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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.820.31527" ID-GBIF-Dataset="19e4c0b1-d8aa-4ff8-a8e0-d639580dc280" ID-PMC="PMC6365488" ID-Pensoft-Pub="1313-2970-820-139" ID-PubMed="30740017" ID-ZBK="83F75B4D6E544E2B94B86525010CF725" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-820-139" ModsDocOrigin="ZooKeys 820" ModsDocTitle="Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)" checkinTime="1548887832952" checkinUser="pensoft" docAuthor="Yoshitomi, Hiroyuki &amp; Sogoh, Koichi" docDate="2019" docId="6FBA5739D5889499217A283009E63161" docLanguage="en" docName="ZooKeys 820: 139-149" docOrigin="ZooKeys 820" docSource="http://dx.doi.org/10.3897/zookeys.820.31527" docTitle="Eumorphus costatus Gorham 1873" docType="treatment" docVersion="4" lastPageNumber="142" masterDocId="ED1CFF82FF84FF97FF8CEE45FFCAFF91" masterDocTitle="Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)" masterLastPageNumber="149" masterPageNumber="139" pageNumber="141" updateTime="1668166862078" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Eumorphusmarginatus group in Sulawesi, Indonesia (Coleoptera, Endomychidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshitomi, Hiroyuki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sogoh, Koichi</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>820</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>149</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.820.31527</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.820.31527</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-820-139</mods:identifier>
<mods:identifier type="ZBK">83F75B4D6E544E2B94B86525010CF725</mods:identifier>
<mods:identifier type="ZooBank">83F75B4D6E544E2B94B86525010CF725</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154126364" LSID="urn:lsid:plazi:treatment:6FBA5739D5889499217A283009E63161" httpUri="http://treatment.plazi.org/id/6FBA5739D5889499217A283009E63161" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
<subSubSection pageId="2" pageNumber="141" type="nomenclature">
<paragraph pageId="2" pageNumber="141">
<taxonomicName authority="Gorham, 1873" authorityName="Gorham" authorityYear="1873" class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus costatus" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="costatus">Eumorphus costatus Gorham, 1873</taxonomicName>
Figs 1A, B; 3A, B; 4A, B; 5A, B; 6
<normalizedToken originalValue="AD">A-D</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="141" type="reference_group">
<paragraph pageId="2" pageNumber="141">
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus costatus" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="costatus">Eumorphus costatus</taxonomicName>
Gorham, 1873: 34, fig. 6.
<bibRefCitation author="Strohecker, HF" journalOrPublisher="Pacific Insect" pageId="9" pageNumber="148" pagination="79 - 112" title="A synopsis of the genus Eumorphus (Coleoptera: Endomychidae)." volume="10" year="1968">Strohecker 1968</bibRefCitation>
: 84, figs. 11, 65 (short description and figures of habitus and male genitalia).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="141" type="materials_examined">
<paragraph pageId="2" pageNumber="141">Material examined.</paragraph>
<paragraph pageId="2" pageNumber="141">
1 male and 2 females (EUMJ), &quot;C. Sulawesi: Puncak Palopo Puncak, Indonesia, 23-25.XI.2012, Kiyoshi Ando leg.&quot;; 1 male (EUMJ), &quot;(C. Sulawesi) Dongidongi, 10 km S of Palolo, alt. ca 1000 m, 18.VI.1986, S. Nagai leg.&quot;; 2 female (EUMJ), &quot;INDONESIA Puncak Palopo Sulawesi Selatan 30-31.XII.1999 N. Ohbayashi leg.&quot;; 1 male &amp; 1 female, &quot;[INDONESIA] Bantu Kaya Angin Battang Palopo Sulawesi Selatan Alt. ca. 350 m
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-2.95">02°57'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="120.13333">120°08'E</geoCoordinate>
31.I.2013 J. Yamasako leg.&quot;; 1 female (EUMJ), &quot;[BONEBOLANGO: INDONESIA] Mt. Tilongkabila (Gunung Tilonkabila) N. Sulawesi, alt. ca. 1300-1500 m
<geoCoordinate direction="north" orientation="latitude" precision="1" value="0.58837223">0°35'18.14&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="123.22298">123°13'22.71&quot;E</geoCoordinate>
-
<geoCoordinate direction="north" orientation="latitude" precision="1" value="0.58843607">0°35'18.37&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="123.222946">123°13'22.61&quot;E</geoCoordinate>
10.VI.2012 Ryo Ogawa leg.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="141" type="diagnosis">
<paragraph pageId="2" pageNumber="141">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="141">
This species is similar to
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus marginatus" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="marginatus">Eumorphus marginatus</taxonomicName>
Fabricius, 1801 but differs from the latter in the following characteristics: elytral spots smaller, each distant from suture by its own diameter; in
<taxonomicName lsidName="E. marginatus" pageId="2" pageNumber="141" rank="species" species="marginatus">E. marginatus</taxonomicName>
elytral spots larger and distant from suture by 1/2 of its own diameter).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141" type="description">
<paragraph pageId="2" pageNumber="141">Description.</paragraph>
<paragraph pageId="2" pageNumber="141">Male.</paragraph>
<paragraph pageId="2" pageNumber="141">Coloration of body black, with blueish luster; elytral spots yellowish orange.</paragraph>
<paragraph pageId="2" pageNumber="141">Antennae long, reaching to ca. basal 1/4 of elytra.</paragraph>
<paragraph lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
Pronotum (Fig. 4A) glossy, narrowly depressed in lateral parts; lateral margins subparallel-sided in basal 1/2, then gently tapered anteriorly; lateral sulci relatively long, reaching ca. basal 1/3 of PLM; anterior angles ca. 60°; posterior angles sharply pointed, projecting postero-laterally; PLM/PLS 0.85-0.86 (0.86); PWP/PWA 1.84-1.95 (1.89); PWP/PLM 1.79-1.99 (1.89); PWP/PLS 1.55-1.69 (1.62). Elytra (Fig. 1A) ovate, bluntly rounded behind, widest at middle; lateral flattened margins wide, flat dorsally; elytral spots two pairs, relatively large, each distant from suture by its own diameter; anterior
<pageBreakToken pageId="3" pageNumber="142" start="start">ones</pageBreakToken>
transverse, situated in anterior 1/4 of elytra; posterior ones oval, situated in posterior 1/3 of elytra; elytra along suture (Fig. 3A) distinctly elevated in mesal part; EL/EW 1.06-1.09 (1.07); EL/PLM 4.09-4.40 (4.24); EW/PWP 2.04-2.14 (2.09); TL/EW 1.32-1.33 (1.33). Fore tibiae (Fig. 5A) stout; apical carinae distinct, covered with yellowish brown pubescence on inner surface; fore tibial tooth long, triangular, situated in apical 1/4. Metatibial apices widely prolonged and flattened, sharply pointed at outer angles.
</paragraph>
<caption pageId="3" pageNumber="142">
<paragraph pageId="3" pageNumber="142">
Figure 1. Habitus of
<taxonomicName class="Insecta" family="Endomychidae" genus="Eumorphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eumorphus" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">Eumorphus</taxonomicName>
spp. A
<taxonomicName lsidName="E. costatus" pageId="3" pageNumber="142" rank="species" species="costatus">E. costatus</taxonomicName>
, male B
<taxonomicName lsidName="E. costatus" pageId="3" pageNumber="142" rank="species" species="costatus">E. costatus</taxonomicName>
, female C
<taxonomicName lsidName="E. wegneri" pageId="3" pageNumber="142" rank="species" species="wegneri">E. wegneri</taxonomicName>
, male. Scale bar: 5.0 mm.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="142">
Aedeagus (Fig. 6
<normalizedToken originalValue="AD">A-D</normalizedToken>
) 5.4 mm long; median lobe stout, expanded ventrally in median part; apical branch short, obtuse and bisinuate at apex; subapical branch long, evenly curved apically; tegmen not movable, with short tegminal strut; endophallus consisting of bifid long right membranous lobe and left membranous lobe, with rather sclerotized spinous plate at base of left lobe.
</paragraph>
<paragraph pageId="3" pageNumber="142">Female. Similar to male, but sexual dimorphism distinct in the following characteristics: posterior angles of pronotum (Fig. 4B) nearly right-angled; elytra with lateral flattened margins narrower than in male, gently arcuate dorsally along suture (Fig. 3B); fore tibiae (Fig. 5B) slender and straight, covered with yellowish brown pubescence in apical half; metatibial apices simple; PLM/PLS 0.80-0.84 (0.82); PWP/PWA 1.46-1.69 (1.59); PWP/PLM 1.59-1.77 (1.66); PWP/PLS 1.31-1.44 (1.37); EL/EW 1.10-1.32 (1.24); EL/PLM 3.86-4.25 (4.07); EW/PWP 1.93-2.13 (1.98); TL/EW 1.37-1.63 (1.55).</paragraph>
<paragraph pageId="3" pageNumber="142">Measurements. Male (n = 2). TL 16.68-18.53 (17.61) mm; PW 5.88-6.83 (6.36) mm; PW 3.20-3.50 (3.35) mm; PL 3.28-3.43 (3.36) mm; PL 3.80-4.05 (3.93) mm; EL 13.40-15.10 (14.25) mm; EW 12.6-13.9 (13.25) mm. Female (n = 6). TL 13.40-16.85 (15.17) mm; PW 4.55-5.60 (4.96) mm; PW 2.90-3.35 (3.12) mm; PL 2.60-3.30 (2.99) mm; PL 3.20-4.00 (3.63) mm; EL 10.80-13.55 (12.18) mm; EW 8.85-10.85 (9.79) mm.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="142" type="distribution">
<paragraph pageId="3" pageNumber="142">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="142">Indonesia (Sulawesi: South, Central, and North).</paragraph>
</subSubSection>
</treatment>
</document>