195 lines
18 KiB
XML
195 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6454048" ID-ISSN="2073-2333" ID-Zenodo-Dep="6454048" LSID="urn:lsid:zoobank.org:act:CDD03952-9506-4EC9-841A-C28C89AF6F56" approvalRequired="2" approvalRequired_for_textStreams="1" approvalRequired_for_treatments="1" checkinTime="1649781036213" checkinUser="felipe" docAuthor="Fedotova, Z. A. & Perkovsky, E. E." docDate="2017" docId="7C0D1A6DFFC8FFF7E6A6AAB4FCB8FC65" docLanguage="en" docName="VestnikZoologii.51.1.23-30.pdf" docOrigin="Vestnik Zoologii (Vestn. Zool.) 51 (1)" docSource="http://dx.doi.org/10.1515/vzoo-2017-0004" docStyle="DocumentStyle:9FE6554C034D0A4D8EA9CEAE255F4368.2:VestnikZoologii.2014-2019.journal_article" docStyleId="9FE6554C034D0A4D8EA9CEAE255F4368" docStyleName="VestnikZoologii.2014-2019.journal_article" docStyleVersion="2" docTitle="Rovnoholoneurus Fedotova et Perkovsky 2017, gen. n." docType="treatment" docUuid="CDD03952-9506-4EC9-841A-C28C89AF6F56" docUuidSource="ZooBank" docVersion="2" lastPageId="2" lastPageNumber="25" masterDocId="80346215FFC9FFF5E60BAF74FF86FFAF" masterDocTitle="New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine)" masterLastPageNumber="30" masterPageNumber="23" pageId="1" pageNumber="24" updateTime="1649781748146" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Genus And Species Of Gall Midges (Diptera, Cecidomyiidae: Porricondyinae, Holoneurini) From The Late Eocene Amber Of Olevsk (Zhitomir Region, Ukraine)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fedotova, Z. A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perkovsky, E. E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Vestnik Zoologii</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="series">
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<mods:title>Vestn. Zool.</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2017-02-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>51</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>30</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1515/vzoo-2017-0004</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1515/vzoo-2017-0004</mods:identifier>
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<mods:identifier type="ISSN">2073-2333</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6454042</mods:identifier>
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<mods:identifier type="ZooBank">43910383-B9E1-44AB-812A-51D406A9D4DA</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6454048" ID-Zenodo-Dep="6454048" LSID="urn:lsid:zoobank.org:act:CDD03952-9506-4EC9-841A-C28C89AF6F56" httpUri="http://treatment.plazi.org/id/7C0D1A6DFFC8FFF7E6A6AAB4FCB8FC65" lastPageId="2" lastPageNumber="25" pageId="1" pageNumber="24">
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<subSubSection pageId="1" pageNumber="24" type="nomenclature">
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<paragraph blockId="1.[173,837,1472,1500]" box="[173,837,1472,1500]" pageId="1" pageNumber="24">
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<heading box="[173,837,1472,1500]" fontSize="11" level="2" pageId="1" pageNumber="24" reason="4">
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<emphasis bold="true" box="[173,251,1472,1498]" pageId="1" pageNumber="24">Genus</emphasis>
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<taxonomicName authority="Fedotova et Perkovsky" authorityName="Fedotova et Perkovsky" authorityYear="2017" box="[258,742,1472,1500]" class="Insecta" family="Cecidomyiidae" genus="Rovnoholoneurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[258,470,1472,1499]" italics="true" pageId="1" pageNumber="24">Rovnoholoneurus</emphasis>
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Fedotova et Perkovsky
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</taxonomicName>
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,
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||
<emphasis bold="true" box="[755,837,1473,1499]" pageId="1" pageNumber="24">
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||
<taxonomicNameLabel box="[755,837,1473,1499]" pageId="1" pageNumber="24" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph blockId="1.[226,912,1519,1541]" box="[226,912,1519,1541]" pageId="1" pageNumber="24">
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urn:lsid:zoobank.org:act:
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="25" pageId="1" pageNumber="24" type="description">
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<paragraph blockId="1.[173,888,1562,1584]" box="[173,888,1562,1584]" pageId="1" pageNumber="24">
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T y p e s p e c i e s:
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<taxonomicName authority="Fedotova et Perkovsky" authorityName="Fedotova et Perkovsky" authorityYear="2017" box="[369,826,1562,1584]" class="Insecta" family="Cecidomyiidae" genus="Rovnoholoneurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="davidi" status="sp. n.">
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<emphasis box="[369,601,1562,1583]" italics="true" pageId="1" pageNumber="24">Rovnoholoneurus davidi</emphasis>
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Fedotova et Perkovsky
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</taxonomicName>
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,
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<emphasis bold="true" box="[834,888,1562,1583]" pageId="1" pageNumber="24">
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<taxonomicNameLabel box="[834,888,1562,1583]" pageId="1" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph blockId="1.[173,1232,1617,1836]" lastBlockId="2.[166,1225,175,970]" lastPageId="2" lastPageNumber="25" pageId="1" pageNumber="24">
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D e s c r i p t i o n (
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<figureCitation box="[449,550,1617,1644]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[163,196,1762,1784]" captionStartId-1="3.[174,207,1700,1722]" captionStartId-2="4.[166,199,936,958]" captionStartId-3="5.[174,207,1732,1754]" captionTargetBox-0="[178,1206,985,1724]" captionTargetBox-1="[173,1230,199,1671]" captionTargetBox-2="[170,1220,169,923]" captionTargetBox-3="[175,1232,212,1681]" captionTargetId-0="figure-372@2.[163,1226,985,1759]" captionTargetId-1="figure-10@3.[173,1231,198,1672]" captionTargetId-2="figure-380@4.[166,1225,155,926]" captionTargetId-3="figure-10@5.[174,1232,209,1683]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig. 1. Rovnoholoneurus davidi, male, holotype SIZK, OL-8: 1 — general appearance; 2 — head and antenna; 3 — flagellomeres 10–11; 4, flagellomere 5; 5 — mouth parts and palpi; 6 — the abdomen and part of the thorax with swollen scutellum. Scale bar 0.1 mm." captionText-1="Fig. 2. Rovnoholoneurus davidi, male, holotype SIZK, OL-8: 1 — general appearance (×20); 2 — head, scapus, pedicel and palpi (×127); 3 — flagellomeres 10–11 (×140); 4 — flagellomere 5 (×165); 5 — head and antenna (×57.4); 6 — head, palpi and notum with swollen scutellum (×100); 7 — veins of the base of the wing: C, R1+2, R, m+rm, rs, CuA1 (×80); 8 — veins CuA1, CuA2 and fold of wing (×109); 9 — abdominal tergites 4–8 (×60); 5 10 — palpi (×155); 11 — middle leg (×60)." captionText-2="Fig. 3. Rovnoholoneurus miyae, female, holotype SIZK, OL-2: 1 — general appearance; 2 — mouth parts and palpi; 3 — flagellomeres 8–10; 4 — flagellomeres 4–6; 5 — scapus, pedicel, flagellomeres 1 and 2; 6 — abdomen distally; 7 — apex of abdomen and ovipositor. Scale bar, 0.1 mm." captionText-3="Fig. 4. Rovnoholoneurus miyae, female, holotype SIZK, OL-2: 1 — head and antenna (×69); 2 — apex of abdomen and ovipositor (×360); 3 — fore tarsi (×39); 4 — general appearance (×30.6); 5 — head, palpi, scapus, pedicel, flagellomeres 1 and 2 (×90); 6 — distal portion of the abdomen (×90); 7 — flagellomeres 8–10 (×180); 8 — flagellomeres 4–6 (×190)." figureDoi-0="http://doi.org/10.5281/zenodo.6454044" figureDoi-1="http://doi.org/10.5281/zenodo.6454046" figureDoi-2="http://doi.org/10.5281/zenodo.6454050" figureDoi-3="http://doi.org/10.5281/zenodo.6454054" httpUri-0="https://zenodo.org/record/6454044/files/figure.png" httpUri-1="https://zenodo.org/record/6454046/files/figure.png" httpUri-2="https://zenodo.org/record/6454050/files/figure.png" httpUri-3="https://zenodo.org/record/6454054/files/figure.png" pageId="1" pageNumber="24">figs 1–4</figureCitation>
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). Body slightly shorter than wing. Head elongate; eyes fused, wide bridge positioned on 2/3 part of head, occiput wide, rounded. Head basally with very long setae. Male antenna 2+11-segmented; female antenna 2+10–11-segmented; scape slightly enlarged apically; flagellomeres narrow, basal enlargement with constriction near base or middle, neck prolonged (in male and female), necks of distal flagellomeres much shorter than those of medial ones. Basal enlargement with whorls of long setae of different sizes, especially with long setae in basal, medial whorls. 10th flagellomere sometimes fused, consisting of two segments. Proximal flagellomeres slightly longer than distal ones. Last flagellomere ovoid or conical, without apical projection. All female flagellomeres bearing two rings of sensorial filae with apical, basal round loops. Mouthparts strongly elongated, wide. Palpus 4-segmented, longer than head, with segments slender, almost parallel-sided, 4th segment longest. Scutellum strongly swollen. Wing longer than body, evenly and very strongly widened distally; vein
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<emphasis box="[527,553,336,368]" italics="true" pageId="2" pageNumber="25">
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R
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<subScript attach="left" box="[545,553,354,368]" fontSize="6" pageId="2" pageNumber="25">5</subScript>
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</emphasis>
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almost straight, slightly curved near apex, joining margin almost at wing tip;
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<emphasis box="[403,424,368,393]" italics="true" pageId="2" pageNumber="25">rs</emphasis>
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strong, short; vein
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<emphasis box="[663,738,368,393]" italics="true" pageId="2" pageNumber="25">m+rm</emphasis>
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slightly curved;
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<emphasis box="[940,982,368,400]" italics="true" pageId="2" pageNumber="25">
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Cu
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<subScript attach="left" box="[974,982,386,400]" fontSize="6" pageId="2" pageNumber="25">2</subScript>
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</emphasis>
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strongly developed, positioned far from margin of wing, but fork indistinct, not fused, with two poorly visible folds near
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<emphasis box="[288,350,432,464]" italics="true" pageId="2" pageNumber="25">
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CuA
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<subScript attach="left" box="[342,350,450,464]" fontSize="6" pageId="2" pageNumber="25">2</subScript>
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</emphasis>
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; vein
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<emphasis box="[418,469,432,458]" italics="true" pageId="2" pageNumber="25">CuP</emphasis>
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well developed. Legs with long setae ventrally. Abdomen slightly swollen near base, narrowed apically. Abdominal segments covered by very long setae. Ovipositor consisting of one pair of dorso-apical plates.
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</paragraph>
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<paragraph blockId="2.[166,1225,175,970]" pageId="2" pageNumber="25">
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C o m p a r i s o n. The new genus appears closely related to
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<taxonomicName authority="Kieffer, 1894" authorityName="Kieffer" authorityYear="1894" box="[930,1224,527,554]" class="Insecta" family="Cecidomyiidae" genus="Holoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis box="[930,1066,527,553]" italics="true" pageId="2" pageNumber="25">Holoneurus</emphasis>
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Kieffer, 1894
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</taxonomicName>
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(
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<bibRefCitation author="Mamaev, B. M." box="[174,350,559,586]" pageId="2" pageNumber="25" pagination="1 - 11" refId="ref4430" refString="Mamaev, B. M. 2001. The tribe Holoneurini in the Palearctic (Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, 17, 1 - 11 [In Russian]." type="journal article" year="2001">Mamaev, 2001</bibRefCitation>
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;
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<bibRefCitation author="Fedotova, Z. A. & Sidorenko, V. S." box="[359,674,559,586]" pageId="2" pageNumber="25" pagination="235 - 277" refId="ref4217" refString="Fedotova, Z. A., Sidorenko, V. S. 2007. New taxa of gall midges (Diptera, Cecidomyiidae, Porricondylinae) with reduced wing venation and flagellomeres from Russia. An International Journal of Dipterological Research, 18 (4), 235 - 277." type="journal article" year="2007">Fedotova, Sidorenko, 2007</bibRefCitation>
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), as evidenced by the body shape, 2+10–11-segmented antennae, long legs, and male and female flagellomere morphology. It differs from
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<taxonomicName authorityName="Kieffer" authorityYear="1894" box="[166,302,623,649]" class="Insecta" family="Cecidomyiidae" genus="Holoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis box="[166,302,623,649]" italics="true" pageId="2" pageNumber="25">Holoneurus</emphasis>
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</taxonomicName>
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in the following characters: wide eye bridge, wing venation, elongated and swollen mouthparts, palpi very thin and much longer than head, different shape of flagellomeres, vein
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<emphasis box="[329,355,688,720]" italics="true" pageId="2" pageNumber="25">
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R
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<subScript attach="left" box="[347,355,706,720]" fontSize="6" pageId="2" pageNumber="25">5</subScript>
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</emphasis>
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joining wing margin at apex,
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<emphasis box="[710,785,688,713]" italics="true" pageId="2" pageNumber="25">m+rm</emphasis>
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curved, vein
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<emphasis box="[942,963,688,713]" italics="true" pageId="2" pageNumber="25">rs</emphasis>
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very short,
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<emphasis box="[1101,1135,688,720]" italics="true" pageId="2" pageNumber="25">
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M
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<subScript attach="left" box="[1127,1135,706,720]" fontSize="6" pageId="2" pageNumber="25">3</subScript>
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</emphasis>
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absent,
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<emphasis box="[166,228,720,752]" italics="true" pageId="2" pageNumber="25">
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CuA
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<subScript attach="left" box="[220,228,738,752]" fontSize="6" pageId="2" pageNumber="25">2</subScript>
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</emphasis>
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well developed and reaching near wing margin, male hind femur longer than tibia, tarsi very long (almost as long as or longer than femur+tibia), long setae on ventral side of legs, and smaller body.
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</paragraph>
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<paragraph blockId="2.[166,1225,175,970]" pageId="2" pageNumber="25">
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S p e c i e s i n c l u d e d.
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<taxonomicName authority="Fedotova & Perkovsky, 2017" authorityName="Fedotova & Perkovsky" authorityYear="2017" box="[519,803,815,841]" class="Insecta" family="Cecidomyiidae" genus="Rovnoholoneurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="davidi" status="sp. n.">
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<emphasis box="[519,803,815,841]" italics="true" pageId="2" pageNumber="25">Rovnoholoneurus davidi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[811,876,815,842]" pageId="2" pageNumber="25" rank="species">
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<emphasis bold="true" box="[811,870,815,841]" pageId="2" pageNumber="25">sp. n</emphasis>
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.
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</taxonomicNameLabel>
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,
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<taxonomicName authority="Fedotova & Perkovsky, 2017" authorityName="Fedotova & Perkovsky" authorityYear="2017" box="[891,993,816,841]" class="Insecta" family="Cecidomyiidae" genus="Rovnoholoneurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="miyae" status="sp. n.">
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<emphasis box="[891,993,816,841]" italics="true" pageId="2" pageNumber="25">R. miyae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1001,1065,815,842]" pageId="2" pageNumber="25" rank="species">
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<emphasis bold="true" box="[1001,1060,815,841]" pageId="2" pageNumber="25">sp. n</emphasis>
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.
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</taxonomicNameLabel>
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,
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<taxonomicName authority="(Fedotova, 2005)" authorityName="Fedotova & Perkovsky" authorityYear="2017" baseAuthorityName="Fedotova" baseAuthorityYear="2005" class="Insecta" family="Cecidomyiidae" genus="Rovnoholoneurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="laqueatus" status="comb. n.">
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<emphasis box="[1080,1224,815,841]" italics="true" pageId="2" pageNumber="25">R. laqueatus</emphasis>
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(Fedotova, 2005)
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</taxonomicName>
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,
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<emphasis bold="true" box="[386,496,847,873]" pageId="2" pageNumber="25">
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<taxonomicNameLabel box="[386,496,847,873]" pageId="2" pageNumber="25" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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(for
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<taxonomicName authority="Fedotova" authorityName="Fedotova" authorityYear="2005" box="[561,919,847,874]" class="Insecta" family="Cecidomyiidae" genus="Bryocrypta" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="laqueata">
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<emphasis box="[561,800,847,873]" italics="true" pageId="2" pageNumber="25">Bryocrypta laqueata</emphasis>
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Fedotova
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</taxonomicName>
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in:
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<bibRefCitation author="Fedotova, Z. A. & Perkovsky, E. E." pageId="2" pageNumber="25" pagination="41 - 51" refId="ref3817" refString="Fedotova, Z. A., Perkovsky, E. E. 2005. New gall midges (Diptera, Cecidomyiidae) from the Rovno Amber: Subfamily Porricondylinae (Tribes Bryocryptini and Winnertziini) and subfamily Lasiopterinae (Tribes Brachineurini and Oligotrophini). Paleontologicheskii Zhurnal, 1, 42 - 53 [In Russian, English translation: Paleontological Journal, 39 (1), 41 - 51]." type="journal article" year="2005">Fedotova, Perkovsky, 2005</bibRefCitation>
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). All species are only known from
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<collectingRegion box="[632,710,879,906]" country="Ukraine" name="Rivne" pageId="2" pageNumber="25">Rovno</collectingRegion>
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amber.
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</paragraph>
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<paragraph blockId="2.[166,1225,175,970]" pageId="2" pageNumber="25">
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E t y m o l o g y. The generic name is formed from a combination of
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<collectingRegion box="[1047,1125,911,938]" country="Ukraine" name="Rivne" pageId="2" pageNumber="25">Rovno</collectingRegion>
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and the gall midge generic name
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<taxonomicName authorityName="Kieffer" authorityYear="1894" box="[459,595,943,969]" class="Insecta" family="Cecidomyiidae" genus="Holoneurus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis box="[459,595,943,969]" italics="true" pageId="2" pageNumber="25">Holoneurus</emphasis>
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</taxonomicName>
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. Gender masculine.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |