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<document id="26A765ADABC5B24885ED115120A280CC" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes &amp; Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC2F5A506BBF19B6FE2FFB54" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys rectangula Frey 1969" docType="treatment" docVersion="7" lastPageNumber="45" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="43" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5F4B0CC48DB5CC94F7E31E4FA91CE8C0">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="439E4F586DAE37367C9200FBE73C538B">Cherman, Mariana Alejandra</mods:namePart>
<mods:affiliation id="CD08D33C9DAA528B019E2E994A626FD8">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="545CA22B76FDB6AF96FE6242E7933F19">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="D753C5B2D7301C8C50A5CC54480322DF">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
<mods:nameIdentifier id="674FC14A292A2F48322C62D8725E5744" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B5DDCE9F5BF4D4206226693E0A1D2FC2">Mise, Kleber Makoto</mods:namePart>
<mods:nameIdentifier id="28F9848C03D7A53FFB6B3334ADB2CE5E" type="ORCID">0000-0003-4391-9589</mods:nameIdentifier>
<mods:affiliation id="5441E14FCC4CE7E3EA8A1E2016A25C26">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
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<mods:namePart id="A0286C7524C001880A80B48F49DAD696">Frisch, Johannes</mods:namePart>
<mods:affiliation id="2A31847C5C5A6DB1C5C863ECD6DA3224">Museum für Naturkunde Berlin, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="1DBF3A77903FCF442EC84B50813600F4">Almeida, Lúcia Massutti De</mods:namePart>
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<mods:affiliation id="752547FE5187ACFCC4467CD3A5EB1751">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; lalmeida @ ufpr. br; https: // orcid. org / 0000 - 0003 - 4277 - 711 X</mods:affiliation>
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<mods:date id="431532D6D98B971808AC2C6DB3F61680">2021</mods:date>
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<heading id="1E19581CEC2F5A566BBF19B6FDB2F9B5" bold="true" box="[151,536,1518,1545]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<taxonomicName id="82EE94F3EC2F5A566BBF19B6FDB2F9B5" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[151,536,1518,1545]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC2F5A566BBF19B6FDB2F9B5" bold="true" box="[151,536,1518,1545]" pageId="42" pageNumber="43">
<emphasis id="779A3362EC2F5A566BBF19B6FE24F9B4" bold="true" box="[151,398,1518,1545]" italics="true" pageId="42" pageNumber="43">Liogenys rectangula</emphasis>
<bibRefCitation id="217F9281EC2F5A566ABD19B6FDB2F9B5" author="Frey, G." box="[405,536,1518,1545]" pageId="42" pageNumber="43" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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<paragraph id="4551EF70EC2F5A566BBF1A4AFEB6F990" blockId="42.[151,536,1518,1581]" box="[151,284,1554,1581]" pageId="42" pageNumber="43">
<figureCitation id="DDD5F3F5EC2F5A566BBF1A4AFF5EF990" box="[151,244,1555,1581]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="42" pageNumber="43">Figs. 19</figureCitation>
;
<figureCitation id="DDD5F3F5EC2F5A566A281A4BFEB6F990" box="[256,284,1554,1581]" captionStart="FIGURE 25" captionStartId="53.[151,250,1459,1483]" captionTargetBox="[171,1416,194,1421]" captionTargetId="figure-20@53.[151,1435,181,1434]" captionTargetPageId="53" captionText="FIGURE 25. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561328" httpUri="https://zenodo.org/record/4561328/files/figure.png" pageId="42" pageNumber="43">25</figureCitation>
</paragraph>
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<paragraph id="4551EF70EC2F5A566BBF1A01FA92F9D2" blockId="42.[151,1437,1623,1709]" box="[151,1336,1623,1648]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="65FEC85EEC2F5A566BBF1A01FA92F9D2" box="[151,1336,1623,1648]" pageId="42" pageNumber="43">
<treatmentCitation id="C44FC961EC2F5A566BBF1A01FDA1F9D2" author="Frey, G." box="[151,523,1623,1648]" page="47" pageId="42" pageNumber="43" year="1969">
<taxonomicName id="82EE94F3EC2F5A566BBF1A01FDA1F9D2" authority="Frey, 1969: 47" authorityName="Frey" authorityPageNumber="47" authorityYear="1969" box="[151,523,1623,1648]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rectangulus">
<emphasis id="779A3362EC2F5A566BBF1A01FEDAF9D2" box="[151,368,1623,1647]" italics="true" pageId="42" pageNumber="43">Liogenys rectangulus</emphasis>
<bibRefCitation id="217F9281EC2F5A566A5E1A01FDA1F9D2" author="Frey, G." box="[374,523,1623,1648]" pageId="42" pageNumber="43" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969: 47</bibRefCitation>
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, 59–60;
<treatmentCitation id="C44FC961EC2F5A56694A1A0EFCB1F9D2" author="Krajcik, M." box="[610,795,1623,1647]" page="145" pageId="42" pageNumber="43" year="2012">
<bibRefCitation id="217F9281EC2F5A56694A1A0EFCB1F9D2" author="Krajcik, M." box="[610,795,1623,1647]" pageId="42" pageNumber="43" refId="ref36405" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp." type="book" year="2012">Krajčík 2012: 145</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="C44FC961EC2F5A5668B11A0EFB28F9D2" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[921,1154,1623,1647]" page="5" pageId="42" pageNumber="43" year="2017">
<bibRefCitation id="217F9281EC2F5A5668B11A0EFB28F9D2" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[921,1154,1623,1647]" pageId="42" pageNumber="43" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC2F5A5668D41A01FB9AF9D2" box="[1020,1072,1623,1647]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2017: 5
</bibRefCitation>
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(generic history);
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</paragraph>
<paragraph id="4551EF70EC2F5A566BBF1A2FFEB7F910" blockId="42.[151,1437,1623,1709]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="65FEC85EEC2F5A566BBF1A2FFEB7F910" pageId="42" pageNumber="43">
<treatmentCitation id="C44FC961EC2F5A566BBF1A2FFDD9F933" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[151,627,1654,1678]" page="13" pageId="42" pageNumber="43" year="2017">
<taxonomicName id="82EE94F3EC2F5A566BBF1A2FFDD9F933" authority=": Cherman et al. 2017: 13" authorityName="Cherman" authorityPageNumber="13" authorityYear="2017" box="[151,627,1654,1678]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC2F5A566BBF1A2FFECDF933" box="[151,359,1654,1678]" italics="true" pageId="42" pageNumber="43">Liogenys rectangula</emphasis>
:
<bibRefCitation id="217F9281EC2F5A566A5E1A2FFDD9F933" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[374,627,1654,1678]" pageId="42" pageNumber="43" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC2F5A566AF21A2EFDA5F933" box="[474,527,1654,1678]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2017: 13
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 20, 75 (illustration; natural history, taxonomy); Cherman
<emphasis id="779A3362EC2F5A566FE61A2EFAA9F933" box="[1230,1283,1654,1678]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2019: 36 (taxonomy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC2F5A506BBF1A82FB56FED8" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="4551EF70EC2F5A506BBF1A82FEABFF44" blockId="42.[151,1437,1754,2033]" lastBlockId="44.[151,1437,150,1257]" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
<emphasis id="779A3362EC2F5A566BBF1A82FEEAF949" bold="true" box="[151,320,1754,1781]" pageId="42" pageNumber="43">Type material</emphasis>
.
<taxonomicName id="82EE94F3EC2F5A566A781A82FDE8F948" authorityName="Frey" authorityYear="1969" box="[336,578,1755,1781]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="rectangulus">
<emphasis id="779A3362EC2F5A566A781A82FDE8F948" box="[336,578,1755,1781]" italics="true" pageId="42" pageNumber="43">Liogenys rectangulus</emphasis>
</taxonomicName>
<materialsCitation id="F586E52DEC2F5A5069631A82FF57FF44" ID-GBIF-Occurrence="3047558396" collectingDate="1949-12" collectionCode="USNM, NHMB, MZSP" collectorName="D. Chaco-Exped. &amp; A. Martinez" country="Argentina / Formosa" determinerName="G. Frey" lastPageId="44" lastPageNumber="45" location="One" municipality="One" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Salta" typeStatus="holotype">
male
<typeStatus id="9A5551D2EC2F5A5669A31A82FD47F948" box="[651,749,1755,1781]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
(
<collectionCode id="23FF77B5EC2F5A5669D61A82FCFCF948" box="[766,854,1755,1781]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43" type="Museum">USNM</collectionCode>
): [white handwritten] “Formosa / P [Puerto] Pilcomayo / Argent. 1950”, [red typeset] “
<typeStatus id="9A5551D2EC2F5A56691C1AA6FD62F8A4" box="[564,712,1791,1817]" pageId="42" pageNumber="43" type="paratype">PARATYPE</typeStatus>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC2F5A566F701AA6FB69F8A4" box="[1112,1219,1791,1817]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
rectangu / lus n. sp / det.
<determinerName id="B08D03B4EC2F5A566BF01B7AFE85F880" box="[216,303,1827,1853]" pageId="42" pageNumber="43">G. Frey</determinerName>
, 1968”, [white, typeset in red] “Smithson., / Cartwr.”, [white handwritten] “ +1”, genitalia mounted.
<typeStatus id="9A5551D2EC2F5A566BBF1B1EFEAEF8DC" box="[151,260,1863,1889]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
(3):
<location id="4031B9ABEC2F5A566A681B1EFEC5F8DC" LSID="urn:lsid:plazi:treatment:CD475E66EC2F5A506BBF19B6FE2FFB54:4031B9ABEC2F5A566A681B1EFEC5F8DC" box="[320,367,1863,1889]" country="Argentina / Formosa" municipality="One" name="One" pageId="42" pageNumber="43" stateProvince="Salta">One</location>
male (
<collectionCode id="23FF77B5EC2F5A566AE91B1EFDB6F8DC" box="[449,540,1863,1889]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="42" pageNumber="43">NHMB</collectionCode>
): [white typeset] “San José Chiq. / X.26. Lind. [Lindner] /
<collectorName id="E81B8AA6EC2F5A566FF01B1EFA37F8DC" box="[1240,1437,1863,1889]" pageId="42" pageNumber="43">D. Chaco-Exped.</collectorName>
[Deutchen Chaco-Expedition]”, [red typeset] “TYPUS”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC2F5A566FAC1B32FB45F838" box="[1156,1263,1899,1925]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
rectangu / lus n sp / Type / det.
<determinerName id="B08D03B4EC2F5A566A691BD6FE3DF814" box="[321,407,1935,1961]" pageId="42" pageNumber="43">G. Frey</determinerName>
, 1968”, genitalia mounted.
<collectingMunicipality id="A535750AEC2F5A5669E31BD6FD50F814" box="[715,762,1935,1961]" pageId="42" pageNumber="43">One</collectingMunicipality>
female (
<collectionCode id="23FF77B5EC2F5A5668741BD6FC1DF814" box="[860,951,1935,1961]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="42" pageNumber="43">NHMB</collectionCode>
): [white handwritten] “
<collectingRegion id="872A2192EC2F5A566F9F1BD6FB5EF814" box="[1207,1268,1935,1961]" country="Argentina" name="Salta" pageId="42" pageNumber="43">Salta</collectingRegion>
, Argent. / Dep. Rivadavia”, [red typeset] “
<typeStatus id="9A5551D2EC2F5A566AE11BEAFDF7F870" box="[457,605,1971,1997]" pageId="42" pageNumber="43" type="paratype">PARATYPE</typeStatus>
”, [white typeset] “
<specimenCount id="53E824F9EC2F5A5668181BEAFCE9F870" box="[816,835,1971,1997]" pageId="42" pageNumber="43" type="female">♀</specimenCount>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC2F5A566FF41BEAFAEDF870" box="[1244,1351,1971,1997]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
rectangu- / lus n sp / det.
<determinerName id="B08D03B4EC2F5A566A4F1B8EFE16F84C" box="[359,444,2007,2033]" pageId="42" pageNumber="43">G. Frey</determinerName>
, 1968”, [white, typeset in red] “Smithson., / Cartwright”, [white typeset] “Museum Frey / Tutzing”. One male (
<collectionCode id="23FF77B5EC295A506AA81CCEFE7AFF0C" box="[384,464,151,177]" country="Brazil" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="44" pageNumber="45" type="Museum">MZSP</collectionCode>
): [white typeset] “
<collectingCountry id="3DF9AFE0EC295A5069BE1CCEFCD5FF0C" box="[662,895,151,177]" isEnumeration="true" pageId="44" pageNumber="45">Argentina / Formosa</collectingCountry>
/ Ciudad
<date id="3150C9B0EC295A5068C11CCEFBE5FF0C" box="[1001,1103,151,177]" pageId="44" pageNumber="45" value="1949-12">
<collectingDate id="21143058EC295A5068C11CCEFBE5FF0C" box="[1001,1103,151,177]" pageId="44" pageNumber="45" value="1949-12">XII.1949</collectingDate>
</date>
/
<collectorName id="E81B8AA6EC295A506F491CCEFB4DFF0C" box="[1121,1255,151,177]" pageId="44" pageNumber="45">A. Martinez</collectorName>
leg.”, [red typeset] “
<typeStatus id="9A5551D2EC295A506BE71CE2FEC9FF68" box="[207,355,187,213]" pageId="44" pageNumber="45" type="paratype">PARATYPE</typeStatus>
”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC295A5069D91CE2FCF6FF68" box="[753,860,187,213]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ rectangu- / lus n sp / det.
<determinerName id="B08D03B4EC295A506FA11CE2FB75FF68" box="[1161,1247,187,213]" pageId="44" pageNumber="45">G. Frey</determinerName>
, 1968”, genitalia mounted
</materialsCitation>
.
</paragraph>
<caption id="1191BFF8EC2E5A576BBF1BCEFD2EF869" ID-DOI="http://doi.org/10.5281/zenodo.4561316" ID-Zenodo-Dep="4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="43">
<paragraph id="4551EF70EC2E5A576BBF1BCEFD2EF869" blockId="43.[151,1437,1943,2004]" pageId="43" pageNumber="44">
<emphasis id="779A3362EC2E5A576BBF1BCEFE8AF812" bold="true" box="[151,288,1943,1967]" pageId="43" pageNumber="44">FIGURE 19.</emphasis>
<taxonomicName id="82EE94F3EC2E5A576A0E1BC1FDC2F80D" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[294,616,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC2E5A576A0E1BC1FE5FF80D" box="[294,501,1944,1968]" italics="true" pageId="43" pageNumber="44">Liogenys rectangula</emphasis>
<bibRefCitation id="217F9281EC2E5A576AD31BC1FDC2F80D" author="Frey, G." box="[507,616,1944,1968]" pageId="43" pageNumber="44" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral.
</paragraph>
</caption>
<paragraph id="4551EF70EC295A506BEF1D5AFB56FED8" blockId="44.[151,1437,150,1257]" pageId="44" pageNumber="45">
<emphasis id="779A3362EC295A506BEF1D5AFE08FEA1" bold="true" box="[199,418,258,285]" pageId="44" pageNumber="45">Non-type material</emphasis>
(8).
<materialsCitation id="F586E52DEC295A506AF21D5AFE60FEFC" ID-GBIF-Occurrence="3047558356" collectionCode="NHMB" country="Bolivia" latitude="-17.834612" location="Lindner" longLatPrecision="1" longitude="-60.750473" municipality="San Jose de Chiquitos" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz">
<collectingCountry id="3DF9AFE0EC295A506AF21D5AFDFBFEA0" box="[474,593,259,285]" name="Bolivia" pageId="44" pageNumber="45">BOLIVIA</collectingCountry>
.
<collectingRegion id="872A2192EC295A5069741D5AFD71FEA0" box="[604,731,259,285]" country="Bolivia" name="Santa Cruz" pageId="44" pageNumber="45">Santa Cruz</collectingRegion>
,
<collectingMunicipality id="A535750AEC295A5069CF1D5AFC4AFEA0" box="[743,992,259,285]" pageId="44" pageNumber="45">San José de Chiquitos</collectingMunicipality>
[
<geoCoordinate id="20DA89B7EC295A5068C71D5BFB3BFEA0" box="[1007,1169,258,285]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="04.6" value="-17.834612">17°50’04.6’’S</geoCoordinate>
,
<geoCoordinate id="20DA89B7EC295A506FB51D5BFAE1FEA0" box="[1181,1355,258,285]" degrees="60" direction="west" minutes="45" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="01.7" value="-60.750473">60°45’01.7’’W</geoCoordinate>
], X.26,
<location id="4031B9ABEC295A506BBF1D7EFF5BFEFC" LSID="urn:lsid:plazi:treatment:CD475E66EC2F5A506BBF19B6FE2FFB54:4031B9ABEC295A506BBF1D7EFF5BFEFC" box="[151,241,295,321]" country="Bolivia" latitude="-17.834612" longLatPrecision="1" longitude="-60.750473" municipality="San Jose de Chiquitos" name="Lindner" pageId="44" pageNumber="45" stateProvince="Santa Cruz">Lindner</location>
,
<specimenCount id="53E824F9EC295A506A2A1D7EFEF9FEFC" box="[258,339,295,321]" pageId="44" pageNumber="45" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC295A506A4F1D7EFE6BFEFC" box="[359,449,295,321]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="44" pageNumber="45">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F586E52DEC295A506AF01D7EFB53FED8" ID-GBIF-Occurrence="3047558312" collectingDate="2008-12-13" collectionCode="IADIZA" country="Argentina" county="Ingeniero Juarez" latitude="-24.090834" location="San Blas" longLatPrecision="21" longitude="-61.946945" municipality="Ocampo" pageId="44" pageNumber="45" specimenCount="7" specimenCount-female="3" specimenCount-male="4">
<collectingCountry id="3DF9AFE0EC295A506AF01D7EFDD5FEFC" box="[472,639,295,321]" name="Argentina" pageId="44" pageNumber="45">ARGENTINA</collectingCountry>
. Formosa:
<collectingCounty id="AC3097FCEC295A50682D1D7EFC6DFEFC" box="[773,967,295,321]" pageId="44" pageNumber="45">Ingeniero Juárez</collectingCounty>
,
<geoCoordinate id="20DA89B7EC295A5068F01D7FFBF4FEFC" box="[984,1118,294,321]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="44" pageNumber="45" precision="15" seconds="27" value="-24.090834">24º05’27’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC295A506F421D7FFB51FEFC" box="[1130,1275,294,321]" degrees="61" direction="west" minutes="56" orientation="longitude" pageId="44" pageNumber="45" precision="15" seconds="49" value="-61.946945">61º56’49’’W</geoCoordinate>
,
<date id="3150C9B0EC295A506E241D7EFA32FEFC" box="[1292,1432,295,321]" pageId="44" pageNumber="45" value="2008-12-13">
<collectingDate id="21143058EC295A506E241D7EFA32FEFC" box="[1292,1432,295,321]" pageId="44" pageNumber="45" value="2008-12-13">13.XII.2008</collectingDate>
</date>
,
<collectingMunicipality id="A535750AEC295A506BBF1D12FF53FED8" box="[151,249,331,357]" pageId="44" pageNumber="45">Ocampo</collectingMunicipality>
,
<location id="4031B9ABEC295A506A2C1D12FEC3FED8" LSID="urn:lsid:plazi:treatment:CD475E66EC2F5A506BBF19B6FE2FFB54:4031B9ABEC295A506A2C1D12FEC3FED8" box="[260,361,331,357]" country="Argentina" county="Ingeniero Juarez" latitude="-24.090834" longLatPrecision="21" longitude="-61.946945" municipality="Ocampo" name="San Blas" pageId="44" pageNumber="45">San Blas</location>
, &amp;
<location id="4031B9ABEC295A506ABA1D12FE5EFED8" LSID="urn:lsid:plazi:treatment:CD475E66EC2F5A506BBF19B6FE2FFB54:4031B9ABEC295A506ABA1D12FE5EFED8" box="[402,500,331,357]" country="Argentina" county="Ingeniero Juarez" latitude="-24.090834" longLatPrecision="21" longitude="-61.946945" municipality="Ocampo" name="Campon" pageId="44" pageNumber="45">Campón</location>
, mercury vapor &amp; ultraviolet light,
<specimenCount id="53E824F9EC295A5068AA1D12FC72FED8" box="[898,984,331,357]" pageId="44" pageNumber="45" type="male">4 males</specimenCount>
and
<specimenCount id="53E824F9EC295A506F271D12FBD1FED8" box="[1039,1147,331,357]" pageId="44" pageNumber="45" type="female">3 females</specimenCount>
(
<collectionCode id="23FF77B5EC295A506FA11D12FB5AFED8" box="[1161,1264,331,357]" pageId="44" pageNumber="45">IADIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC295A506BEF1D37FDE8FD70" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="4551EF70EC295A506BEF1D37FDE8FD70" blockId="44.[151,1437,150,1257]" pageId="44" pageNumber="45">
<emphasis id="779A3362EC295A506BEF1D37FE90FE35" bold="true" box="[199,314,366,392]" pageId="44" pageNumber="45">Diagnosis</emphasis>
. Length: 9.0–
<quantity id="82164295EC295A506AC91D36FDE1FE34" box="[481,587,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="44" pageNumber="45" unit="mm" value="10.2">10.2 mm</quantity>
; width:
<quantity id="82164295EC295A5069851D36FC91FE34" box="[685,827,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" pageId="44" pageNumber="45" unit="mm" value="4.45" valueMax="4.7" valueMin="4.2">4.2–4.7 mm</quantity>
. Body and pronotum reddish brown, elongate, sides almost parallel; elytra testaceous, glabrous, opaque, somewhat pruinose (
<figureCitation id="DDD5F3F5EC295A5068EA1DCAFB80FE10" box="[962,1066,403,429]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="44" pageNumber="45">Fig. 19A</figureCitation>
); clypeal emargination very shallow, rounded and wide, in males as wide as the distance between eyes; outer sides of anterior teeth following the clypeal lateral margin; clypeal lateral margin straight or concave; antennae with 10 antennomeres; pronotal posterior corners sharp, in obtuse angle (
<figureCitation id="DDD5F3F5EC295A5069021DA6FD3BFDA4" box="[554,657,511,537]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="44" pageNumber="45">Fig. 19B</figureCitation>
); mesotibia quadrate in cross section; two transverse carinae posteriorly, the apical incomplete or weakly complete; pygidium flat; subtrapezoidal; pygidial disc with short, thick, erect bristles throughout (
<figureCitation id="DDD5F3F5EC295A506A561E1EFE43FDDC" box="[382,489,583,609]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="44" pageNumber="45">Fig. 19D</figureCitation>
); in males, inner margin of metatibia strongly carinate straight towards apex; tarsi abundantly with bristles dorsally, protarsomere II as long as it is wide; parameres widened medially and abruptly narrowed sub-basally; parameral split slightly beyond the midline; apex spatulate (
<figureCitation id="DDD5F3F5EC295A506F691ED6FB02FD14" box="[1089,1192,655,681]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="44" pageNumber="45">Fig. 19E</figureCitation>
); parameres in lateral view straight, not coplanar (
<figureCitation id="DDD5F3F5EC295A506AF91EEAFD9FFD70" box="[465,565,691,717]" captionStart="FIGURE 19" captionStartId="43.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@43.[203,1385,181,1920]" captionTargetPageId="43" captionText="FIGURE 19. Liogenys rectangula Frey, 1969 male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561316" httpUri="https://zenodo.org/record/4561316/files/figure.png" pageId="44" pageNumber="45">Fig. 19F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC295A506BEF1E8EFB0EFCA8" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="4551EF70EC295A506BEF1E8EFCF7FD4C" blockId="44.[151,1437,150,1257]" box="[199,861,726,753]" pageId="44" pageNumber="45">
<materialsCitation id="F586E52DEC295A506BEF1E8EFCF0FD4C" ID-GBIF-Occurrence="3047558418" box="[199,858,726,753]" country="Argentina" location="Puerto Pilcomayo" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Formosa" typeStatus="holotype">
<emphasis id="779A3362EC295A506BEF1E8EFECAFD4D" bold="true" box="[199,352,726,753]" pageId="44" pageNumber="45">
<typeStatus id="9A5551D2EC295A506BEF1E8EFEA8FD4C" box="[199,258,727,753]" pageId="44" pageNumber="45">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="3DF9AFE0EC295A506A441E8EFDB8FD4C" box="[364,530,727,753]" name="Argentina" pageId="44" pageNumber="45">ARGENTINA</collectingCountry>
,
<collectingRegion id="872A2192EC295A5069361E8EFD29FD4C" box="[542,643,727,753]" country="Argentina" name="Formosa" pageId="44" pageNumber="45">Formosa</collectingRegion>
,
<location id="4031B9ABEC295A5069A61E8EFCF0FD4C" LSID="urn:lsid:plazi:treatment:CD475E66EC2F5A506BBF19B6FE2FFB54:4031B9ABEC295A5069A61E8EFCF0FD4C" box="[654,858,727,753]" country="Argentina" name="Puerto Pilcomayo" pageId="44" pageNumber="45" stateProvince="Formosa">Puerto Pilcomayo</location>
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC295A506BEF1EA3FB0EFCA8" blockId="44.[151,1437,150,1257]" box="[199,1188,762,789]" pageId="44" pageNumber="45">
<emphasis id="779A3362EC295A506BEF1EA3FE54FCA9" bold="true" box="[199,510,762,788]" pageId="44" pageNumber="45">Geographical distribution</emphasis>
.
<emphasis id="779A3362EC295A5069241EA3FC8DFCA9" bold="true" box="[524,807,762,788]" pageId="44" pageNumber="45">
<collectingCountry id="3DF9AFE0EC295A5069241EA3FD2FFCA9" box="[524,645,762,788]" name="Bolivia" pageId="44" pageNumber="45">BOLIVIA</collectingCountry>
(
<collectingRegion id="872A2192EC295A5069BC1EA3FCB5FCA9" box="[660,799,762,788]" country="Bolivia" name="Santa Cruz" pageId="44" pageNumber="45">Santa Cruz</collectingRegion>
)
</emphasis>
; ARGENTINA (
<emphasis id="779A3362EC295A5068CE1EA3FB89FCA9" bold="true" box="[998,1059,762,788]" pageId="44" pageNumber="45">Salta</emphasis>
, Formosa).
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC295A506BEF1F46FE2FFB54" pageId="44" pageNumber="45" type="discussion">
<paragraph id="4551EF70EC295A506BEF1F46FE2FFB54" blockId="44.[151,1437,150,1257]" pageId="44" pageNumber="45">
<emphasis id="779A3362EC295A506BEF1F46FE9EFC84" bold="true" box="[199,308,799,825]" pageId="44" pageNumber="45">Remarks</emphasis>
.
<taxonomicName id="82EE94F3EC295A506A6B1F46FD8DFC84" authorityName="Frey" authorityYear="1969" box="[323,551,799,825]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC295A506A6B1F46FD8DFC84" box="[323,551,799,825]" italics="true" pageId="44" pageNumber="45">Liogenys rectangula</emphasis>
</taxonomicName>
resembles
<taxonomicName id="82EE94F3EC295A50698E1F46FC3AFC84" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[678,912,799,825]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="779A3362EC295A50698E1F46FCBAFC85" box="[678,784,799,824]" italics="true" pageId="44" pageNumber="45">L. moseri</emphasis>
<bibRefCitation id="217F9281EC295A5068301F46FC3AFC84" author="Frey, G." box="[792,912,799,825]" pageId="44" pageNumber="45" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="217F9281EC295A5068B71F46FB20FC84" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[927,1162,799,825]" pageId="44" pageNumber="45" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC295A506F271F46FBE2FC84" box="[1039,1096,799,825]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2017
</bibRefCitation>
,
<bibRefCitation id="217F9281EC295A506FBD1F46FB71FC84" author="Cherman, M. A. &amp; Basilio, D. S. &amp; Mise, K. M. &amp; Almeida, L. M." box="[1173,1243,799,825]" pageId="44" pageNumber="45" pagination="1 - 69" refId="ref35724" refString="Cherman, M. A., Basilio, D. S., Mise, K. M. &amp; Almeida, L. M. (2019 a) Unraveling the puzzle of Liogenys biodiversity: fifteen new species, nomenclatural acts and new geographical records (Coleoptera: Scarabaeidae). Insect Systematics &amp; Evolution, 51, 1 - 69. https: // doi. org / 10.1163 / 1876312 X- 00001040" type="journal article" year="2019">2019a</bibRefCitation>
) in the body size and color, in the shape and vestiture of pygidium, and in the protarsi abundantly with bristles.
<taxonomicName id="82EE94F3EC295A506F901F1AFA36FCE0" authorityName="Frey" authorityYear="1969" box="[1208,1436,835,861]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC295A506F901F1AFA36FCE0" box="[1208,1436,835,861]" italics="true" pageId="44" pageNumber="45">Liogenys rectangula</emphasis>
</taxonomicName>
differs from
<taxonomicName id="82EE94F3EC295A506A0F1F3EFE38FC3D" authorityName="Frey" authorityYear="1969" box="[295,402,871,896]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="moseri">
<emphasis id="779A3362EC295A506A0F1F3EFE38FC3D" box="[295,402,871,896]" italics="true" pageId="44" pageNumber="45">L. moseri</emphasis>
</taxonomicName>
(in parenthesis) mainly in the clypeus laterally straight, without tooth-like projection (convex, with a sharp tooth-like projection) and in the elytra in general pruinose (shiny or slightly semiopaque in some specimens).
<bibRefCitation id="217F9281EC295A506A0E1FF6FE04FC77" author="Frey, G." box="[294,430,943,970]" pageId="44" pageNumber="45" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
described
<taxonomicName id="82EE94F3EC295A5069051FF6FD6CFC74" authorityName="Frey" authorityYear="1969" box="[557,710,943,969]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC295A5069051FF6FD6CFC74" box="[557,710,943,969]" italics="true" pageId="44" pageNumber="45">L. rectangula</emphasis>
</taxonomicName>
from a series collected in Pilcomayo,
<collectingCountry id="3DF9AFE0EC295A506FAF1FF6FB46FC74" box="[1159,1260,943,969]" name="Taiwan" pageId="44" pageNumber="45">Formosa</collectingCountry>
,
<collectingCountry id="3DF9AFE0EC295A506FD11FF6FAC6FC74" box="[1273,1388,943,969]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
; the
<typeStatus id="9A5551D2EC295A506BBF1F8AFF53FC50" box="[151,249,979,1005]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
was deposited at the USNM and the
<typeStatus id="9A5551D2EC295A5069BA1F8AFD57FC50" box="[658,765,979,1005]" pageId="44" pageNumber="45" type="paratype">paratypes</typeStatus>
(Frey did not state the number) at NHMB. At the USNM the specimen deposited by Frey bears a
<typeStatus id="9A5551D2EC295A50690F1FAEFD2DFBAC" box="[551,647,1015,1041]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
label instead. At the NHMB, specimens with type label found are from other localities but the one stated by
<bibRefCitation id="217F9281EC295A5069131842FD6FFB8B" author="Frey, G." box="[571,709,1051,1078]" pageId="44" pageNumber="45" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
: one specimen is from “San José Chiquitos” (
<collectingCountry id="3DF9AFE0EC295A506FE51842FA89FB8B" box="[1229,1315,1051,1078]" name="Bolivia" pageId="44" pageNumber="45">Bolivia</collectingCountry>
) and other is from “
<collectingRegion id="872A2192EC295A506BD21866FE9DFBE4" box="[250,311,1087,1113]" country="Argentina" name="Salta" pageId="44" pageNumber="45">Salta</collectingRegion>
, Dep. Rivadavia” (
<collectingCountry id="3DF9AFE0EC295A5069251866FD29FBE7" box="[525,643,1087,1114]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
), with
<typeStatus id="9A5551D2EC295A5069E71866FC9BFBE4" box="[719,817,1087,1113]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
and
<typeStatus id="9A5551D2EC295A5068401866FC62FBE4" box="[872,968,1087,1113]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
labels, respectively. As they are not from the type locality stated by the author, they are not considered types. There is another
<typeStatus id="9A5551D2EC295A506F7F183AFB1DFBC0" box="[1111,1207,1123,1149]" pageId="44" pageNumber="45" type="paratype">paratype</typeStatus>
specimen deposited at the MZSP, which is from
<collectingCountry id="3DF9AFE0EC295A506AF818DEFD98FB1C" box="[464,562,1159,1185]" name="Taiwan" pageId="44" pageNumber="45">Formosa</collectingCountry>
Ciudad (
<collectingCountry id="3DF9AFE0EC295A5069B018DEFCA4FB1F" box="[664,782,1159,1186]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
). As the author did not state the ocurrences from Chiquitos (
<collectingCountry id="3DF9AFE0EC295A506BB718F2FF5FFB7B" box="[159,245,1195,1222]" name="Bolivia" pageId="44" pageNumber="45">Bolivia</collectingCountry>
) and
<collectingRegion id="872A2192EC295A506A1A18F2FEC0FB78" box="[306,362,1195,1221]" country="Argentina" name="Salta" pageId="44" pageNumber="45">Salta</collectingRegion>
(
<collectingCountry id="3DF9AFE0EC295A506A5F18F2FE40FB7B" box="[375,490,1195,1222]" name="Argentina" pageId="44" pageNumber="45">Argentina</collectingCountry>
) in the original description, we consider them as new
<taxonomicName id="82EE94F3EC295A506F6A18F2FB7DFB78" authorityName="Frey" authorityYear="1969" box="[1090,1239,1195,1221]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="779A3362EC295A506F6A18F2FB7DFB78" box="[1090,1239,1195,1221]" italics="true" pageId="44" pageNumber="45">L. rectangula</emphasis>
</taxonomicName>
state and country records, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>
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