81968FC9BFAF7" box="[640,817,1328,1355]" country="Argentina" latitude="-24.35582" longLatPrecision="1" longitude="-61.315342" municipality="Reserva de Biosfera Teuquito" name="Ruta Provincial" pageId="49" pageNumber="50" stateProvince="Formosa">Ruta Provincial</location>
37, 1 km S “Canal”,
<geoCoordinate id="20DA89B7EC345A4D6F191969FB70FAF6" box="[1073,1242,1328,1355]" degrees="24" direction="south" minutes="21" orientation="latitude" pageId="49" pageNumber="50" precision="1" seconds="20,95" value="-24.35582">24º21’20,95’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC345A4D6FCC1969FA32FAF6" box="[1252,1432,1328,1355]" degrees="61" direction="west" minutes="18" orientation="longitude" pageId="49" pageNumber="50" precision="1" seconds="55,23" value="-61.315342">61º18’55,23’’W</geoCoordinate>
,
<date id="3150C9B0EC345A4D6BBF190DFE8BFAD3" box="[151,289,1364,1390]" pageId="49" pageNumber="50" value="2008-12-11">
<collectingDate id="21143058EC345A4D6BBF190DFE8BFAD3" box="[151,289,1364,1390]" pageId="49" pageNumber="50" value="2008-12-11">11.XII.2008</collectingDate>
</date>
,
<specimenCount id="53E824F9EC345A4D6A18190DFED4FAD3" box="[304,382,1364,1390]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
and
<specimenCount id="53E824F9EC345A4D6A93190DFDB5FAD3" box="[443,543,1364,1390]" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
(
<collectionCode id="23FF77B5EC345A4D6919190DFD32FAD3" box="[561,664,1364,1390]" pageId="49" pageNumber="50">IADIZA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC345A4D6986190DFE09FA2F" ID-GBIF-Occurrence="3047558357" collectingDate="1978-11" collectionCode="CMNC" collectorName="A. Martinez &amp; Fritz" country="Argentina" location="Fuerte Esperanza" pageId="49" pageNumber="50" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chaco">
<collectingRegion id="872A2192EC345A4D6986190DFD50FAD3" box="[686,762,1364,1390]" country="Argentina" name="Chaco" pageId="49" pageNumber="50">Chaco</collectingRegion>
:
<location id="4031B9ABEC345A4D682F190CFC65FAD2" LSID="urn:lsid:plazi:treatment:CD475E66EC345A4D6BBF1EA2FAF7F84A:4031B9ABEC345A4D682F190CFC65FAD2" box="[775,975,1365,1391]" country="Argentina" name="Fuerte Esperanza" pageId="49" pageNumber="50" stateProvince="Chaco">Fuerte Esperanza</location>
,
<date id="3150C9B0EC345A4D68F5190DFB96FAD3" box="[989,1084,1364,1390]" pageId="49" pageNumber="50" value="1978-11">
<collectingDate id="21143058EC345A4D68F5190DFB96FAD3" box="[989,1084,1364,1390]" pageId="49" pageNumber="50" value="1978-11">XI.1978</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC345A4D6F61190DFB79FAD3" box="[1097,1235,1364,1390]" pageId="49" pageNumber="50">A. Martínez</collectorName>
&amp;
<collectorName id="E81B8AA6EC345A4D6FD5190DFA9FFAD3" box="[1277,1333,1364,1390]" pageId="49" pageNumber="50">Fritz</collectorName>
,
<specimenCount id="53E824F9EC345A4D6E6B190DFA36FAD3" box="[1347,1436,1364,1390]" pageId="49" pageNumber="50" type="male">2 males</specimenCount>
and
<specimenCount id="53E824F9EC345A4D6BEF1921FE99FA2F" box="[199,307,1400,1426]" pageId="49" pageNumber="50" type="female">2 females</specimenCount>
(
<collectionCode id="23FF77B5EC345A4D6A6A1921FE31FA2F" box="[322,411,1400,1426]" pageId="49" pageNumber="50">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC345A4D6A871921FF4DFA0B" ID-GBIF-Occurrence="3047558390" collectingDate="1960-02" country="Argentina" location="Santiago del Estero" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Santiago del Estero">
<collectingRegion id="872A2192EC345A4D6A871921FD27FA2E" box="[431,653,1400,1427]" country="Argentina" name="Santiago del Estero" pageId="49" pageNumber="50">Santiago del Estero</collectingRegion>
: without locality,
<date id="3150C9B0EC345A4D687D1921FC03FA2F" box="[853,937,1400,1426]" pageId="49" pageNumber="50" value="1960-02">
<collectingDate id="21143058EC345A4D687D1921FC03FA2F" box="[853,937,1400,1426]" pageId="49" pageNumber="50" value="1960-02">II.1960</collectingDate>
</date>
, without collector,
<specimenCount id="53E824F9EC345A4D6FA91921FB66FA2F" box="[1153,1228,1400,1426]" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
and
<specimenCount id="53E824F9EC345A4D6E2B1921FACFFA2F" box="[1283,1381,1400,1426]" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
(IADIZA)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BEF1999FBD4F8FE" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="4551EF70EC345A4D6BEF1999FEBFF8A3" blockId="49.[151,1437,967,2039]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1999FE90FA67" bold="true" box="[199,314,1472,1498]" pageId="49" pageNumber="50">Diagnosis</emphasis>
. Length
<quantity id="82164295EC345A4D6A8A1999FDE3FA66" box="[418,585,1472,1499]" metricMagnitude="-2" metricUnit="m" metricValue="1.135" metricValueMax="1.19" metricValueMin="1.08" pageId="49" pageNumber="50" unit="mm" value="11.35" valueMax="11.9" valueMin="10.8">10.8–11.9 mm</quantity>
; width:
<quantity id="82164295EC345A4D698E1999FC9BFA66" box="[678,817,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.9" metricValueMin="4.8" pageId="49" pageNumber="50" unit="mm" value="5.35" valueMax="5.9" valueMin="4.8">4.8–5.9 mm</quantity>
. Body elongate, brown to dark brown, shiny; coarsely punctate (
<figureCitation id="DDD5F3F5EC345A4D6A2119BDFEDCFA43" box="[265,374,1508,1534]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Fig. 22A</figureCitation>
); clypeal emargination deep but not reaching the base of the teeth, rounded and narrowed; clypeal lateral margin strongly convex and produced, forming a tooth-like projection; pronotal posterior corners subangulate to rounded (
<figureCitation id="DDD5F3F5EC345A4D6A851A75FDBEF9FB" box="[429,532,1580,1606]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Fig. 22B</figureCitation>
); antennae with 10 antennomeres; first two pair of elytral ridges equally defined and more noticeable than the other two; basal apophysis of metacoxa produced; metafemur with bristles, sometimes abundantly; metatibia not carinate along the inner margin; pygidium convex, wider than it is long; pygidial width wider than the distance between spiracles of propygidium, disc with bristles throughout, erect bristles (
<figureCitation id="DDD5F3F5EC345A4D6E231AC1FADEF90F" box="[1291,1396,1688,1714]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Fig. 22D</figureCitation>
); in males protarsomeres and mesotarsomeres strongly enlarged, metatarsomeres slightly enlarged; parameres strongly expanded at the midline and abruptly narrowed at the apex (
<figureCitation id="DDD5F3F5EC345A4D681B1AB9FC1FF946" box="[819,949,1760,1787]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Fig. 22E, F</figureCitation>
); parameres strongly convex in lateral view (
<figureCitation id="DDD5F3F5EC345A4D6B881B5DFEA3F8A3" box="[160,265,1796,1822]" captionStart="FIGURE 22" captionStartId="50.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@50.[203,1385,181,1920]" captionTargetPageId="50" captionText="FIGURE 22. Liogenys vicina Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, F) dorsolateral, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561322" httpUri="https://zenodo.org/record/4561322/files/figure.png" pageId="49" pageNumber="50">Fig. 22G</figureCitation>
).
</paragraph>
<paragraph id="4551EF70EC345A4D6BEF1B71FBD4F8FE" blockId="49.[151,1437,967,2039]" box="[199,1150,1832,1859]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1B71FE54F8FF" bold="true" box="[199,510,1832,1858]" pageId="49" pageNumber="50">Geographical distribution</emphasis>
.
<materialsCitation id="F586E52DEC345A4D69221B71FBD3F8FE" ID-GBIF-Occurrence="3047558328" box="[522,1145,1832,1859]" country="Argentina" location="Formosa" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Formosa">
<collectingCountry id="3DF9AFE0EC345A4D69221B71FD06F8FF" box="[522,684,1832,1858]" name="Argentina" pageId="49" pageNumber="50">ARGENTINA</collectingCountry>
(
<collectingRegion id="872A2192EC345A4D69921B71FC8BF8FF" box="[698,801,1832,1858]" country="Argentina" name="Formosa" pageId="49" pageNumber="50">Formosa</collectingRegion>
,
<emphasis id="779A3362EC345A4D68041B71FCD2F8FF" bold="true" box="[812,888,1832,1858]" pageId="49" pageNumber="50">
<collectingRegion id="872A2192EC345A4D68041B71FCD2F8FF" box="[812,888,1832,1858]" country="Argentina" name="Chaco" pageId="49" pageNumber="50">Chaco</collectingRegion>
</emphasis>
,
<emphasis id="779A3362EC345A4D68AE1B71FBC7F8FF" bold="true" box="[902,1133,1832,1858]" pageId="49" pageNumber="50">
<collectingRegion id="872A2192EC345A4D68AE1B71FBC7F8FF" box="[902,1133,1832,1858]" country="Argentina" name="Santiago del Estero" pageId="49" pageNumber="50">Santiago del Estero</collectingRegion>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BEF1B15FAF7F84A" pageId="49" pageNumber="50" type="discussion">
<paragraph id="4551EF70EC345A4D6BEF1B15FAF7F84A" blockId="49.[151,1437,967,2039]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1B15FE9EF8DB" bold="true" box="[199,308,1868,1894]" pageId="49" pageNumber="50">Remarks</emphasis>
.
<taxonomicName id="82EE94F3EC345A4D6A6B1B14FE5EF8DB" authorityName=": Evans" authorityYear="2003" box="[323,500,1869,1894]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="779A3362EC345A4D6A6B1B14FE5EF8DB" box="[323,500,1869,1894]" italics="true" pageId="49" pageNumber="50">Liogenys vicina</emphasis>
</taxonomicName>
resembles the sympatric
<taxonomicName id="82EE94F3EC345A4D683D1B14FC3DF8DB" authorityName="Moser" authorityYear="1918" box="[789,919,1868,1894]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="latitarsis">
<emphasis id="779A3362EC345A4D683D1B14FC3DF8DB" box="[789,919,1868,1894]" italics="true" pageId="49" pageNumber="50">L. latitarsis</emphasis>
</taxonomicName>
(
<figureCitation id="DDD5F3F5EC345A4D688F1B15FC51F8DB" box="[935,1019,1868,1894]" captionStart="FIGURE 10" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@24.[203,1385,181,1920]" captionTargetPageId="24" captionText="FIGURE 10. Liogenys latitarsis Moser, 1918 male lectotype views: A) dorsal, B) labels, C) head and pronotum, D) lateral, E) pygidium, and parameres in views: F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561288" httpUri="https://zenodo.org/record/4561288/files/figure.png" pageId="49" pageNumber="50">Fig. 10</figureCitation>
), and these similarities difficult their identification.
<taxonomicName id="82EE94F3EC345A4D6A151B28FE45F837" authorityName=": Evans" authorityYear="2003" box="[317,495,1905,1930]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="779A3362EC345A4D6A151B28FE45F837" box="[317,495,1905,1930]" italics="true" pageId="49" pageNumber="50">Liogenys vicina</emphasis>
</taxonomicName>
differs from
<taxonomicName id="82EE94F3EC345A4D69A21B28FCA4F837" authorityName="Moser" authorityYear="1918" box="[650,782,1904,1930]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="latitarsis">
<emphasis id="779A3362EC345A4D69A21B28FCA4F837" box="[650,782,1904,1930]" italics="true" pageId="49" pageNumber="50">L. latitarsis</emphasis>
</taxonomicName>
(in parenthesis) in the punctures on pronotum and elytra coarser; clypeal emargination slightly narrower, lateral margins of pronotum slightly rounder, elytral inner ridges evenly elevated (sometimes the second ridge less elevated), metafemur more with bristles; and in the shape of parameres inflated basally and abruptly narrowed at the apex (not inflated basally and apical constriction absent).
</paragraph>
</subSubSection>
</treatment>
</document>
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