314 lines
36 KiB
XML
314 lines
36 KiB
XML
<document id="F531BBAD6319B5DA1E821BB1B629240C" ID-CLB-Dataset="45703" ID-DOI="10.5281/zenodo.211332" ID-GBIF-Dataset="7ee29426-108f-4393-ab96-28df5a11092b" ID-ISSN="1175-5326" ID-Zenodo-Dep="211332" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460424169765" checkinUser="plazi" docAuthor="Purchart, Luboš" docDate="2012" docId="732487F5BD486E7CEBEFA113FEECFBA1" docLanguage="en" docName="zt03153p068.pdf" docOrigin="Zootaxa 3153" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Deretus maderai Purchart, 2012, sp. nov." docType="treatment" docVersion="11" lastPageNumber="63" masterDocId="8F1DFF8DBD4D6E7AEB78A553FFF4FFF9" masterDocTitle="Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae)" masterLastPageNumber="68" masterPageNumber="57" pageNumber="62" updateTime="1698330379695" updateUser="ExternalLinkService">
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<mods:mods id="E04E6EA4CE1015B40894EAF1586E3B4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5F53298B347F6347CF81D8D98B2A96BB">
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<mods:title id="10FFE881D49B42379C5BFDBC53832BAD">Biodiversity research of darkling beetles on Socotra Island. Part I. The genus Deretus Gahan, 1900 (Coleoptera: Tenebrionidae)</mods:title>
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<mods:name id="D13EBFEFBB050C18CF465D9EE6620D48" type="personal">
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<mods:namePart id="8031E4316098E9FE88343EBA5FBA7252">Purchart, Luboš</mods:namePart>
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<mods:title id="C84229C64E2F5196DAC11CA770A31F66">Zootaxa</mods:title>
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<mods:part id="A470A092788AFBD30665D1674B4FEBC9">
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<mods:date id="A378CF80D6DB08521AC405614F1BD964">2012</mods:date>
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<mods:number id="77182A5F61EA51CA8DCA42DABD2CF48E">3153</mods:number>
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<mods:classification id="3FD93558A865F1074BC2BFF438A59A4A">journal article</mods:classification>
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<mods:identifier id="45B3319655DFE90A35CF00C801670477" type="CLB-Dataset">45703</mods:identifier>
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<mods:identifier id="8FD0728163127F2A0AE2C6DEB89409E6" type="DOI">10.5281/zenodo.211332</mods:identifier>
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<treatment id="732487F5BD486E7CEBEFA113FEECFBA1" ID-DOI="http://doi.org/10.5281/zenodo.6180206" ID-GBIF-Taxon="119428110" ID-Zenodo-Dep="6180206" LSID="urn:lsid:plazi:treatment:732487F5BD486E7CEBEFA113FEECFBA1" httpUri="http://treatment.plazi.org/id/732487F5BD486E7CEBEFA113FEECFBA1" lastPageId="6" lastPageNumber="63" pageId="5" pageNumber="62">
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<subSubSection id="B3976568BD486E7FEBEFA113FE71FB85" pageId="5" pageNumber="62" type="nomenclature">
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<paragraph id="FB3236E3BD486E7FEBEFA113FE43FBA1" blockId="5.[151,439,1087,1148]" box="[151,439,1087,1112]" pageId="5" pageNumber="62">
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<heading id="A07A818FBD486E7FEBEFA113FE43FBA1" bold="true" box="[151,439,1087,1112]" fontSize="10" level="2" pageId="5" pageNumber="62" reason="2">
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<emphasis id="C9F9EAF1BD486E7FEBEFA113FE43FBA1" bold="true" box="[151,439,1087,1112]" pageId="5" pageNumber="62">
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<taxonomicName id="3C8D4D60BD486E7FEBEFA113FEA3FBA1" box="[151,343,1087,1112]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
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<emphasis id="C9F9EAF1BD486E7FEBEFA113FEA3FBA1" bold="true" box="[151,343,1087,1112]" italics="true" pageId="5" pageNumber="62">Deretus maderai</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="D2CA578ABD486E7FEA26A113FE43FBA1" box="[350,439,1088,1112]" pageId="5" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="FB3236E3BD486E7FEBEFA137FE71FB85" blockId="5.[151,439,1087,1148]" box="[151,389,1124,1148]" pageId="5" pageNumber="62">
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(
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<figureCitation id="63B62A66BD486E7FEBE7A137FF0BFB85" box="[159,255,1124,1148]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1945,1968]" captionTargetBox="[161,1425,339,1921]" captionTargetId="figure@2.[161,1426,339,1921]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Habitus, dorsal view. 1. Deretus bezdeki sp. nov. (paratype), male; 2. D. bezdeki sp. nov. (paratype), female; 3. D. denticollis Gahan, 1900, male; 4. D. denticollis Gahan, 1900, female; 5. D. hulai sp. nov. (paratype), female; 6. D. maderai sp. nov. (holotype), male; 7. D. maderai sp. nov. (paratype), female." httpUri="https://zenodo.org/record/211333/files/figure.png" pageId="5" pageNumber="62">Fig. 6, 7</figureCitation>
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,
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<figureCitation id="63B62A66BD486E7FEA74A137FEDEFB85" box="[268,298,1124,1148]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 – 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="5" pageNumber="62">16</figureCitation>
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,
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<figureCitation id="63B62A66BD486E7FEA4DA137FEA7FB85" box="[309,339,1124,1148]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[220,1367,193,1738]" captionTargetId="figure@9.[220,1367,193,1739]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Aedeagus, dorsal view. 19. Deretus bezdeki sp. nov. (paratype), male; 20. D. denticollis Gahan, 1900, male; 21. D. maderai sp. nov. (holotype), male; 22. Deretus necopinatus sp. nov. (paratype), male; 23. D. spinicollis Schawaller, 2004, male." httpUri="https://zenodo.org/record/211335/files/figure.png" pageId="5" pageNumber="62">21</figureCitation>
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,
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<figureCitation id="63B62A66BD486E7FEA27A137FE88FB85" box="[351,380,1124,1148]" captionStart="FIGURES 24 – 28" captionStartId="9.[151,264,1937,1960]" captionText="FIGURES 24 – 28. Aedeagus, lateral view. 24. Deretus bezdeki sp. nov. (paratype), male; 25. D. denticollis Gahan, 1900, male; 26. D. maderai sp. nov. (holotype), male; 27. Deretus necopinatus sp. nov. (paratype), male; 28. D. spinicollis Schawaller, 2004, male." pageId="5" pageNumber="62">26</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B3976568BD486E7FEBEFA1FEFBD1FB3B" box="[151,1061,1195,1218]" pageId="5" pageNumber="62" type="reference_group">
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||
<paragraph id="FB3236E3BD486E7FEBEFA1FEFBD1FB3B" blockId="5.[151,1061,1195,1218]" box="[151,1061,1195,1218]" pageId="5" pageNumber="62">
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||
<treatmentCitationGroup id="DB9D11CDBD486E7FEBEFA1FEFBD1FB3B" box="[151,1061,1195,1218]" pageId="5" pageNumber="62">
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||
<taxonomicName id="3C8D4D60BD486E7FEBEFA1FEFDD0FB38" authority="Schawaller 2004: 457" authorityName="Schawaller" authorityPageNumber="457" authorityYear="2004" box="[151,548,1195,1218]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="wraniki">
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<emphasis id="C9F9EAF1BD486E7FEBEFA1FEFECDFB3B" box="[151,313,1196,1218]" italics="true" pageId="5" pageNumber="62">Deretus wraniki</emphasis>
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||
:
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<treatmentCitation id="7A2C10F2BD486E7FEA3FA1F8FDD0FB38" author="Schawaller" box="[327,548,1195,1217]" page="457" pageId="5" pageNumber="62" year="2004">
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||
<bibRefCitation id="9F1C4B12BD486E7FEA3FA1F8FE00FB38" author="Schawaller" box="[327,500,1195,1217]" pageId="5" pageNumber="62" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller 2004</bibRefCitation>
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||
: 457
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(with figure of habitus) (partim, female
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||
<typeStatus id="24368841BD486E7FE8C7A1FEFBEFFB3B" box="[959,1051,1197,1218]" pageId="5" pageNumber="62" type="paratype">paratype</typeStatus>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B3976568BD486E7FEBEFA1BCFF05F8BF" pageId="5" pageNumber="62" type="materials_examined">
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||
<paragraph id="FB3236E3BD486E7FEBEFA1BCFD58FAFF" blockId="5.[151,1437,1262,2007]" box="[151,684,1262,1287]" pageId="5" pageNumber="62">
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||
<emphasis id="C9F9EAF1BD486E7FEBEFA1BCFEC1FAFE" bold="true" box="[151,309,1262,1287]" pageId="5" pageNumber="62">
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||
<typeStatus id="24368841BD486E7FEBEFA1BCFF26FAFE" box="[151,210,1263,1287]" pageId="5" pageNumber="62">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<collectingCountry id="839A7673BD486E7FEA44A1BCFE79FAFF" box="[316,397,1263,1286]" name="Yemen" pageId="5" pageNumber="62">Yemen</collectingCountry>
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||
,
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||
<collectingCountry id="839A7673BD486E7FEAE0A1BDFE19FAFF" box="[408,493,1262,1286]" name="Yemen" pageId="5" pageNumber="62">Socotra</collectingCountry>
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Island, Shibhon.
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||
</paragraph>
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||
<paragraph id="FB3236E3BD486E7FEBBDA040FCB9FA27" blockId="5.[151,1437,1262,2007]" pageId="5" pageNumber="62">
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||
<emphasis id="C9F9EAF1BD486E7FEBBDA040FE8BFAD2" bold="true" box="[197,383,1298,1323]" pageId="5" pageNumber="62">
|
||
<typeStatus id="24368841BD486E7FEBBDA040FEF4FAD2" box="[197,256,1299,1323]" pageId="5" pageNumber="62">Type</typeStatus>
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||
material.
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||
</emphasis>
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||
<typeStatus id="24368841BD486E7FEAEAA040FE0DFAD2" box="[402,505,1299,1323]" pageId="5" pageNumber="62" type="holotype">
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||
<emphasis id="C9F9EAF1BD486E7FEAEAA040FE0DFAD2" box="[402,505,1299,1323]" italics="true" pageId="5" pageNumber="62">Holotype</emphasis>
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||
</typeStatus>
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||
—(13
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||
<collectionCode id="9D9CAE26BD486E7FE938A041FD61FAD3" box="[576,661,1298,1322]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="62">NMPC</collectionCode>
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||
):
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||
<collectingCountry id="839A7673BD486E7FE9CCA040FCF1FAD3" box="[692,773,1299,1322]" name="Yemen" pageId="5" pageNumber="62">Yemen</collectingCountry>
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||
,
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||
<collectingCountry id="839A7673BD486E7FE864A041FC85FAD3" box="[796,881,1298,1322]" name="Yemen" pageId="5" pageNumber="62">Socotra</collectingCountry>
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||
Island, Shibhon,
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||
<quantity id="3C759B06BD486E7FEF23A041FB5BFAD3" box="[1115,1199,1298,1322]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="5" pageNumber="62" unit="m" value="680.0">680 m</quantity>
|
||
,
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||
<geoCoordinate id="9EB95024BD486E7FEFBFA040FA87FAD3" box="[1223,1395,1298,1322]" direction="north" orientation="latitude" pageId="5" pageNumber="62" precision="1" value="12.467083">N 12°28´1.5´´</geoCoordinate>
|
||
,
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||
<geoCoordinate id="9EB95024BD486E7FEEF3A040FEDEFAB7" direction="east" orientation="longitude" pageId="5" pageNumber="62" precision="1" value="53.975388">E 53°58´31.4´´</geoCoordinate>
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||
,
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||
<date id="8F331023BD486E7FEA4EA065FE35FAB7" box="[310,449,1334,1358]" pageId="5" pageNumber="62" value="2009-06-13">13. vi. 2009</date>
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, L. Purchart lgt.
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||
<emphasis id="C9F9EAF1BD486E7FE9FBA065FCD0FAB6" box="[643,804,1334,1359]" italics="true" pageId="5" pageNumber="62">
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||
The
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||
<typeStatus id="24368841BD486E7FE9CDA06BFCD0FAB6" box="[693,804,1336,1359]" pageId="5" pageNumber="62" type="paratype">paratypes</typeStatus>
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||
</emphasis>
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||
—(23 + 2Ƥ
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||
<collectionCode id="9D9CAE26BD486E7FE8C0A065FBF0FAB7" box="[952,1028,1334,1358]" pageId="5" pageNumber="62">LPCB</collectionCode>
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||
): same data as
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<typeStatus id="24368841BD486E7FEFC8A065FAE2FAB7" box="[1200,1302,1334,1358]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
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||
; (1
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||
<collectionCode id="9D9CAE26BD486E7FEE42A064FA6CFAB6" box="[1338,1432,1335,1359]" pageId="5" pageNumber="62">ƤLPCB</collectionCode>
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||
, 1Ƥ
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||
<collectionCode id="9D9CAE26BD486E7FEBBAA009FEE3FA8B" box="[194,279,1370,1394]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="62">NMPC</collectionCode>
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||
):
|
||
<collectingCountry id="839A7673BD486E7FEA52A008FE8FFA8B" box="[298,379,1371,1394]" name="Yemen" pageId="5" pageNumber="62">Yemen</collectingCountry>
|
||
,
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||
<collectingCountry id="839A7673BD486E7FEAFEA009FE2FFA8B" box="[390,475,1370,1394]" name="Yemen" pageId="5" pageNumber="62">Socotra</collectingCountry>
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||
Island, Firmihin plato,
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||
<quantity id="3C759B06BD486E7FE99FA009FC9DFA8B" box="[743,873,1370,1394]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="62" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
|
||
,
|
||
<geoCoordinate id="9EB95024BD486E7FE80DA008FBFAFA8B" box="[885,1038,1370,1394]" direction="north" orientation="latitude" pageId="5" pageNumber="62" precision="15" value="12.479166">N 12°28´45´´</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9EB95024BD486E7FEF63A008FB45FA8B" box="[1051,1201,1370,1394]" direction="east" orientation="longitude" pageId="5" pageNumber="62" precision="15" value="54.024723">E 54°00´89´´</geoCoordinate>
|
||
,
|
||
<date id="8F331023BD486E7FEFC5A009FA8DFA8B" box="[1213,1401,1370,1394]" pageId="5" pageNumber="62" value="2010-06-18" valueMax="2010-06-19" valueMin="2010-06-18">18.–19. vi. 2010</date>
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||
, V. Hula & J. Niedobová leg.; (1Ƥ
|
||
<collectionCode id="9D9CAE26BD486E7FE973A02DFDACFA6F" box="[523,600,1406,1430]" pageId="5" pageNumber="62">SBCB</collectionCode>
|
||
):
|
||
<collectingCountry id="839A7673BD486E7FE915A02CFD25FA6F" box="[621,721,1407,1430]" name="Yemen" pageId="5" pageNumber="62">YEMEN</collectingCountry>
|
||
E,
|
||
<collectingCountry id="839A7673BD486E7FE985A02DFCA6FA6F" box="[765,850,1406,1430]" name="Yemen" pageId="5" pageNumber="62">Socotra</collectingCountry>
|
||
Isl.,
|
||
<date id="8F331023BD486E7FE8E9A02DFBDBFA6F" box="[913,1071,1406,1430]" pageId="5" pageNumber="62" value="1999-09-06" valueMax="1999-09-24" valueMin="1999-09-06">6.–24.ix.1999</date>
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||
, V. Bejček & K. Šťastný lgt. // COLLECTION STANISLAV BEČVÁŘ {yellow label} //
|
||
<typeStatus id="24368841BD486E7FE854A0F1FC39FA43" box="[812,973,1442,1466]" pageId="5" pageNumber="62" type="paratype">PARATYPUS</typeStatus>
|
||
,
|
||
<taxonomicName id="3C8D4D60BD486E7FE8A1A0F0FB7AFA43" box="[985,1166,1442,1466]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="wraniki" status="sp. nov.">Deretus wraniki</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FEFEEA0F0FB21FA43" box="[1174,1237,1443,1466]" pageId="5" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
, SCHAWALLER 2000 {red label} // Collection Stanislav Bečvář {green label}.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD486E7FEBBDA0B9FF05F8BF" blockId="5.[151,1437,1262,2007]" pageId="5" pageNumber="62">
|
||
<emphasis id="C9F9EAF1BD486E7FEBBDA0B9FE38F9FA" bold="true" box="[197,460,1514,1539]" pageId="5" pageNumber="62">Differential diagnosis.</emphasis>
|
||
This species can be separated from
|
||
<taxonomicName id="3C8D4D60BD486E7FE811A0BFFC32F9FA" box="[873,966,1515,1539]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="hulai" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FE811A0BFFC32F9FA" box="[873,966,1515,1539]" italics="true" pageId="5" pageNumber="62">D. hulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FE8B6A0B8FBDDF9FA" box="[974,1065,1515,1539]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FE8B6A0B8FBDDF9FA" bold="true" box="[974,1065,1515,1539]" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by its transverse pronotum (
|
||
<figureCitation id="63B62A66BD486E7FEE16A0B9FF40F9DF" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 – 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="5" pageNumber="62">Fig. 16</figureCitation>
|
||
) and smooth elytra (in
|
||
<taxonomicName id="3C8D4D60BD486E7FEACFA343FDE7F9DF" box="[439,531,1550,1575]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="hulai" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FEACFA343FDE7F9DF" box="[439,531,1550,1575]" italics="true" pageId="5" pageNumber="62">D. hulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FE961A35CFD87F9DE" box="[537,627,1551,1575]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FE961A35CFD87F9DE" bold="true" box="[537,627,1551,1575]" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
subquadrate—Fig. 15 and sculptured respectively). From
|
||
<taxonomicName id="3C8D4D60BD486E7FEF86A343FA87F9DF" box="[1278,1395,1550,1575]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FEF86A343FA87F9DF" box="[1278,1395,1550,1575]" italics="true" pageId="5" pageNumber="62">D. bezdeki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FEE02A35CFF3CF9B2" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FEE02A35CFF3CF9B2" bold="true" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
it differs by having smooth elytra, a non dentate pronotum which also lacks extended anterior corners (elytra sculptured, pronotal sides dentate and with extremely protruded anterior corners of pronotum in
|
||
<taxonomicName id="3C8D4D60BD486E7FEF8FA30BFA84F997" box="[1271,1392,1622,1647]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FEF8FA30BFA84F997" box="[1271,1392,1622,1647]" italics="true" pageId="5" pageNumber="62">D. bezdeki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FEE02A304FF3CF96A" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FEE02A304FF3CF96A" bold="true" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
). From
|
||
<taxonomicName id="3C8D4D60BD486E7FEA51A32FFE37F96A" box="[297,451,1659,1683]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="denticollis">
|
||
<emphasis id="C9F9EAF1BD486E7FEA51A32FFE37F96A" box="[297,451,1659,1683]" italics="true" pageId="5" pageNumber="62">D. denticollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3C8D4D60BD486E7FEA87A32FFD5FF96A" box="[511,683,1660,1683]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="necopintaus" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FEA87A32FFD5FF96A" box="[511,683,1660,1683]" italics="true" pageId="5" pageNumber="62">D. necopintaus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FE9CEA328FCE6F96A" box="[694,786,1659,1683]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FE9CEA328FCE6F96A" bold="true" box="[694,786,1659,1683]" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
it differs by having a distinctly convex pronotum when viewed laterally (more or less flat in
|
||
<taxonomicName id="3C8D4D60BD486E7FE94CA3F3FD3FF94F" box="[564,715,1694,1719]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="denticollis">
|
||
<emphasis id="C9F9EAF1BD486E7FE94CA3F3FD3FF94F" box="[564,715,1694,1719]" italics="true" pageId="5" pageNumber="62">D. denticollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3C8D4D60BD486E7FE879A3F3FC5FF94F" box="[769,939,1695,1719]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="necopinatus" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FE879A3F3FC5FF94F" box="[769,939,1695,1719]" italics="true" pageId="5" pageNumber="62">D. necopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FE8CAA3CCFBFFF94E" box="[946,1035,1695,1719]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FE8CAA3CCFBFFF94E" bold="true" box="[946,1035,1695,1719]" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
), by elytra length / pronotum width ratio less than 1.8 and by the sixth elytral interval with a longitudinal impression at the elytral base (no impressions on elytra in the latter two species). It also differs from
|
||
<taxonomicName id="3C8D4D60BD486E7FE879A3BBFC5FF907" box="[769,939,1767,1791]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="necopinatus" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD486E7FE879A3BBFC5FF907" box="[769,939,1767,1791]" italics="true" pageId="5" pageNumber="62">D. necopinatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD486E7FE8CBA3B4FBF9F906" box="[947,1037,1767,1791]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="C9F9EAF1BD486E7FE8CBA3B4FBF9F906" bold="true" box="[947,1037,1767,1791]" pageId="5" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by its shiny body (dull in the latter species). From
|
||
<taxonomicName id="3C8D4D60BD486E7FEA31A25FFE15F8DA" box="[329,481,1803,1827]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="spinicollis">
|
||
<emphasis id="C9F9EAF1BD486E7FEA31A25FFE15F8DA" box="[329,481,1803,1827]" italics="true" pageId="5" pageNumber="62">D. spinicollis</emphasis>
|
||
</taxonomicName>
|
||
it can be distinguished by its much smaller size and unextended anterior pronotal corners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3976568BD486E7CEBBDA201FE12FCCE" lastPageId="6" lastPageNumber="63" pageId="5" pageNumber="62" type="description">
|
||
<paragraph id="FB3236E3BD486E7FEBBDA201FBF2F877" blockId="5.[151,1437,1262,2007]" pageId="5" pageNumber="62">
|
||
<emphasis id="C9F9EAF1BD486E7FEBBDA201FEA2F893" bold="true" box="[197,342,1874,1899]" pageId="5" pageNumber="62">Description.</emphasis>
|
||
<typeStatus id="24368841BD486E7FEA25A201FE3CF893" box="[349,456,1874,1898]" pageId="5" pageNumber="62" type="holotype">Holotype</typeStatus>
|
||
, body length
|
||
<quantity id="3C759B06BD486E7FE918A201FD4CF893" box="[608,696,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="62" unit="mm" value="7.2">7.2 mm</quantity>
|
||
, width
|
||
<quantity id="3C759B06BD486E7FE873A201FC90F893" box="[779,868,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="62" unit="mm" value="2.7">2.7 mm</quantity>
|
||
. Variability of size in paratypes—females: 5.8–8.3 x
|
||
<quantity id="3C759B06BD486E7FEBD4A225FEC2F877" box="[172,310,1910,1934]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.6" metricValueMin="2.5" pageId="5" pageNumber="62" unit="mm" value="3.05" valueMax="3.6" valueMin="2.5">2.5–3.6 mm</quantity>
|
||
; males: 5.6–7.8 x 2.3–3.0 mm. Body dark brown, shiny (
|
||
<figureCitation id="63B62A66BD486E7FE8CBA225FC0CF877" box="[947,1016,1910,1934]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1945,1968]" captionTargetBox="[161,1425,339,1921]" captionTargetId="figure@2.[161,1426,339,1921]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Habitus, dorsal view. 1. Deretus bezdeki sp. nov. (paratype), male; 2. D. bezdeki sp. nov. (paratype), female; 3. D. denticollis Gahan, 1900, male; 4. D. denticollis Gahan, 1900, female; 5. D. hulai sp. nov. (paratype), female; 6. D. maderai sp. nov. (holotype), male; 7. D. maderai sp. nov. (paratype), female." httpUri="https://zenodo.org/record/211333/files/figure.png" pageId="5" pageNumber="62">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD486E7CEBBDA2C8FE55FEC6" blockId="5.[151,1437,1262,2007]" lastBlockId="6.[151,1437,151,1112]" lastPageId="6" lastPageNumber="63" pageId="5" pageNumber="62">
|
||
<emphasis id="C9F9EAF1BD486E7FEBBDA2C8FEF6F84A" box="[197,258,1947,1971]" italics="true" pageId="5" pageNumber="62">Head</emphasis>
|
||
densely punctate, punctures as large as eye facets. Space between punctures narrower or equal to their diameter. Eyes reniform, not divided by genae. Clypeus with almost straight anterior margin and with several long yellowish setae anteriorly. Labrum transverse, punctate and covered with long yellowish setae. Ventral side of head shiny and punctate. Mentum transverse with midlongitudinal and smooth gibbosity. Apical labial palpomere broadened, triangular in both sexes. Antennae filiform, relatively long, covered by whitish setae. Third antennomere prolonged, approximately as long as the two following joints combined or slightly shorter. Apical antennomere pale brown, longer in males.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA41EFBFBFE55" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA41EFEC0FE9D" box="[197,308,333,356]" italics="true" pageId="6" pageNumber="63">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="63B62A66BD4B6E7CEA3CA41FFE62FE9D" box="[324,406,332,356]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1753,1776]" captionText="FIGURES 13 – 18. Pronotum, dorsal view. 13. Deretus bezdeki sp. nov. (paratype), male; 14. D. denticollis Gahan, 1900, male; 15. D. hulai sp. nov. (paratype), female; 16. D. maderai sp. nov. (holotype), male; 17. Deretus necopinatus sp. nov. (paratype), male; 18. D. spinicollis Schawaller, 2004, male." pageId="6" pageNumber="63">Fig. 16</figureCitation>
|
||
) transverse, distinctly convex (lateral view), punctate, punctures same as on head. Anterior margin broadly and shallowly emarginated. Both lateral margins irregularly serrate. Basal, lateral, and anterior margins of pronotum distinctly bordered. Anterior margin medially obliterated.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA4EBFCA1FDA6" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA4EBFEFFFE29" box="[197,267,440,464]" italics="true" pageId="6" pageNumber="63">Elytra</emphasis>
|
||
subparallel in males and subovoid in females. Slightly convex (lateral view) with moderate apical declivity (lateral view). Base of elytra bordered only around humeri. Scutellum triangular, finely punctate. Elytral striae conspicuous, composed of punctures, puncture diameter subequal to eye facet. Elytral interstriae glabrous, convex and punctate with punctures smaller than eye facet. Sixth elytral interval with longitudinal impression at elytral base. Elytral epipleuron broad and very finely punctate.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA73EFC22FD35" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<taxonomicName id="3C8D4D60BD4B6E7CEBBDA73EFEBCFD7D" box="[197,328,621,644]" class="Insecta" family="Elateridae" genus="Prosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA73EFEBCFD7D" box="[197,328,621,644]" italics="true" pageId="6" pageNumber="63">Prosternum</emphasis>
|
||
</taxonomicName>
|
||
, hypomeron, mesoventrite, mesepisternum, mesepimeron and metaventrite smooth, glabrous, densely and coarsely punctate. Prosternal process widened behind coxae (ventral view). Abdominal ventrites shallowly punctate, moderately pubescent. Males without patches of sensillae.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA78AFBF3FCED" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA78AFF0EFD09" box="[197,250,729,752]" italics="true" pageId="6" pageNumber="63">Legs</emphasis>
|
||
punctate, covered with yellowish setae. Males with first three protarsomeres distinctly wider than penultimate and apical protarsomeres. Further modifications not present in either sex.
|
||
</paragraph>
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA64CFE12FCCE" blockId="6.[151,1437,151,1112]" box="[197,486,799,824]" pageId="6" pageNumber="63">
|
||
Aedeagus see
|
||
<figureCitation id="63B62A66BD4B6E7CEA1DA64CFE42FCCE" box="[357,438,799,823]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1845,1868]" captionTargetBox="[220,1367,193,1738]" captionTargetId="figure@9.[220,1367,193,1739]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Aedeagus, dorsal view. 19. Deretus bezdeki sp. nov. (paratype), male; 20. D. denticollis Gahan, 1900, male; 21. D. maderai sp. nov. (holotype), male; 22. Deretus necopinatus sp. nov. (paratype), male; 23. D. spinicollis Schawaller, 2004, male." httpUri="https://zenodo.org/record/211335/files/figure.png" pageId="6" pageNumber="63">Fig. 21</figureCitation>
|
||
,
|
||
<figureCitation id="63B62A66BD4B6E7CEABAA64CFE15FCCE" box="[450,481,799,823]" captionStart="FIGURES 24 – 28" captionStartId="9.[151,264,1937,1960]" captionText="FIGURES 24 – 28. Aedeagus, lateral view. 24. Deretus bezdeki sp. nov. (paratype), male; 25. D. denticollis Gahan, 1900, male; 26. D. maderai sp. nov. (holotype), male; 27. Deretus necopinatus sp. nov. (paratype), male; 28. D. spinicollis Schawaller, 2004, male." pageId="6" pageNumber="63">26</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3976568BD4B6E7CEBBDA617FB23FC86" pageId="6" pageNumber="63" type="etymology">
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA617FB23FC86" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA617FEB8FCA2" bold="true" box="[197,332,836,860]" pageId="6" pageNumber="63">Etymology.</emphasis>
|
||
Dedicated to Dr. Petr Madĕra (
|
||
<collectingCountry id="839A7673BD4B6E7CE9C9A617FC9FFCA5" box="[689,875,836,860]" name="Czech Republic" pageId="6" pageNumber="63">Czech Republic</collectingCountry>
|
||
), botanist and dendrologist. Without his generous offer to join his educational project I would never have had the possibility to visit
|
||
<collectingCountry id="839A7673BD4B6E7CEF56A634FB70FC86" box="[1070,1156,871,895]" name="Yemen" pageId="6" pageNumber="63">Socotra</collectingCountry>
|
||
Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3976568BD4B6E7CEBBDA6DFFEECFBA1" pageId="6" pageNumber="63" type="discussion">
|
||
<paragraph id="FB3236E3BD4B6E7CEBBDA6DFFEECFBA1" blockId="6.[151,1437,151,1112]" pageId="6" pageNumber="63">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBBDA6DFFECDFC5A" bold="true" box="[197,313,908,932]" pageId="6" pageNumber="63">Remarks.</emphasis>
|
||
<bibRefCitation id="9F1C4B12BD4B6E7CEA39A6DFFDE4FC5D" author="Schawaller" box="[321,528,908,932]" pageId="6" pageNumber="63" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller (2004)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="3C8D4D60BD4B6E7CE9F4A6DEFC3BFC5D" authority="Schawaller, 2004" authorityName="Schawaller" authorityYear="2004" box="[652,975,908,932]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="wraniki">
|
||
<emphasis id="C9F9EAF1BD4B6E7CE9F4A6DEFCF1FC5D" box="[652,773,908,932]" italics="true" pageId="6" pageNumber="63">D. wraniki</emphasis>
|
||
<bibRefCitation id="9F1C4B12BD4B6E7CE874A6DFFC3BFC5D" author="Schawaller" box="[780,975,908,932]" pageId="6" pageNumber="63" refString="Schawaller, W. (2004) New species and records of Tenebrionidae (Coleoptera) from the Socotra Archipelago. Fauna of Arabia, 20, 439 - 458." type="journal article" year="2004">Schawaller, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
based on two specimens (male
|
||
<typeStatus id="24368841BD4B6E7CEE42A6DFFA68FC5D" box="[1338,1436,908,932]" pageId="6" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
and female
|
||
<typeStatus id="24368841BD4B6E7CEA61A6FCFE8AFC3E" box="[281,382,943,967]" pageId="6" pageNumber="63" type="paratype">paratype</typeStatus>
|
||
), which were very similar to each other, and which he believed belonged together. During this work it was discovered that the
|
||
<typeStatus id="24368841BD4B6E7CE976A687FD84FC15" box="[526,624,980,1004]" pageId="6" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
is identical to the
|
||
<typeStatus id="24368841BD4B6E7CE836A687FC41FC15" box="[846,949,980,1004]" pageId="6" pageNumber="63" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="3C8D4D60BD4B6E7CE899A686FB8FFC15" box="[993,1147,980,1004]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="denticollis">
|
||
<emphasis id="C9F9EAF1BD4B6E7CE899A686FB8FFC15" box="[993,1147,980,1004]" italics="true" pageId="6" pageNumber="63">D. denticollis</emphasis>
|
||
</taxonomicName>
|
||
and therefore it is synonymised (see also remarks in
|
||
<taxonomicName id="3C8D4D60BD4B6E7CEA98A6AAFD82FBE9" box="[480,630,1016,1040]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="denticollis">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEA98A6AAFD82FBE9" box="[480,630,1016,1040]" italics="true" pageId="6" pageNumber="63">D. denticollis</emphasis>
|
||
</taxonomicName>
|
||
). Further examination revealed that the female
|
||
<typeStatus id="24368841BD4B6E7CEFFFA6AAFB1CFBE8" box="[1159,1256,1017,1041]" pageId="6" pageNumber="63" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="3C8D4D60BD4B6E7CEE74A6AAFA77FBE9" box="[1292,1411,1016,1040]" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="wraniki">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEE74A6AAFA77FBE9" box="[1292,1411,1016,1040]" italics="true" pageId="6" pageNumber="63">D. wraniki</emphasis>
|
||
</taxonomicName>
|
||
is a different species, identical to specimens collected during this study, which is described in this paper as
|
||
<taxonomicName id="3C8D4D60BD4B6E7CEE4BA14EFF4CFBA1" class="Insecta" family="Tenebrionidae" genus="Deretus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="maderai" status="sp. nov.">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEE4BA14EFF4CFBA1" italics="true" pageId="6" pageNumber="63">D. maderai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D2CA578ABD4B6E7CEBC6A113FEECFBA1" box="[190,280,1088,1112]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="C9F9EAF1BD4B6E7CEBC6A113FEECFBA1" bold="true" box="[190,280,1088,1112]" pageId="6" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |