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<document id="516A4A250CAC072D8FD37885C07D04B7" ID-CLB-Dataset="7835" ID-DOI="10.11646/zootaxa.4938.1.1" ID-GBIF-Dataset="045b347a-4a5b-474f-b69f-1cd3fedf7678" ID-ISSN="1175-5326" ID-Zenodo-Dep="4561258" ID-ZooBank="074E95CD-E7C0-4098-93F0-70DE03CAD0D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614236395824" checkinUser="plazi" docAuthor="Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes &amp; Almeida, Lúcia Massutti De" docDate="2021" docId="CD475E66EC2A5A4D6BBF1E3CFDA8FD17" docLanguage="en" docName="zootaxa.4938.1.1.pdf" docOrigin="Zootaxa 4938 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys truncata Cherman 2021, new species" docType="treatment" docUuid="D09F18D9-F0DA-429B-A7DF-C70B253EAC08" docUuidSource="ZooBank" docVersion="7" lastPageNumber="50" masterDocId="317E261EEC055A7C6B281C59FFAAFFBD" masterDocTitle="Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="48" updateTime="1698931070950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2E8B3E1636128805B31D77C1256CF809">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="C37D62F2E7AA9950AC6012A113C853EB">Cherman, Mariana Alejandra</mods:namePart>
<mods:affiliation id="BA2ECA8874002918C8A2C945BFE46CD3">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="58B954F9B05B80CE3B1CF9B2DDD7DE17">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="ACD2F485322C24F5037494C09A3E6760">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
<mods:nameIdentifier id="6F72F498CB1DF2155E66CF7C3BE721A3" type="email">basilio.dsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="37470BECB1D3B129F6E69BB968CC8A99">Mise, Kleber Makoto</mods:namePart>
<mods:nameIdentifier id="60B9A2C8F3E10E50F24AA830068FCCD7" type="ORCID">0000-0003-4391-9589</mods:nameIdentifier>
<mods:affiliation id="EE40095FF4798BEEA813F4DD6B208BD4">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
<mods:nameIdentifier id="D8E8CC6BF29E77C828BCD0D7683E6664" type="email">klebermise@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="455F5FADD68A42710E8DC01C26C0A21F">Frisch, Johannes</mods:namePart>
<mods:affiliation id="3A091F605AC6EDC48F9E41E0D99B547F">Museum für Naturkunde Berlin, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:namePart id="B7D34F7E1909C4A45D80D45CEB22B428">Almeida, Lúcia Massutti De</mods:namePart>
<mods:nameIdentifier id="072AA7316B575594DFEA2F2057E3A2B4" type="ORCID">0000-0003-4277-711X</mods:nameIdentifier>
<mods:affiliation id="95ACDE6FDB9BFDA025F42728E3170948">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; lalmeida @ ufpr. br; https: // orcid. org / 0000 - 0003 - 4277 - 711 X</mods:affiliation>
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<mods:date id="DAB3D067012AF97C26F86B3DAA4F4BDC">2021</mods:date>
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<treatment id="CD475E66EC2A5A4D6BBF1E3CFDA8FD17" ID-DOI="http://doi.org/10.5281/zenodo.4561306" ID-GBIF-Taxon="180301825" ID-Zenodo-Dep="4561306" LSID="urn:lsid:zoobank.org:act:D09F18D9-F0DA-429B-A7DF-C70B253EAC08" httpUri="http://treatment.plazi.org/id/CD475E66EC2A5A4D6BBF1E3CFDA8FD17" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
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<heading id="1E19581CEC2A5A536BBF1E3CFD25FDC2" bold="true" box="[151,655,613,640]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
<emphasis id="779A3362EC2A5A536BBF1E3CFD25FDC2" bold="true" box="[151,655,613,640]" pageId="47" pageNumber="48">
<taxonomicName id="82EE94F3EC2A5A536BBF1E3CFE59FDC2" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[151,499,613,640]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="truncata" status="sp. nov.">
<emphasis id="779A3362EC2A5A536BBF1E3CFED9FD3D" bold="true" box="[151,371,613,640]" italics="true" pageId="47" pageNumber="48">Liogenys truncata</emphasis>
Cherman
</taxonomicName>
,
<taxonomicNameLabel id="6CA98E19EC2A5A536AD71E3CFD25FDC2" box="[511,655,613,639]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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<paragraph id="4551EF70EC2A5A536BBF1ED0FE89FD1E" blockId="47.[151,919,613,712]" box="[151,291,649,675]" pageId="47" pageNumber="48">
<figureCitation id="DDD5F3F5EC2A5A536BBF1ED0FF5EFD1E" box="[151,244,649,675]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Figs. 21</figureCitation>
;
<figureCitation id="DDD5F3F5EC2A5A536A281ED0FEB5FD1E" box="[256,287,649,675]" captionStart="FIGURE 27" captionStartId="55.[151,250,1459,1483]" captionTargetBox="[171,1416,192,1422]" captionTargetId="figure-20@55.[151,1435,181,1434]" captionTargetPageId="55" captionText="FIGURE 27. Geographical distribution of the Liogenys new species. Shaped points: new species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561332" httpUri="https://zenodo.org/record/4561332/files/figure.png" pageId="47" pageNumber="48">27</figureCitation>
.
</paragraph>
<paragraph id="4551EF70EC2A5A536BBF1EF4FC3DFD75" blockId="47.[151,919,613,712]" box="[151,919,685,712]" pageId="47" pageNumber="48">
http://zoobank.org/
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</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC2A5A536BBF1EA9FCDAFC03" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="4551EF70EC2A5A536BBF1EA9FEF3FC27" blockId="47.[151,1437,751,2039]" pageId="47" pageNumber="48">
<emphasis id="779A3362EC2A5A536BBF1EA9FE97FCB7" bold="true" box="[151,317,752,778]" pageId="47" pageNumber="48">Type material</emphasis>
.
<materialsCitation id="F586E52DEC2A5A536A611EA9FE6EFCEF" ID-GBIF-Occurrence="3047558423" collectingDate="1982-01-30" collectionCode="CMNC" country="Argentina" location="Cabra Corral Dam" pageId="47" pageNumber="48" specimenCount="2" stateProvince="Salta" typeStatus="holotype">
<typeStatus id="9A5551D2EC2A5A536A611EA9FE1BFCB7" box="[329,433,752,778]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
labeled (
<collectionCode id="23FF77B5EC2A5A53693E1EA9FDDAFCB7" box="[534,624,752,778]" pageId="47" pageNumber="48">CMNC</collectionCode>
): [white typeset] “
<collectingCountry id="3DF9AFE0EC2A5A53681D1EA9FC74FCB7" box="[821,990,752,778]" name="Argentina" pageId="47" pageNumber="48">ARGENTINA</collectingCountry>
, Prov.
<collectingRegion id="872A2192EC2A5A536F001EA9FBCAFCB7" box="[1064,1120,752,778]" country="Argentina" name="Salta" pageId="47" pageNumber="48">Salta</collectingRegion>
/
<quantity id="82164295EC2A5A536F5C1EA9FB01FCB7" box="[1140,1195,752,778]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="48" unit="km" value="3.0">3 km</quantity>
W
<location id="4031B9ABEC2A5A536FF81EA9FA36FCB6" LSID="urn:lsid:plazi:treatment:CD475E66EC2A5A4D6BBF1E3CFDA8FD17:4031B9ABEC2A5A536FF81EA9FA36FCB6" box="[1232,1436,752,779]" country="Argentina" name="Cabra Corral Dam" pageId="47" pageNumber="48" stateProvince="Salta">Cabra Corral Dam</location>
/
<date id="3150C9B0EC2A5A536B8E1F4DFEBBFC93" box="[166,273,788,814]" pageId="47" pageNumber="48" value="1982-01-30">
<collectingDate id="21143058EC2A5A536B8E1F4DFEBBFC93" box="[166,273,788,814]" pageId="47" pageNumber="48" value="1982-01-30">30.I.1982</collectingDate>
</date>
/ H &amp; A Howden”, [red, typeset and handwritten] “
<taxonomicName id="82EE94F3EC2A5A5368781F4DFBDDFC93" authority="TRUNCATA" authorityName="TRUNCATA" box="[848,1143,788,814]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">LIOGENYS TRUNCATA</taxonomicName>
/
<typeStatus id="9A5551D2EC2A5A536FA41F4DFA8AFC93" box="[1164,1312,788,814]" pageId="47" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
/ Cherman M. A.”, genitalia mounted
</materialsCitation>
.
<materialsCitation id="F586E52DEC2A5A536AF81F60FEFEFC27" ID-GBIF-Occurrence="3047558387" collectingDate="1958-12" collectionCode="CNCI" country="Argentina" location="Stgo. del Estero" municipality="Choya" pageId="47" pageNumber="48" specimenCount="2" typeStatus="paratype">
<typeStatus id="9A5551D2EC2A5A536AF81F60FD97FCEE" box="[464,573,825,851]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
(3), one male and two females (
<collectionCode id="23FF77B5EC2A5A5368991F61FC51FCEF" box="[945,1019,824,850]" country="Canada" name="Canadian National Collection Insects" pageId="47" pageNumber="48">CNCI</collectionCode>
): “
<collectingCountry id="3DF9AFE0EC2A5A536F301F61FB17FCEF" box="[1048,1213,824,850]" name="Argentina" pageId="47" pageNumber="48">ARGENTINA</collectingCountry>
/
<location id="4031B9ABEC2A5A536FFE1F61FA26FCEE" LSID="urn:lsid:plazi:treatment:CD475E66EC2A5A4D6BBF1E3CFDA8FD17:4031B9ABEC2A5A536FFE1F61FA26FCEE" box="[1238,1420,824,851]" country="Argentina" municipality="Choya" name="Stgo. del Estero" pageId="47" pageNumber="48">Stgo. del Estero</location>
/
<collectingMunicipality id="A535750AEC2A5A536BBF1F05FF4AFCCB" box="[151,224,860,886]" pageId="47" pageNumber="48">Choya</collectingMunicipality>
[strikethrough]
<date id="3150C9B0EC2A5A536AB51F05FDA1FCCA" box="[413,523,860,887]" pageId="47" pageNumber="48" value="1958-12">
<collectingDate id="21143058EC2A5A536AB51F05FDA1FCCA" box="[413,523,860,887]" pageId="47" pageNumber="48" value="1958-12">XII-1958</collectingDate>
</date>
”, [yellow, typeset and handwritten] “
<taxonomicName id="82EE94F3EC2A5A5368901F05FB49FCCB" authority="TRUNCATA" authorityName="TRUNCATA" box="[952,1251,860,886]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">LIOGENYS TRUNCATA</taxonomicName>
/
<typeStatus id="9A5551D2EC2A5A536FD71F05FA20FCCB" box="[1279,1418,860,886]" pageId="47" pageNumber="48" type="paratype">PARATYPE</typeStatus>
/ Cherman M. A.”
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC2A5A536BEF1FFDFCDAFC03" blockId="47.[151,1437,751,2039]" box="[199,880,932,959]" pageId="47" pageNumber="48">
<typeStatus id="9A5551D2EC2A5A536BEF1FFDFE85FC03" box="[199,303,932,958]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
deposited at CMNC.
<typeStatus id="9A5551D2EC2A5A53690C1FFCFD3BFC02" box="[548,657,933,959]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
deposited at CNCI.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC2A5A536BEF1F91FEB8FB2B" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="4551EF70EC2A5A536BEF1F91FEB8FB2B" blockId="47.[151,1437,751,2039]" pageId="47" pageNumber="48">
<emphasis id="779A3362EC2A5A536BEF1F91FE90FC5F" bold="true" box="[199,314,968,994]" pageId="47" pageNumber="48">Diagnosis</emphasis>
. Body elongate; yellowish brown, pronotum and elytra unicolored, elytra uniformly with bristles (
<figureCitation id="DDD5F3F5EC2A5A536B881FB5FEA3FBBB" box="[160,265,1004,1030]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21A</figureCitation>
); clypeal emargination very wide and shallow, almost truncate (
<figureCitation id="DDD5F3F5EC2A5A5368F51FB5FBEEFBBB" box="[989,1092,1004,1030]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21B</figureCitation>
); procoxal and mesotibial surface smooth, metatibial surface coarsely punctate; apical inner margin of metatibia with sparse, long bristles (
<figureCitation id="DDD5F3F5EC2A5A536E471849FF60FBF3" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21C</figureCitation>
); pygidium with bristles and as wide as it is long (
<figureCitation id="DDD5F3F5EC2A5A536820186DFCD8FBF3" box="[776,882,1076,1102]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21D</figureCitation>
); in males protarsomeres I to IV weakly enlarge; parameres narrowed subapically; apex flap shaped, outer margin angulate inferiorly, angle in a plane below the apex (
<figureCitation id="DDD5F3F5EC2A5A536B881825FEAFFB2B" box="[160,261,1148,1174]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC2A5A536BEF18F9FB02F84B" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="4551EF70EC2A5A536BEF18F9FB02F84B" blockId="47.[151,1437,751,2039]" pageId="47" pageNumber="48">
<emphasis id="779A3362EC2A5A536BEF18F9FE9EFB07" bold="true" box="[199,308,1184,1210]" pageId="47" pageNumber="48">
<typeStatus id="9A5551D2EC2A5A536BEF18F9FE9EFB07" box="[199,308,1184,1210]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="779A3362EC2A5A536A6C18F9FE2BFB07" bold="true" box="[324,385,1184,1210]" pageId="47" pageNumber="48">Male</emphasis>
. Length:
<quantity id="82164295EC2A5A536ADA18F8FDE7FB06" box="[498,589,1185,1211]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="47" pageNumber="48" unit="mm" value="7.7">7.7 mm</quantity>
; width:
<quantity id="82164295EC2A5A53698418F9FCAEFB06" box="[684,772,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="47" pageNumber="48" unit="mm" value="3.5">3.5 mm</quantity>
. Yellowish brown, head and margins of pronotum darker.
<emphasis id="779A3362EC2A5A536BBF189DFF77FB63" box="[151,221,1220,1246]" italics="true" pageId="47" pageNumber="48">Head:</emphasis>
distance between eyes almost three times the width of one eye; frons with bristles; anterior margin of clypeus apparently truncate; clypeal emargination very shallow, almost straight, as wide as the distance between eyes; lateral margin of clypeus straight, slightly convex near the canthus; distal maxillary palpomere, maximum width less than twice the apical width; fovea shallow, extending up to the transverse midline; antenna with 10 antennomeres, club equal in color and longer than the funicle.
<emphasis id="779A3362EC2A5A536959190DFD61FAD3" box="[625,715,1364,1390]" italics="true" pageId="47" pageNumber="48">Thorax:</emphasis>
anterior margin of pronotum slightly produced medially; pronotum anteriorly narrowed forming a depressed ring; pronotal disc glabrous, punctures coarse and very sparse; posterior corners subangulate, angle obtuse; hypomere with long bristles, mesepisternum and sides of metaventrite with thick and fine bristles; distance between mesocoxae and metacoxae slightly longer than the metacoxa; scutellum ogival, finely punctate throughout.
<emphasis id="779A3362EC2A5A53693719BDFDC4FA43" box="[543,622,1508,1534]" italics="true" pageId="47" pageNumber="48">Elytra:</emphasis>
shiny, uniformly with bristles, bristles longer at the margins of the elytra, unicolored with the pronotum; elytral suture unicolored with the elytron and not elevated; all elytral ridges weakly defined.
<emphasis id="779A3362EC2A5A536BD41A74FE90F9FB" box="[252,314,1581,1606]" italics="true" pageId="47" pageNumber="48">Legs:</emphasis>
procoxa with bristles, smooth; three protibial teeth, the apical the longest; the three teeth equally spaced; mesofemural surface almost glabrous, setae on margins; mesotibia cylindrical in cross section, surface smooth; two transverse carinae, the apical carina incomplete; metacoxa with sparse, thick bristles; basal apophysis of metacoxa produced; metatibia carinate along the inner margin towards the apex, apical inner surface sparsely setose, bristles long; metatibial surface coarsely punctate; two metatibial transverse carinae present posteriorly; metatibial apical spurs of different lengths; the larger equal in length to the diameter of the tibial apex; protarsomeres I to IV enlarged; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, superior tooth of a claw longer and equal in width to the inferior, distance between teeth shorter than the inferior tooth.
<emphasis id="779A3362EC2A5A536FEA1B71FA9FF8FF" box="[1218,1333,1832,1858]" italics="true" pageId="47" pageNumber="48">Abdomen:</emphasis>
ventrites and propygidium with bristles; pygidium slightly convex, subquadrate, as wide as it is long; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc with bristles throughout; pygidial apex quadrate.
<emphasis id="779A3362EC2A5A536BBF1BCCFEBBF813" box="[151,273,1941,1966]" italics="true" pageId="47" pageNumber="48">Parameres</emphasis>
: basal region slightly wider than both sections of the parameres at the transverse midline, parameral split at 2/3; inner margins convergent at the apex, narrowed subapically; apex flap shaped, outer margin angulate inferiorly, angle in a plane below the apex; parameres in lateral view slightly convex, coplanar (
<figureCitation id="DDD5F3F5EC2A5A536F121B85FB36F84B" box="[1082,1180,2012,2038]" captionStart="FIGURE 21" captionStartId="48.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@48.[203,1385,181,1920]" captionTargetPageId="48" captionText="FIGURE 21. Liogenys truncata Cherman, new species, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="47" pageNumber="48">Fig. 21F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1191BFF8EC355A4C6BBF1BCEFD52F869" ID-DOI="http://doi.org/10.5281/zenodo.4561320" ID-Zenodo-Dep="4561320" httpUri="https://zenodo.org/record/4561320/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="48">
<paragraph id="4551EF70EC355A4C6BBF1BCEFD52F869" blockId="48.[151,1436,1943,2004]" pageId="48" pageNumber="49">
<emphasis id="779A3362EC355A4C6BBF1BCEFEB3F812" bold="true" box="[151,281,1943,1967]" pageId="48" pageNumber="49">FIGURE 21</emphasis>
.
<taxonomicName id="82EE94F3EC355A4C6A0F1BC1FDEFF80D" authority="Cherman" authorityName="Cherman" authorityYear="2021" box="[295,581,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="truncata" status="sp. nov.">
<emphasis id="779A3362EC355A4C6A0F1BC1FE75F812" box="[295,479,1944,1967]" italics="true" pageId="48" pageNumber="49">Liogenys truncata</emphasis>
Cherman
</taxonomicName>
,
<emphasis id="779A3362EC355A4C69671BC1FD66F812" bold="true" box="[591,716,1943,1968]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="6CA98E19EC355A4C69671BC1FD66F812" box="[591,716,1943,1968]" pageId="48" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</emphasis>
, male holotype views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, and parameres in views: E) dorsal, and F) lateral.
</paragraph>
</caption>
<subSubSection id="0DF4BCFBEC345A4D6BEF1CCFFB9DFF44" pageId="49" pageNumber="50" type="description">
<paragraph id="4551EF70EC345A4D6BEF1CCFFB9DFF44" blockId="49.[151,1437,150,682]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1CCFFE92FF0D" bold="true" box="[199,312,150,176]" pageId="49" pageNumber="50">Variation</emphasis>
. Male
<typeStatus id="9A5551D2EC345A4D6AAF1CCEFE46FF0C" box="[391,492,151,177]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
: length:
<quantity id="82164295EC345A4D69651CCEFD0CFF0C" box="[589,678,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="49" pageNumber="50" unit="mm" value="8.2">8.2 mm</quantity>
; width:
<quantity id="82164295EC345A4D68291CCEFCF2FF0C" box="[769,856,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="49" pageNumber="50" unit="mm" value="4.2">4.2 mm</quantity>
. Equal to the
<typeStatus id="9A5551D2EC345A4D68DE1CCEFBF2FF0C" box="[1014,1112,151,177]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
except in the pronotal punctures finer and sparser and pygidial apex rounder. Female
<typeStatus id="9A5551D2EC345A4D68321CE2FC20FF68" box="[794,906,187,213]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
: length: 7.8–8.0 mm; width:
<quantity id="82164295EC345A4D6FE01CE2FAE4FF68" box="[1224,1358,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="49" pageNumber="50" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6–3.9 mm</quantity>
. Equal to the
<typeStatus id="9A5551D2EC345A4D6BF51C86FE95FF44" box="[221,319,223,249]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
except in the abdomen with bristles less abundant and more sparse.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BEF1D5AFCA6FED8" pageId="49" pageNumber="50" type="etymology">
<paragraph id="4551EF70EC345A4D6BEF1D5AFCA6FED8" blockId="49.[151,1437,150,682]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1D5AFEE2FEA0" bold="true" box="[199,328,259,285]" pageId="49" pageNumber="50">Etymology</emphasis>
. Adjective in the nominative singular. From Latin
<emphasis id="779A3362EC345A4D68BB1D5AFC57FEA0" box="[915,1021,259,285]" italics="true" pageId="49" pageNumber="50">truncâtus</emphasis>
, perfect passive participle of
<emphasis id="779A3362EC345A4D6E7B1D5AFA36FEA0" box="[1363,1436,259,285]" italics="true" pageId="49" pageNumber="50">truncô</emphasis>
(“maim, reduce to a trunk”), meaning that a structure is ending abruptly as if cut across at the base or tip. This term is referring to the clypeus truncate in
<taxonomicName id="82EE94F3EC345A4D69121D12FCAFFED9" authorityName="Cherman" authorityYear="2021" box="[570,773,331,356]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="779A3362EC345A4D69121D12FCAFFED9" box="[570,773,331,356]" italics="true" pageId="49" pageNumber="50">Liogenys truncata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BEF1D36FC47FE10" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="4551EF70EC345A4D6BEF1D36FB68FE34" blockId="49.[151,1437,150,682]" box="[199,1218,366,393]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1D36FECCFE35" bold="true" box="[199,358,366,393]" pageId="49" pageNumber="50">Type locality.</emphasis>
<materialsCitation id="F586E52DEC345A4D6A431D36FB14FE34" ID-GBIF-Occurrence="3047558332" box="[363,1214,366,393]" country="Argentina" latitude="-25.277111" location="Cabra Corral" longLatPrecision="1" longitude="-65.389" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Salta">
<collectingCountry id="3DF9AFE0EC345A4D6A431D36FDBBFE34" box="[363,529,367,393]" name="Argentina" pageId="49" pageNumber="50">ARGENTINA</collectingCountry>
,
<collectingRegion id="872A2192EC345A4D69341D36FDFDFE34" box="[540,599,367,393]" country="Argentina" name="Salta" pageId="49" pageNumber="50">Salta</collectingRegion>
,
<quantity id="82164295EC345A4D694A1D36FD31FE34" box="[610,667,367,393]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="km" value="3.0">3 km</quantity>
W
<location id="4031B9ABEC345A4D69EA1D36FCFEFE34" LSID="urn:lsid:plazi:treatment:CD475E66EC2A5A4D6BBF1E3CFDA8FD17:4031B9ABEC345A4D69EA1D36FCFEFE34" box="[706,852,367,393]" country="Argentina" latitude="-25.277111" longLatPrecision="1" longitude="-65.389" name="Cabra Corral" pageId="49" pageNumber="50" stateProvince="Salta">Cabra Corral</location>
[
<geoCoordinate id="20DA89B7EC345A4D684A1D37FBABFE34" box="[866,1025,366,393]" degrees="25" direction="south" minutes="16" orientation="latitude" pageId="49" pageNumber="50" precision="1" seconds="37.6" value="-25.277111">25°16’37.6’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC345A4D6F211D37FB1DFE34" box="[1033,1207,366,393]" degrees="65" direction="west" minutes="23" orientation="longitude" pageId="49" pageNumber="50" precision="1" seconds="20.4" value="-65.389">65°23’20.4’’W</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC345A4D6BEF1DCBFC47FE10" blockId="49.[151,1437,150,682]" box="[199,1005,402,429]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1DCBFE54FE11" bold="true" box="[199,510,402,428]" pageId="49" pageNumber="50">Geographical distribution</emphasis>
.
<materialsCitation id="F586E52DEC345A4D69221DCAFC43FE10" ID-GBIF-Occurrence="3047558305" box="[522,1001,403,429]" country="Argentina" location="Salta" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Salta">
<collectingCountry id="3DF9AFE0EC345A4D69221DCAFD06FE10" box="[522,684,403,429]" name="Argentina" pageId="49" pageNumber="50">ARGENTINA</collectingCountry>
(
<collectingRegion id="872A2192EC345A4D69931DCAFD5DFE10" box="[699,759,403,429]" country="Argentina" name="Salta" pageId="49" pageNumber="50">Salta</collectingRegion>
,
<collectingRegion id="872A2192EC345A4D682A1DCAFC4BFE10" box="[770,993,403,429]" country="Argentina" name="Santiago del Estero" pageId="49" pageNumber="50">Santiago del Estero</collectingRegion>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC345A4D6BEF1DEEFDA8FD17" pageId="49" pageNumber="50" type="discussion">
<paragraph id="4551EF70EC345A4D6BEF1DEEFDA8FD17" blockId="49.[151,1437,150,682]" pageId="49" pageNumber="50">
<emphasis id="779A3362EC345A4D6BEF1DEEFE9EFE6C" bold="true" box="[199,308,439,465]" pageId="49" pageNumber="50">Remarks</emphasis>
.
<taxonomicName id="82EE94F3EC345A4D6A6C1DEEFDBBFE6D" authorityName="Cherman" authorityYear="2021" box="[324,529,439,464]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="779A3362EC345A4D6A6C1DEEFDBBFE6D" box="[324,529,439,464]" italics="true" pageId="49" pageNumber="50">Liogenys truncata</emphasis>
</taxonomicName>
resembles
<taxonomicName id="82EE94F3EC345A4D69BA1DEEFC05FE6C" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[658,943,439,465]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rufoflava">
<emphasis id="779A3362EC345A4D69BA1DEEFCB2FE6C" box="[658,792,439,465]" italics="true" pageId="49" pageNumber="50">L. rufoflava</emphasis>
<bibRefCitation id="217F9281EC345A4D68081DEEFC05FE6C" author="Moser, J." box="[800,943,439,465]" pageId="49" pageNumber="50" pagination="95 - 167" refId="ref36796" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">
Moser,
<quantity id="82164295EC345A4D685F1DEEFC05FE6C" box="[887,943,439,465]" metricMagnitude="1" metricUnit="m" metricValue="4.87172" pageId="49" pageNumber="50" unit="in" value="1918.0">1918</quantity>
</bibRefCitation>
</taxonomicName>
in the body color and the bristle pattern of pronotum, elytra, and pygidium.
<taxonomicName id="82EE94F3EC345A4D692D1D82FD65FE49" authorityName="Cherman" authorityYear="2021" box="[517,719,475,500]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="779A3362EC345A4D692D1D82FD65FE49" box="[517,719,475,500]" italics="true" pageId="49" pageNumber="50">Liogenys truncata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="82EE94F3EC345A4D68771D82FC49FE48" authorityName="Moser" authorityYear="1918" box="[863,995,475,501]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rufoflava">
<emphasis id="779A3362EC345A4D68771D82FC49FE48" box="[863,995,475,501]" italics="true" pageId="49" pageNumber="50">L. rufoflava</emphasis>
</taxonomicName>
(in parenthesis) mainly in the body size smaller, the clypeus almost truncate, not forming noticeable anterior teeth (two distinct teeth and a wide emargination between them), lateral margin of clypeus straight (convex, with a rounded projection); the pronotal punctures denser at the disc; the elytra entirely shiny (pruinose at the margins); the pygidium as long as it is wide (longer than it is wide); and in males the protarsi and mesotarsi slightly enlarged (distinctly enlarged); and the parameres longer, with apex spatulate (lanceolate).
</paragraph>
</subSubSection>
</treatment>
</document>
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