treatments-xml/data/6F/C1/96/6FC196345F8D530AAE799175863D5D60.xml
2024-06-21 12:39:53 +02:00

416 lines
65 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1083.75624" ID-GBIF-Dataset="66ba5520-2b6d-4925-8dea-ce6abf9a724b" ID-PMC="PMC8803752" ID-Pensoft-Pub="1313-2970-1083-13" ID-Pensoft-UUID="958BB719DA1D54DB801A9004E3D80C99" ID-PubMed="35115872" ID-ZooBank="D263A9C3D2EB4A2D9D7FECAC41AFD710" ModsDocID="1313-2970-1083-13" checkinTime="1643093437741" checkinUser="pensoft" docAuthor="Kolcsar, Levente-Peter, Paramonov, Nikolai, Imada, Yume, Kato, Daichi, Gamboa, Maribet, Shinoka, Dai, Kato, Makoto &amp; Watanabe, Kozo" docDate="2022" docId="6FC196345F8D530AAE799175863D5D60" docLanguage="en" docName="ZooKeys 1083: 13-88" docOrigin="ZooKeys 1083" docPubDate="2022-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1083.75624" docTitle="Diogma caudata Takahashi 1960" docType="treatment" docVersion="4" id="958BB719DA1D54DB801A9004E3D80C99" lastPageNumber="13" masterDocId="958BB719DA1D54DB801A9004E3D80C99" masterDocTitle="Notes on the taxonomic status and distribution of some Cylindrotomidae (Diptera, Tipuloidea), with emphasis on Japanese species" masterLastPageNumber="88" masterPageNumber="13" pageNumber="13" updateTime="1668151423222" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Notes on the taxonomic status and distribution of some Cylindrotomidae (Diptera, Tipuloidea), with emphasis on Japanese species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kolcsar, Levente-Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7784-2386</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kolcsar.peter@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paramonov, Nikolai</mods:namePart>
<mods:affiliation>Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Imada, Yume</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2173-7389</mods:nameIdentifier>
<mods:affiliation>Graduate School of Science and Engineering, Ehime University, 2 - 5 Bunkyo-cho, Matsuyama, Ehime, 790 - 8577 Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kato, Daichi</mods:namePart>
<mods:affiliation>Echigo-Matsunoyama Museum of Natural Sciences, ' Kyororo', 1712 - 2 Matsunoyama, Tokamachi, 942 - 1411, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gamboa, Maribet</mods:namePart>
<mods:affiliation>Department of Ecology, Faculty of Sciences, Universidad Catolica de la Santisima Concepcion, 409054 Concepcion, Chile</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shinoka, Dai</mods:namePart>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kato, Makoto</mods:namePart>
<mods:affiliation>Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Yoshida-nihonmatsu-cho, Kyoto, 606 - 8501 Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watanabe, Kozo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7062-595X</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1083</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>88</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1083.75624</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1083.75624</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1083-13</mods:identifier>
<mods:identifier type="ZooBank">D263A9C3D2EB4A2D9D7FECAC41AFD710</mods:identifier>
<mods:identifier type="Pensoft-UUID">958BB719DA1D54DB801A9004E3D80C99</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="192586631" LSID="urn:lsid:plazi:treatment:6FC196345F8D530AAE799175863D5D60" httpUri="http://treatment.plazi.org/id/6FC196345F8D530AAE799175863D5D60" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection pageId="0" pageNumber="13" type="nomenclature">
<paragraph pageId="0" pageNumber="13">
<taxonomicName LSID="6FC19634-5F8D-530A-AE79-9175863D5D60" authority="Takahashi, 1960" authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">Diogma caudata Takahashi, 1960</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">Figs 4B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of &quot; Triogma kuwanai &quot; form H Triogma kuwanai (Alexander, 1913) of &quot; Triogma limbinervis &quot; form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">, 5C</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Occurrence data in Japan and surrounding areas of A Cylindrotoma distinctissima (Meigen, 1818) B Diogma glabrata (Meigen, 1818) and D. caudata Takahashi, 1960. Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data. Green dot indicate approximate location of type locality of D. caudata Takahashi, 1960." figureDoi="10.3897/zookeys.1083.75624.figure8" httpUri="https://binary.pensoft.net/fig/637481" pageId="0" pageNumber="13">, 8B</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="reference_group">
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">Diogma caudata</taxonomicName>
in
<bibRefCitation author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
: 82-84: original description;
<bibRefCitation author="Siitonen, O" journalOrPublisher="Notulae entomologica" pageId="0" pageNumber="13" pagination=": 203" refId="B59" refString="Siitonen, O, 1984. Diogma caudata Takahashi, 1960 Suomella uusi vaaksiainen (Cylindrotomidae). Notulae entomologica 64 (4): 203" title="Diogma caudata Takahashi, 1960 Suomella uusi vaaksiainen (Cylindrotomidae)." volume="64" year="1984">Siitonen 1984</bibRefCitation>
: 203: faunistic record;
<bibRefCitation author="Sidorenko, VS" editor="Ler, PA" journalOrPublisher="Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="68 - 70" refId="B58" refString="Sidorenko, VS, 1999. Cylindrotomidae. In: Ler, PA, Ed., Key to the insects of Russian Far East. Vol. VI. Diptera and Siphonaptera. Part 1. Dalnauka, Vladivostok: 68 - 70" title="Cylindrotomidae." volumeTitle="Key to the insects of Russian Far East." year="1999">Sidorenko 1999</bibRefCitation>
: 68-70: identification key, illustration, distribution;
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" pagination="119 - 123" refId="B38" refString="Oosterbroek, P, Dufour, C, Pilipenko, VE, 2001. On the presence of Dolichopeza (subgenus Oropeza) in the westpalaearctic (Diptera, Tipulidae). Bulletin de la Societe Neuchateloise des Sciences Naturelles 124: 119 - 123" title="On the presence of Dolichopeza (subgenus Oropeza) in the westpalaearctic (Diptera, Tipulidae)." volume="124" year="2001">Oosterbroek et al. 2001</bibRefCitation>
: 122: distribution;
<bibRefCitation author="Paramonov, NM" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" refId="B40" refString="Paramonov, NM, 2004a. Diogma caudata Takahashi, 1960 new to the fauna of Russia (Diptera, Cylindrotomidae). Zoosystematica Rossica 12: e258." title="Diogma caudata Takahashi, 1960 new to the fauna of Russia (Diptera, Cylindrotomidae). Zoosystematica Rossica 12: e 258." year="2004 a">Paramonov 2004a</bibRefCitation>
: 258: faunistic record;
<bibRefCitation author="Paramonov, NM" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="13" pagination="215 - 218" refId="B42" refString="Paramonov, NM, 2005. Diogma dmitrii sp. n., a new species of cylindrotomid crane-flies (Diptera, Cylindrotomidae) from the Caucasus. Entomologicheskoe Obozrenie 84: 215 - 218" title="Diogma dmitrii sp. n., a new species of cylindrotomid crane-flies (Diptera, Cylindrotomidae) from the Caucasus." volume="84" year="2005">Paramonov 2005</bibRefCitation>
: 211: comparison;
<bibRefCitation DOI="https://doi.org/10.13140/RG.2.1.1584.4320" author="Mukkala, VM" journalOrPublisher="W-album" pageId="0" pageNumber="13" pagination="1 - 34" refId="B34" refString="Mukkala, VM, Haarto, A, Koponen, S, Mukkala, L, Rinne, V, Salmela, J, 2005. On insects, arachnids and other invertebrates in Kivistoenmaeki, Ilmajoki. W-album 2: 1 - 34, DOI: https://doi.org/10.13140/RG.2.1.1584.4320" title="On insects, arachnids and other invertebrates in Kivistoenmaeki, Ilmajoki." url="https://doi.org/10.13140/RG.2.1.1584.4320" volume="2" year="2005">Mukkala et al. 2005</bibRefCitation>
: 7: faunistic record;
<bibRefCitation author="Bartsch, H" editor="Gardenfors, U" journalOrPublisher="Swedish Species Information Centre, Uppsala" pageId="0" pageNumber="13" pagination="325 - 337" refId="B15" refString="Bartsch, H, Cederberg, B, Engelmark, R, Hedmark, K, Viklund, B, 2005. Diptera. In: Gardenfors, U, Ed., The 2005 red list of Swedish species. Swedish Species Information Centre, Uppsala: 325 - 337" title="Diptera." volumeTitle="The 2005 red list of Swedish species." year="2005">Bartsch et al. 2005</bibRefCitation>
: red list status, faunistic record;
<bibRefCitation author="Paramonov, NM" editor="Leley, AS" journalOrPublisher="6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok" pageId="0" pageNumber="13" pagination="887 - 889" refId="B43" refString="Paramonov, NM, 2006. Cylindrotomidae. In: Leley, AS, Ed., Key to the Insects of Russian Far East. 6. Diptera and Siphonaptera. 4. Supplement. Dalnauka, Vladivostok: 887 - 889" title="Cylindrotomidae." volumeTitle="Key to the Insects of Russian Far East." year="2006">Paramonov 2006</bibRefCitation>
: 888-889: identification key, illustration, distribution;
<bibRefCitation author="Polevoi, AV" journalOrPublisher="Trudy Karelskogo Nauchnogo Tsentra RAN (Transactions of the Karelian Research Centre of the Russian Academy of Science)" pageId="0" pageNumber="13" pagination="95 - 104" refId="B49" refString="Polevoi, AV, 2006. New data on the Diptera fauna of Kivach Nature Reserve. Trudy Karelskogo Nauchnogo Tsentra RAN (Transactions of the Karelian Research Centre of the Russian Academy of Science) 2006: 95 - 104" title="New data on the Diptera fauna of Kivach Nature Reserve." volume="2006" year="2006">Polevoi 2006</bibRefCitation>
: 96: faunistic records;
<bibRefCitation author="Sandstroem, J" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="13" publicationUrl="https://www.artdatabanken.se/" refId="B52" refString="Sandstroem, J, 2008. ArtDatabanken: Faktablad, Diogma caudata - nordlig strimharkrank. https://www.artdatabanken.se/" title="ArtDatabanken: Faktablad, Diogma caudata - nordlig strimharkrank." url="https://www.artdatabanken.se/" year="2008">
<normalizedToken originalValue="Sandström">Sandstroem</normalizedToken>
2008
</bibRefCitation>
: red list status;
<bibRefCitation author="Salmela, J" journalOrPublisher="W-album" pageId="0" pageNumber="13" pagination="1 - 63" refId="B53" refString="Salmela, J, 2008. Semiaquatic fly (Diptera, Nematocera) fauna of fens, springs, headwater streams and alpine wetlands in the northern boreal ecoregion, Finland. W-album 6: 1 - 63" title="Semiaquatic fly (Diptera, Nematocera) fauna of fens, springs, headwater streams and alpine wetlands in the northern boreal ecoregion, Finland." volume="6" year="2008">Salmela 2008</bibRefCitation>
: 12: ecology;
<bibRefCitation DOI="https://doi.org/10.33338/ef.5002" author="Salmela, J" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="13" pagination="219 - 242" refId="B54" refString="Salmela, J, 2012a. Annotated list of Finnish crane flies (Diptera, Tipulidae, Limoniidae, Pediciidae, Cylindrotomidae). Entomologica Fennica 22: 219 - 242, DOI: https://doi.org/10.33338/ef.5002" title="Annotated list of Finnish crane flies (Diptera, Tipulidae, Limoniidae, Pediciidae, Cylindrotomidae)." url="https://doi.org/10.33338/ef.5002" volume="22" year="2012 a">Salmela 2012a</bibRefCitation>
: 242: distribution;
<bibRefCitation DOI="https://doi.org/10.1155/2012/913710" author="Salmela, J" journalOrPublisher="Psyche" pageId="0" pageNumber="13" pagination="1 - 19" refId="B55" refString="Salmela, J, 2012b. Biogeographic patterns of Finnish crane flies (Diptera, Tipuloidea). Psyche 2012: 1 - 19, DOI: https://doi.org/10.1155/2012/913710" title="Biogeographic patterns of Finnish crane flies (Diptera, Tipuloidea)." url="https://doi.org/10.1155/2012/913710" volume="2012" year="2012 b">Salmela 2012b</bibRefCitation>
: 16: distribution;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.441.7533" author="Salmela, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="13" pagination="21 - 36" refId="B56" refString="Salmela, J, Petrasiūnas, A, 2014. Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland. ZooKeys 441: 21 - 36, DOI: https://doi.org/10.3897/zookeys.441.7533" title="Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland." url="https://doi.org/10.3897/zookeys.441.7533" volume="441" year="2014">
Salmela and
<normalizedToken originalValue="Petrašiūnas">Petrasiūnas</normalizedToken>
2014
</bibRefCitation>
: 31: checklist;
<bibRefCitation author="Nakamura, T" journalOrPublisher="Tokai University Press, Kanawaga" pageId="0" pageNumber="13" refId="B37" refString="Nakamura, T, 2014. Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." title="Cylindrotomidae. In: Nakamura T, Saigusa T, Suwa M (Eds) Catalogue of the insects of Japan, Volume 8 Diptera (Part 1 Nematocera - BrachyceraAschiza), Touka Shobo, Fukuoka, 54." year="2014">Nakamura 2014</bibRefCitation>
: 54: distribution.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type material">
<paragraph pageId="0" pageNumber="13">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="13">Diogma caudata</emphasis>
</taxonomicName>
Takahashi:
<emphasis bold="true" italics="true" pageId="0" pageNumber="13">Holotype</emphasis>
: • ♂; Japan, Hokkaido, Mount Meakandake; 5 Jul. 1958; M. Takahashi leg.; ELUK.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type material">
<paragraph pageId="0" pageNumber="13">Non-type material examined.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Finland</emphasis>
• 3 ♂, 1 ♀; Kaavi, Kalalamminpuro;
<geoCoordinate degrees="63.11458" direction="north" orientation="latitude" precision="1" value="63.11458">63.11458°N</geoCoordinate>
,
<geoCoordinate degrees="28.67255" direction="east" orientation="longitude" precision="1" value="28.67255">28.67255°E</geoCoordinate>
; alt. 140 m; 20 Jun. 2008 - 17 Jul. 2008; J. Salmela leg.; LMM, CKLP.
<emphasis bold="true" pageId="0" pageNumber="13">Russia</emphasis>
• 1 ♂, 1 ♀; Arkhangelsk Oblast, Plesetsk District, Obozersky Settlement, around the settlement;
<geoCoordinate degrees="63.44231" direction="north" orientation="latitude" precision="1" value="63.44231">63.44231°N</geoCoordinate>
,
<geoCoordinate degrees="40.30789" direction="east" orientation="longitude" precision="1" value="40.30789">40.30789°E</geoCoordinate>
; alt. 100 m; 26 Jun. 1959; N.P. Krivosheina leg.; CKLP. • 1 ♂; Karelia Republic, Kon: 6909:550, Kondopoga District, Kivach Nature Reserve, spruce forest;
<geoCoordinate degrees="62.26766" direction="north" orientation="latitude" precision="1" value="62.26766">62.26766°N</geoCoordinate>
,
<geoCoordinate degrees="33.97975" direction="east" orientation="longitude" precision="1" value="33.97975">33.97975°E</geoCoordinate>
, alt. 42 m; 19 May. 1993 - 23 Jun. 1993; A.V. Polevoi leg.; window trap; ZIN. • 1 ♂; Karelia Republic, Karelia, Kon: 6982:570, Medvezhyegorsk Urban Settlement, 3 km NW Medvezhyegorsk City, point
<normalizedToken originalValue="№">No</normalizedToken>
6;
<geoCoordinate degrees="62.93364" direction="north" orientation="latitude" precision="1" value="62.93364">62.93364°N</geoCoordinate>
,
<geoCoordinate degrees="34.38467" direction="east" orientation="longitude" precision="1" value="34.38467">34.38467°E</geoCoordinate>
; alt. 130 m; 19 Jul. 2002; A.V. Polevoi leg.; ZIN. • 1 ♂; Perm Krai, Kungur Urban Okrug, Kungur City, forest station;
<geoCoordinate degrees="57.42881" direction="north" orientation="latitude" precision="1" value="57.42881">57.42881°N</geoCoordinate>
,
<geoCoordinate degrees="56.944206" direction="east" orientation="longitude" precision="1" value="56.944206">56.944206°E</geoCoordinate>
; alt. 219 m; 16 Jun. 1960; K.B. Borisova leg.; ZIN. • 1 ♂; Tuva Republic, Tandinsky District, north slope of Tannu-Ola mountains, near Chagytaj Lake;
<geoCoordinate degrees="50.99591" direction="north" orientation="latitude" precision="1" value="50.99591">50.99591°N</geoCoordinate>
,
<geoCoordinate degrees="94.6764" direction="east" orientation="longitude" precision="5" value="94.6764">94.6764°E</geoCoordinate>
; alt. 1500 m; 24 Jun. 1963; N.A. Violovich leg.; ZIN.
<emphasis bold="true" pageId="0" pageNumber="13">Sweden</emphasis>
• 2 ♂; Lule Lappmark, Kaltbacken bei Messaure;
<geoCoordinate degrees="66.67347" direction="north" orientation="latitude" precision="1" value="66.67347">66.67347°N</geoCoordinate>
,
<geoCoordinate degrees="20.32239" direction="east" orientation="longitude" precision="1" value="20.32239">20.32239°E</geoCoordinate>
; alt. 240 m; 23 Jun. 1969 - 26 Jun. 1969; • 1 ♀; same locality; 22 Jun. 1970 - 24 Jun. 1970; • 2 ♂; same locality; 23 Jun. 1971 - 30 Jun. 1971; • 31 ♂, 4 ♀; same locality; 12 Jun. 1972 - 13 Jul. 1972 / 21. Aug. 1972 - 28 Aug. 1972; • 2 ♂; same locality; 19 Jun. 1973 - 25 Jun. 1973; • 1 ♂, 1 ♀; same locality; 17 Jun. 1974 - 8 Jul. 1974; K.
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
leg.; ZFMK.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="redescription">
<paragraph pageId="0" pageNumber="13">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Head</emphasis>
. Dorsally dark brown, ventrally brown. Frons with white pubescence noticeable only in dry specimens (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11C, D</figureCitation>
). Rostrum pale brown, short, without nasus, with few hairs. Mouthparts pale brown to brown (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11C, D</figureCitation>
). Palpus pale brown to brown, short, five segmented; last segment slightly longer than penultimate segment (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11B</figureCitation>
). Scape cylindrical, 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as pedicel; pedicel ovate, slightly darker than scape; flagellum 14-segmented, gradually darkening from base to tip; segments simple in both sexes, not expanded ventrally (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4B</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11E, F</figureCitation>
); in male, first flagellomere as long as wide, remaining segments cylindrical; last segment 1.2-1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as penultimate segment; flagellomeres with short, relative sparse whitish setae (sensilla), just slightly denser in ventral and lateral sides (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11E</figureCitation>
); in female, last flagellomere 1.8-2
<normalizedToken originalValue="×">x</normalizedToken>
as long as penultimate; last 4-6 segments without sensilla (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Antennae A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1083.75624.figure4" httpUri="https://binary.pensoft.net/fig/637477" pageId="0" pageNumber="13">4B</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Female genital structures of Cylindrotoma nigriventris Loew, 1849 A terminalia, lateral view B sperm ducts C Spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure10" httpUri="https://binary.pensoft.net/fig/637483" pageId="0" pageNumber="13">10F</figureCitation>
). Verticels black, shorter than length of flagellomere; usually one verticel in ventral surface and two or three in dorsal/dorsolateral sides, first segment with 4-6 verticels.
</paragraph>
<caption doi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 11.</emphasis>
<taxonomicName authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="13">Diogma caudata</emphasis>
</taxonomicName>
Takahashi, 1960
<emphasis bold="true" pageId="0" pageNumber="13">A</emphasis>
habitus of the holotype male, lateral view (colouration of wings is artefact)
<emphasis bold="true" pageId="0" pageNumber="13">B</emphasis>
head and thorax of female, lateral view
<emphasis bold="true" pageId="0" pageNumber="13">C</emphasis>
head and thorax of female, lateral and dorsal views
<emphasis bold="true" pageId="0" pageNumber="13">D</emphasis>
head of female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">E</emphasis>
antenna of holotype male
<emphasis bold="true" pageId="0" pageNumber="13">F</emphasis>
antenna of female.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Thorax</emphasis>
. General colouration yellowish brown with contrasting, shiny black markings. Mesonotum pale brown, greenish yellow in fresh, living specimen (
<bibRefCitation author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
), with three separated, broad, longitudinal black markings (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11C</figureCitation>
); several small yellow setae along pale strips. Scutellum yellow. Posterior part of mediotergite black. Anepisternum and katepisternum separated, both ventral parts dark brown to black. Ventral corner of laterotergite black. Additional small darker patch on posterior basalare, and on ventral part of meron. Coxa and trochanter yellowish, darker on anterior- dorsal parts (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11B</figureCitation>
); femur pale brown; tibia gradually darkening from pale brown to dark brown/black; tarsus uniformly black. Wing hyaline; veins pale brown to brown; pterostigma pale (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of &quot; Triogma kuwanai &quot; form H Triogma kuwanai (Alexander, 1913) of &quot; Triogma limbinervis &quot; form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5B</figureCitation>
); three branches of M reaching wing margin; M1 in same level as M1+2; cell a2 narrow,&gt; 7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Wing A Cylindrotoma distinctissima (Meigen, 1818) B Diogma caudata Takahashi, 1960 C Diogma glabrata (Meigen, 1818) D Liogma mikado (Alexander, 1919) E Liogma brevipecten Alexander, 1932 F Liogma serraticornis Alexander, 1919 G Triogma kuwanai (Alexander, 1913) of &quot; Triogma kuwanai &quot; form H Triogma kuwanai (Alexander, 1913) of &quot; Triogma limbinervis &quot; form." figureDoi="10.3897/zookeys.1083.75624.figure5" httpUri="https://binary.pensoft.net/fig/637478" pageId="0" pageNumber="13">5B</figureCitation>
); membrane with interference patterns, visible with dark background (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11A</figureCitation>
). Halter monochrome, yellow or pale brown.
</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Abdomen</emphasis>
. Tergites and sternites pale brown to brown, tergite 8 and sternite 8 darker than others (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diogma caudata Takahashi, 1960 A habitus of the holotype male, lateral view (colouration of wings is artefact) B head and thorax of female, lateral view C head and thorax of female, lateral and dorsal views D head of female, dorsal view E antenna of holotype male F antenna of female." figureDoi="10.3897/zookeys.1083.75624.figure11" httpUri="https://binary.pensoft.net/fig/637484" pageId="0" pageNumber="13">11A</figureCitation>
). Pleural parts greenish in living specimen (
<bibRefCitation author="Takahashi, M" journalOrPublisher="Transactions of the Shikoku Entomological Society" pageId="0" pageNumber="13" pagination="81 - 91" refId="B62" refString="Takahashi, M, 1960. A revision of Japanese Cylindrotominae (Diptera, Tipulidae). Transactions of the Shikoku Entomological Society 6: 81 - 91" title="A revision of Japanese Cylindrotominae (Diptera, Tipulidae)." volume="6" year="1960">Takahashi 1960</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Male terminalia</emphasis>
: Black, large, complex, directed caudally. Tergite 9 not fused with gonocoxite, partly cover gonocoxite (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12C</figureCitation>
); medial part rounded with small tuft of hairs (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12A</figureCitation>
); lateral lobe of tergite 9 greatly extended, complex; as long as basal part of tergite 9; ventral portion of lateral lobe elongated, finger-like in lateral view (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12C</figureCitation>
); lateral margin almost straight or weakly divergent in dorsal view (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12A, B</figureCitation>
); ventral base of lateral lobe with small, black, heavily sclerotised lobe (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12C, E</figureCitation>
) - named lamina by some authors - shape variable; posterior margin of tergite 9 between median round part and lateral lobe covered with dense short, blunt ended setae (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12A</figureCitation>
). Gonocoxite complex; apical lobe with dense hairs (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12B-D</figureCitation>
); ventral lobe round, almost bare (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12B, D</figureCitation>
); inner part of gonocoxite with basally directed lobe, with hairs on margin (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12D</figureCitation>
). Gonostylus simple, wider at middle (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12F</figureCitation>
); with finger-like membranous lobe on inner side, poorly visible in dry specimens (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12G</figureCitation>
). Aedeagus bifid; aedeagus with apical branches long, curved ventrally almost in right angle, then curved in right angle posteriorly, then turn dorsally in right angle in lateral view (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12J</figureCitation>
); dorsal lobe between interbases complex, sclerotised (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12H, J</figureCitation>
); interbase with ventral projection (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12J</figureCitation>
). Sperm pump and ejaculatory apodeme small (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12H-J</figureCitation>
), covered by parameres in lateral view (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, ventral view (aedeagus complex removed) C terminalia, lateral view D gonocoxite, inner lateral view E shape of heavily sclerotised lobe (lamina) of tergite 9 F shape of the gonostylus, caudal view G shape of the gonostylus, inner ventral view H aedeagus complex, dorsal view I aedeagus complex, ventral view J aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t 9 l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe." figureDoi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13">12J</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1083.75624.figure12" httpUri="https://binary.pensoft.net/fig/637485" pageId="0" pageNumber="13" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 12.</emphasis>
Male genital structures of
<taxonomicName authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="13">Diogma caudata</emphasis>
</taxonomicName>
Takahashi, 1960
<emphasis bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, ventral view (aedeagus complex removed)
<emphasis bold="true" pageId="0" pageNumber="13">C</emphasis>
terminalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="13">D</emphasis>
gonocoxite, inner lateral view
<emphasis bold="true" pageId="0" pageNumber="13">E</emphasis>
shape of heavily sclerotised lobe (lamina) of tergite 9
<emphasis bold="true" pageId="0" pageNumber="13">F</emphasis>
shape of the gonostylus, caudal view
<emphasis bold="true" pageId="0" pageNumber="13">G</emphasis>
shape of the gonostylus, inner ventral view
<emphasis bold="true" pageId="0" pageNumber="13">H</emphasis>
aedeagus complex, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">I</emphasis>
aedeagus complex, ventral view
<emphasis bold="true" pageId="0" pageNumber="13">J</emphasis>
aedeagus complex, lateral view. Abbreviations: ae - aedeagus; al - gonocoxite apical lobe; eja - ejaculatory apodeme; gsl - lobe of gonostylus; ib - interbase; ibml - interbase median lobe; ibvl - interbase ventral lobe; pm - paramere; sp - sperm pump; t9l - tergite 9 lateral lobe; vl - gonocoxite ventral lobe.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Female terminalia</emphasis>
: Brown, tip of cercus and hypopygial valve yellowish brown. Tergite 8 separated at middle by membranous area (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13A</figureCitation>
). Tergites 8 and 9 similar in size (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13B</figureCitation>
). Ventral corner of tergite 9 rugged (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13B</figureCitation>
). Triangular sclerite separated from tip of tergite 10 (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13A</figureCitation>
). Lateral lobes of tergite 10 elongated, with few longer hairs (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13A, B</figureCitation>
). Cercus and hypogynial valve simple, wide, blade-shaped, tips rounded (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13B</figureCitation>
). Dorsal apical surface of cercus rough, formed by few blunt, pyramid or round teeth (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13A, B</figureCitation>
). Base of sternite 8, weakly sclerotised, extended laterally at middle, with transverse creases (Fig. 13AC). Two round spermathecae present, duct curved (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13D</figureCitation>
). Lateral sclerite of genital fork elongated; two sperm ducts simple, without any clear markings (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female genital structures of Diogma caudata Takahashi, 1960 A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view D spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t 8 - tergite 8; t 9 - tergite 9; t 10 - tergite 10; t 10 s - tergite 10 triangular sclerite; t 10 ll - tergite 10 lateral lobe; s 8 - stergite 8." figureDoi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13">13D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1083.75624.figure13" httpUri="https://binary.pensoft.net/fig/637486" pageId="0" pageNumber="13" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 13.</emphasis>
Female genital structures of
<taxonomicName authorityName="Takahashi" authorityYear="1960" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma caudata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="13">Diogma caudata</emphasis>
</taxonomicName>
Takahashi, 1960
<emphasis bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="13">C</emphasis>
sternite 8, hypogynial valve, genital fork, and sperm ducts, inner dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">D</emphasis>
spermathecae. Abbreviations: crc - cercus; hv - hypogynial valve; t8 - tergite 8; t9 - tergite 9; t10 - tergite 10; t10s - tergite 10 triangular sclerite; t10ll - tergite 10 lateral lobe; s8 - stergite 8.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="distribution">
<paragraph pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="13">
Finland, Japan (Hokkaido I, Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Occurrence data in Japan and surrounding areas of A Cylindrotoma distinctissima (Meigen, 1818) B Diogma glabrata (Meigen, 1818) and D. caudata Takahashi, 1960. Red dots indicate locations of investigated specimens, white dots indicate approximate locations of literature data. Green dot indicate approximate location of type locality of D. caudata Takahashi, 1960." figureDoi="10.3897/zookeys.1083.75624.figure8" httpUri="https://binary.pensoft.net/fig/637481" pageId="0" pageNumber="13">8B</figureCitation>
), Sweden, and Russia (Kareliya Republic, Perm Krai and Tuva) (
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B39" refString="Oosterbroek, P, 2021. Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea). https://ccw.naturalis.nl/index.php" title="Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea)." url="https://ccw.naturalis.nl/index.php" year="2021">Oosterbroek 2021</bibRefCitation>
). First records from Arkhangelsk Oblast, Russia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="comments">
<paragraph pageId="0" pageNumber="13">Comments.</paragraph>
<paragraph pageId="0" pageNumber="13">
The species was initially described from Hokkaido, Japan. However, no additional Japanese data has been published, and the species was not found in the type locality in our study. The species later was reported from Finland, Sweden, and Russia (Karelia Republic, Perm Krai, and Tuva). Morphologically it is a well separated species from the close related
<taxonomicName baseAuthorityName="Meigen" baseAuthorityYear="1818" class="Insecta" family="Cylindrotomidae" genus="Diogma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diogma glabrata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="glabrata">
<emphasis italics="true" pageId="0" pageNumber="13">Diogma glabrata</emphasis>
</taxonomicName>
, but the Finnish specimens show only a small COI genetic difference from it and form a clade together with the West Palearctic
<taxonomicName family="Cylindrotomidae" kingdom="Animalia" lsidName="D. glabrata" order="Diptera" pageId="0" pageNumber="13" rank="species" species="glabrata">
<emphasis italics="true" pageId="0" pageNumber="13">D. glabrata</emphasis>
</taxonomicName>
. No significant morphological differences were found between the Finnish and Russian specimens and the Japanese holotype.
</paragraph>
</subSubSection>
</treatment>
</document>