plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/7C/0D/87/7C0D87B7FFBAFFAD8987FE768C78F614.xml
maintenance 059f2cc034 added 7C
2024-06-21 12:41:26 +02:00

222 lines
26 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="5E88BFF464352FF7EC6E585D03BACB2B" ID-CLB-Dataset="45436" ID-DOI="10.5281/zenodo.211122" ID-GBIF-Dataset="344f8458-2c80-43b3-a96b-3aca02033f88" ID-ISSN="1175-5326" ID-Zenodo-Dep="211122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460439637460" checkinUser="plazi" docAuthor="Kohlmann, Bert &amp; Solís, Ángel" docDate="2012" docId="7C0D87B7FFBAFFAD8987FE768C78F614" docLanguage="en" docName="zt03193p052.pdf" docOrigin="Zootaxa 3193" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus turgidus Kohlmann &amp; Solis, new species" docType="treatment" docVersion="11" lastPageNumber="40" masterDocId="8034FFCFFFB0FFA18910FFC18D46F523" masterDocTitle="New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama" masterLastPageNumber="52" masterPageNumber="28" pageNumber="38" updateTime="1698332701623" updateUser="ExternalLinkService">
<mods:mods id="335FCEE3E70223808CF7C2C61CF5053D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A540AD99B52AAFAE860C1F84B9F04FCE">
<mods:title id="3FBCE5229508827A2AFA1B407516AEBD">New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama</mods:title>
</mods:titleInfo>
<mods:name id="C7EA9092AE8C271740532E97652AA7A4" type="personal">
<mods:role id="B42589867717213E3821FF7486A05D72">
<mods:roleTerm id="DE31D164AFA8163123D379827899B581">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5A4D1199FC34E94E54CC2A6B482216B">Kohlmann, Bert</mods:namePart>
</mods:name>
<mods:name id="59AFB180DAC45E811DD71CBB7EAD0F22" type="personal">
<mods:role id="0A997EA828764B78B6F4F9F08D60EF11">
<mods:roleTerm id="549200A29D331A08A6430596CB2D363B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EA7349C88DC4761BAF2A731A8A2C0AE">Solís, Ángel</mods:namePart>
</mods:name>
<mods:typeOfResource id="0A0C1965A761551AD885921A28867EB3">text</mods:typeOfResource>
<mods:relatedItem id="76F0129AE8C837E9FFD7480C31456598" type="host">
<mods:titleInfo id="959F7E2A39CC2C16EC9C115DB54A32FD">
<mods:title id="03A837E4F18D2CDBDAE1D1DAF34EAFFD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="563AAF3BEBF80E0A3D334D5D66EDC79C">
<mods:date id="7BFFE381E3C801A2C97DE3B30104F256">2012</mods:date>
<mods:detail id="922713325256253047C98765E6AA2EC1" type="volume">
<mods:number id="D15A2B26F69D8D20FB5EB76B44C9FD80">3193</mods:number>
</mods:detail>
<mods:extent id="38D30AA9281A265C53FB8FF0BDA19D1B" unit="page">
<mods:start id="86DE81E74570B0E6D2289A9D94BE1070">28</mods:start>
<mods:end id="DF648826E2090841AED3E317052F50BD">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3A5A87A8D106E8D3E3B4F8B31D066A94">journal article</mods:classification>
<mods:identifier id="552AC552CF3C63A3A736F279106C2B2D" type="CLB-Dataset">45436</mods:identifier>
<mods:identifier id="6B79DA5259233C104BDE3EFB35CD71D9" type="DOI">10.5281/zenodo.211122</mods:identifier>
<mods:identifier id="6CD9EE722CD57D69B86105B0C3EE2FA1" type="GBIF-Dataset">344f8458-2c80-43b3-a96b-3aca02033f88</mods:identifier>
<mods:identifier id="BE4DE1D1139CB07D5AF223B7F9F69D4B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7A6DD25D1A63B968F785A4690AF364B" type="Zenodo-Dep">211122</mods:identifier>
</mods:mods>
<treatment id="7C0D87B7FFBAFFAD8987FE768C78F614" ID-DOI="http://doi.org/10.5281/zenodo.6168316" ID-GBIF-Taxon="119507523" ID-Zenodo-Dep="6168316" LSID="urn:lsid:plazi:treatment:7C0D87B7FFBAFFAD8987FE768C78F614" httpUri="http://treatment.plazi.org/id/7C0D87B7FFBAFFAD8987FE768C78F614" lastPageId="12" lastPageNumber="40" pageId="10" pageNumber="38">
<subSubSection id="BCBE652AFFBAFFAB8987FE768C4BF4D1" pageId="10" pageNumber="38" type="nomenclature">
<paragraph id="F41B36A1FFBAFFAB8987FE768E72F4F2" blockId="10.[151,820,439,498]" box="[151,820,439,465]" pageId="10" pageNumber="38">
<heading id="AF5381CDFFBAFFAB8987FE768E72F4F2" bold="true" box="[151,820,439,465]" fontSize="11" level="1" pageId="10" pageNumber="38" reason="1">
<emphasis id="C6D0EAB3FFBAFFAB8987FE768E72F4F2" bold="true" box="[151,820,439,465]" pageId="10" pageNumber="38">
<taxonomicName id="33A44D22FFBAFFAB8987FE768FDEF4F2" ID-CoL="7XPF3" authority="Kohlmann &amp; Solis" authorityName="Kohlmann &amp; Solis" box="[151,664,439,465]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="C6D0EAB3FFBAFFAB8987FE768CEEF4F2" bold="true" box="[151,424,439,465]" italics="true" pageId="10" pageNumber="38">Onthophagus turgidus</emphasis>
Kohlmann &amp; Solís
</taxonomicName>
,
<taxonomicNameLabel id="DDE357C8FFBAFFAB8BB5FE768E72F4F2" box="[677,820,439,465]" pageId="10" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F41B36A1FFBAFFAB8987FE1B8C4BF4D1" blockId="10.[151,820,439,498]" box="[151,269,474,498]" pageId="10" pageNumber="38">
<figureCitation id="6C9F2A24FFBAFFAB8987FE1B8DA2F4D1" box="[151,228,474,498]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="10" pageNumber="38">Figs. 8</figureCitation>
,
<figureCitation id="6C9F2A24FFBAFFAB89E1FE1B8C4BF4D1" box="[241,269,474,498]" captionStart="FIGURE 14" captionStartId="21.[151,250,1866,1889]" captionTargetBox="[340,1246,1114,1842]" captionTargetId="figure@21.[340,1246,1114,1842]" captionTargetPageId="21" captionText="FIGURE 14. Known distribution of Ateuchus alutacius Kohlmann &amp; Solís, new species (circle) and Onthophagus turgidus Kohlmann &amp; Solís, new species (triangle). Base map from Google Maps." httpUri="https://zenodo.org/record/211135/files/figure.png" pageId="10" pageNumber="38">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFBAFFAB8987FDE38803F785" pageId="10" pageNumber="38" type="diagnosis">
<paragraph id="F41B36A1FFBAFFAB8987FDE38803F785" blockId="10.[151,1436,546,787]" pageId="10" pageNumber="38">
<emphasis id="C6D0EAB3FFBAFFAB8987FDE38C57F718" bold="true" box="[151,273,546,571]" pageId="10" pageNumber="38">Diagnosis.</emphasis>
This species is distinguished from other species of the
<taxonomicName id="33A44D22FFBAFFAB8A8CFDE2896AF718" box="[924,1068,547,571]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="dicranius">
<emphasis id="C6D0EAB3FFBAFFAB8A8CFDE28EF6F719" box="[924,944,547,570]" italics="true" pageId="10" pageNumber="38">O</emphasis>
.
<emphasis id="C6D0EAB3FFBAFFAB8AD2FDE2896AF718" box="[962,1068,547,571]" italics="true" pageId="10" pageNumber="38">dicranius</emphasis>
</taxonomicName>
species group by the following combination of characters: body dark reddish brown (
<figureCitation id="6C9F2A24FFBAFFAB8BFFFD878E74F77D" box="[751,818,582,606]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="10" pageNumber="38">Fig. 8</figureCitation>
); clypeal horn upright, basal portion wide, forked in its apical third, apical portion nearly parallel-sided; pronotum broadly tumescent anteriorly (
<figureCitation id="6C9F2A24FFBAFFAB8D96FDAB898CF7A1" box="[1158,1226,618,642]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="10" pageNumber="38">Fig. 8</figureCitation>
), apical portion of tumescence with well separated lateral tubercles; anterior pronotal apical bead angularly reflexed medially.
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFBAFFAA89D7FD738F3EF784" lastPageId="11" lastPageNumber="39" pageId="10" pageNumber="38" type="description">
<paragraph id="F41B36A1FFBAFFAB89D7FD738E24F631" blockId="10.[151,1436,546,787]" pageId="10" pageNumber="38">
<emphasis id="C6D0EAB3FFBAFFAB89D7FD738C9EF7E8" bold="true" box="[199,472,690,715]" pageId="10" pageNumber="38">
Description.
<typeStatus id="2B1F8803FFBAFFAB8874FD728C95F7E8" box="[356,467,691,715]" pageId="10" pageNumber="38" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="6C9F2A24FFBAFFAB8B22FD738F3DF7E9" box="[562,635,690,714]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="10" pageNumber="38">Fig. 8</figureCitation>
): Length 10.3 mm. Humeral width 5.9 mm. Body oblong and dark brown. Vertex, pronotum, and elytral intervals closely punctate; vertex and pronotal punctures ocellate, surface between smooth; most elytral punctures with very minute setae.
</paragraph>
<caption id="A0DB6629FFBAFFAB8987F800899EF2F4" box="[151,1240,1985,2008]" httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="10" pageNumber="38" targetBox="[414,1185,837,1970]" targetPageId="10">
<paragraph id="F41B36A1FFBAFFAB8987F800899EF2F4" blockId="10.[151,1240,1985,2008]" box="[151,1240,1985,2008]" pageId="10" pageNumber="38">
<emphasis id="C6D0EAB3FFBAFFAB8987F8008C54F2F5" bold="true" box="[151,274,1985,2008]" pageId="10" pageNumber="38">FIGURE 8.</emphasis>
Dorsal habitus of the male holotype of
<taxonomicName id="33A44D22FFBAFFAB8BB4F800890BF2F4" authority="Kohlmann &amp; Solis" authorityName="Kohlmann &amp; Solis" box="[676,1101,1985,2007]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="C6D0EAB3FFBAFFAB8BB4F8008EC1F2F4" box="[676,903,1985,2007]" italics="true" pageId="10" pageNumber="38">Onthophagus turgidus</emphasis>
Kohlmann &amp; Solís
</taxonomicName>
,
<taxonomicNameLabel id="DDE357C8FFBAFFAB8D48F8038992F2F4" box="[1112,1236,1985,2007]" pageId="10" pageNumber="38" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="F41B36A1FFBBFFAA89D7FF568EA8F41C" blockId="11.[151,1436,151,679]" pageId="11" pageNumber="39">
Clypeus at anterior median edge with an upright, slightly arched, and flattened horn (
<figureCitation id="6C9F2A24FFBBFFAA8D94FF56898EF58C" box="[1156,1224,151,175]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="11" pageNumber="39">Fig. 8</figureCitation>
); horn in its apical third bifurcate, Y-shaped, apical portion nearly parallel-sided (
<figureCitation id="6C9F2A24FFBBFFAA8A47FF7D8EDAF5F7" box="[855,924,188,212]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="11" pageNumber="39">Fig. 8</figureCitation>
). Side of clypeus from rounded edge of gena to base of horn almost straight. Clypeal surface behind the horn concave. Frons surface feebly convex medially and with scattered punctures. Vertex with very small tubercle on each side near anterior inner edge of each eye; surface anterior to and between tubercles punctate, punctures between eyes ocellate.
</paragraph>
<paragraph id="F41B36A1FFBBFFAA89D7FE8C8EABF4EC" blockId="11.[151,1436,151,679]" pageId="11" pageNumber="39">
Pronotum with anterior margin raised medially; anterior median half with large transverse tumosity (
<figureCitation id="6C9F2A24FFBBFFAA8C57FE8D88CAF447" box="[1351,1420,332,356]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="11" pageNumber="39">Fig. 8</figureCitation>
), delimited on each side by a distinct conical tubercle, tubercles evidently separated; anterior face of tumosity almost vertical with a small, slightly convex, tuberculated mid-line, tubercles very small. Pronotal surfaces near anterior lateral angles concave; marginal bead of posterior margin obsolete medially.
</paragraph>
<paragraph id="F41B36A1FFBBFFAA89D7FE1D8C16F71F" blockId="11.[151,1436,151,679]" pageId="11" pageNumber="39">Elytron with striae distinctly impressed with ocellate punctures at regular intervals; intervals on disc with two or more irregular rows of punctures, surface between shiny. Pygidium closely ocellate-punctate, each puncture with a short stiff seta.</paragraph>
<paragraph id="F41B36A1FFBBFFAA89D7FD898F3EF784" blockId="11.[151,1436,151,679]" pageId="11" pageNumber="39">Metasternum with ocellate punctures, except along mid-line. Protibia elongate, with terminal tuft of setae; apical and subapical teeth distinctly closer to each other than second to third or third to fourth teeth. Ventral surface of all femora with regular, shallow punctures.</paragraph>
</subSubSection>
<caption id="A0DB6629FFBBFFAA8987F8088EC0F2C3" box="[151,902,1993,2016]" pageId="11" pageNumber="39">
<paragraph id="F41B36A1FFBBFFAA8987F8088EC0F2C3" blockId="11.[151,902,1993,2016]" box="[151,902,1993,2016]" pageId="11" pageNumber="39">
<emphasis id="C6D0EAB3FFBBFFAA8987F8088C54F2FC" bold="true" box="[151,274,1993,2016]" pageId="11" pageNumber="39">FIGURE 9.</emphasis>
Dorsal habitus of a male
<taxonomicName id="33A44D22FFBBFFAA8B07F80B8EC0F2C3" authority="Howden &amp; Gill." authorityName="Howden &amp; Gill." box="[535,902,1994,2016]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="C6D0EAB3FFBBFFAA8B07F80B8F9DF2C3" box="[535,731,1994,2016]" italics="true" pageId="11" pageNumber="39">Onthophagus solisi</emphasis>
Howden &amp; Gill.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="BCBE652AFFBCFFAD89D7FF568833F7D3" pageId="12" pageNumber="40" type="materials_examined">
<paragraph id="F41B36A1FFBCFFAD89D7FF5689B3F5F7" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FF568CEAF593" bold="true" box="[199,428,151,176]" pageId="12" pageNumber="40">Examined material</emphasis>
(
<specimenCount id="E2A2FD28FFBCFFAD88AEFF568F7BF58C" box="[446,573,151,175]" pageId="12" pageNumber="40" type="generic">1 specimen</specimenCount>
).
<emphasis id="C6D0EAB3FFBCFFAD8B42FF598F80F593" bold="true" box="[594,710,152,176]" pageId="12" pageNumber="40">
<typeStatus id="2B1F8803FFBCFFAD8B42FF598F86F593" box="[594,704,152,176]" pageId="12" pageNumber="40" type="holotype">Holotype</typeStatus>
,
</emphasis>
male:
<collectingCountry id="8CB37631FFBCFFAD8A02FF598ECBF593" box="[786,909,152,176]" name="Panama" pageId="12" pageNumber="40">PANAMA</collectingCountry>
.
<collectingCountry id="8CB37631FFBCFFAD8A87FF568EB4F58C" box="[919,1010,151,175]" name="Panama" pageId="12" pageNumber="40">Panamá</collectingCountry>
, Bocas del Toro. Fila a 1.5 km este de río Tskui,
<specimenCount id="E2A2FD28FFBCFFAD8819FF7D8C11F5F7" box="[265,343,188,212]" pageId="12" pageNumber="40" type="generic">
<quantity id="335C9B44FFBCFFAD8819FF7D8C15F5F7" box="[265,339,188,212]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="40" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<geoCoordinate id="91905066FFBCFFAD884DFF7D8C88F5F7" box="[349,462,188,212]" direction="north" orientation="latitude" pageId="12" pageNumber="40" precision="5" value="9.4453">9.4453º N</geoCoordinate>
-
<geoCoordinate id="91905066FFBCFFAD88CDFF7D8F23F5F7" box="[477,613,188,212]" direction="west" orientation="longitude" pageId="12" pageNumber="40" precision="5" value="-82.8471">82.8471º W</geoCoordinate>
. Col: A. Solís y M. Moraga. Trampa 11. Proyecto Darwin.
</paragraph>
<paragraph id="F41B36A1FFBCFFAD89D7FF1E8FBDF43F" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FF1E8C6CF5DB" bold="true" box="[199,298,223,248]" pageId="12" pageNumber="40">Habitat.</emphasis>
The specimen was collected with a trap baited with pig manure at an altitude of
<quantity id="335C9B44FFBCFFAD8DD4FF1E884AF5D4" box="[1220,1292,223,247]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="40" unit="m" value="800.0">800 m</quantity>
inside a primary tropical rain forest, during the month of October.
</paragraph>
<paragraph id="F41B36A1FFBCFFAD89D7FEE68CE7F447" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FEE68F34F463" bold="true" box="[199,626,295,320]" pageId="12" pageNumber="40">
Geographical distribution (
<figureCitation id="6C9F2A24FFBCFFAD8B1EFEE68F22F463" box="[526,612,295,320]" captionStart="FIGURE 14" captionStartId="21.[151,250,1866,1889]" captionTargetBox="[340,1246,1114,1842]" captionTargetId="figure@21.[340,1246,1114,1842]" captionTargetPageId="21" captionText="FIGURE 14. Known distribution of Ateuchus alutacius Kohlmann &amp; Solís, new species (circle) and Onthophagus turgidus Kohlmann &amp; Solís, new species (triangle). Base map from Google Maps." httpUri="https://zenodo.org/record/211135/files/figure.png" pageId="12" pageNumber="40">Fig. 14</figureCitation>
).
</emphasis>
This species is known so far from the Caribbean slope on the Panamanian Central Cordillera.
</paragraph>
<paragraph id="F41B36A1FFBCFFAD89D7FEAE8F6FF4EC" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FEAE8C9CF4AB" bold="true" box="[199,474,367,392]" pageId="12" pageNumber="40">Chorological affinities.</emphasis>
<taxonomicName id="33A44D22FFBCFFAD88F3FEB18FA6F4AB" box="[483,736,368,392]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="turgidus">
<emphasis id="C6D0EAB3FFBCFFAD88F3FEB18FA6F4AB" box="[483,736,368,392]" italics="true" pageId="12" pageNumber="40">Onthophagus turgidus</emphasis>
</taxonomicName>
is found at similar altitudes in the Chiriquí Cordillera as
<taxonomicName id="33A44D22FFBCFFAD8C91FEB18D94F48F" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="C6D0EAB3FFBCFFAD8C91FEB18D94F48F" italics="true" pageId="12" pageNumber="40">O. solisi</emphasis>
</taxonomicName>
(
<quantity id="335C9B44FFBCFFAD89F4FE558C3EF48F" box="[228,376,404,428]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="12.5" metricValueMin="5.0" pageId="12" pageNumber="40" unit="m" value="875.0" valueMax="1250.0" valueMin="500.0">500–1250 m</quantity>
;
<bibRefCitation id="90354B50FFBCFFAD8897FE558FE3F48F" author="Kohlmann" box="[391,677,404,428]" pageId="12" pageNumber="40" refString="Kohlmann, B. &amp; Solis, A. (2001) El genero Onthophagus (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 49, 159 - 261." type="journal article" year="2001">Kohlmann &amp; Solís 2001</bibRefCitation>
;
<figureCitation id="6C9F2A24FFBCFFAD8BA4FE558FBCF48F" box="[692,762,404,428]" captionStart="FIGURE 9" captionStartId="11.[151,250,1993,2016]" captionText="FIGURE 9. Dorsal habitus of a male Onthophagus solisi Howden &amp; Gill." pageId="12" pageNumber="40">Fig. 9</figureCitation>
), its ecological equivalent (a small dung tunneller), in the Guanacaste and Tilarán Cordilleras.
</paragraph>
<paragraph id="F41B36A1FFBCFFAD89D7FE1A8C15F77C" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FE1A8CB2F4D7" bold="true" box="[199,500,475,500]" pageId="12" pageNumber="40">Taxonomic relationships.</emphasis>
More material is needed, especially females, in order to establish taxonomic relationships. Presently, and using the similar clypeal horn, the anterior pronotal marginal bead, and the pronotal tumescence, the new species would seem to be closely related to
<taxonomicName id="33A44D22FFBCFFAD8A62FDE489C1F71F" authority="Howden &amp; Gill" authorityName="Howden &amp; Gill" box="[882,1159,548,572]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="C6D0EAB3FFBCFFAD8A62FDE48E89F71F" box="[882,975,548,572]" italics="true" pageId="12" pageNumber="40">O. solisi</emphasis>
Howden &amp; Gill
</taxonomicName>
, and it might actually be its sister species.
</paragraph>
<paragraph id="F41B36A1FFBCFFAD89D7FDAD8833F7D3" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
This species will key to
<taxonomicName id="33A44D22FFBCFFAD88C3FDAC8F76F7A7" box="[467,560,620,644]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="C6D0EAB3FFBCFFAD88C3FDAC8F76F7A7" box="[467,560,620,644]" italics="true" pageId="12" pageNumber="40">O. solisi</emphasis>
</taxonomicName>
in Kohlmann &amp; Solís’ (2001) key. The male of the new species (
<figureCitation id="6C9F2A24FFBCFFAD8C12FDAD8802F7A7" box="[1282,1348,620,644]" captionStart="FIGURE 8" captionStartId="10.[151,250,1985,2008]" captionTargetBox="[414,1185,837,1970]" captionTargetId="figure@10.[408,1197,828,1985]" captionTargetPageId="10" captionText="FIGURE 8. Dorsal habitus of the male holotype of Onthophagus turgidus Kohlmann &amp; Solís, new species." httpUri="https://zenodo.org/record/211130/files/figure.png" pageId="12" pageNumber="40">Fig. 8</figureCitation>
) can be easily separated from the males of
<taxonomicName id="33A44D22FFBCFFAD8B3CFD518FCDF78B" box="[556,651,656,680]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="C6D0EAB3FFBCFFAD8B3CFD518FCDF78B" box="[556,651,656,680]" italics="true" pageId="12" pageNumber="40">O. solisi</emphasis>
</taxonomicName>
(
<figureCitation id="6C9F2A24FFBCFFAD8B8DFD4E8FA5F784" box="[669,739,655,679]" captionStart="FIGURE 9" captionStartId="11.[151,250,1993,2016]" captionText="FIGURE 9. Dorsal habitus of a male Onthophagus solisi Howden &amp; Gill." pageId="12" pageNumber="40">Fig. 9</figureCitation>
) by the form of the clypeal horn, which is thicker and more robust in the new species; as well as having a developed pronotal tumescence with more divergent lateral tubercles. With the description of these new species of
<taxonomicName id="33A44D22FFBCFFAD8B9CFD198E65F7D3" box="[652,803,728,752]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="C6D0EAB3FFBCFFAD8B9CFD198E65F7D3" box="[652,803,728,752]" italics="true" pageId="12" pageNumber="40">Onthophagus</emphasis>
</taxonomicName>
in this paper, there are now 24 known from
<collectingCountry id="8CB37631FFBCFFAD8C04FD188837F7D3" box="[1300,1393,729,752]" name="Panama" pageId="12" pageNumber="40">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BCBE652AFFBCFFAD89D7FD3D8C78F614" pageId="12" pageNumber="40" type="etymology">
<paragraph id="F41B36A1FFBCFFAD89D7FD3D8C78F614" blockId="12.[151,1437,151,823]" pageId="12" pageNumber="40">
<emphasis id="C6D0EAB3FFBCFFAD89D7FD3D8C08F637" bold="true" box="[199,334,764,788]" pageId="12" pageNumber="40">Etymology.</emphasis>
<taxonomicName id="33A44D22FFBCFFAD8844FD3D8CF1F637" box="[340,439,764,788]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="turgidus">
<emphasis id="C6D0EAB3FFBCFFAD8844FD3D8CF1F637" box="[340,439,764,788]" italics="true" pageId="12" pageNumber="40">Turgidus</emphasis>
</taxonomicName>
, Latin adjective in the genitive case meaning swollen, in reference to the transverse pronotal tumosity.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink