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<document id="0481C53C3EF739FA878A601A0E4A5237" ID-CLB-Dataset="26164" ID-DOI="10.11646/zootaxa.4639.1" ID-GBIF-Dataset="812e0570-38c6-4f6e-840c-7d52d87a8c12" ID-ISSN="1175-5326" ID-PMC="PMC6910866" ID-PubMed="31712455" ID-Zenodo-Dep="3342161" ID-ZooBank="B1690E30-EC81-46D3-881D-97648DDC7745" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1563526566144" checkinUser="plazi" docAuthor="Madeira, Patrícia, Kroh, Andreas, Cordeiro, Ricardo, De, António M., Martins, Frias &amp; Ávila, Sérgio P." docDate="2019" docId="4148D21204DBFF5FFF33F8F5757411C0" docLanguage="en" docName="zootaxa.4639.1.1.pdf.imf" docOrigin="Zootaxa 4639 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Penilpidia midatlantica $ Gebruk 2008" docType="treatment" docVersion="16" lastPageNumber="141" masterDocId="BD71AA6A0450FFD3FFA4FFCD76121037" masterDocTitle="The Echinoderm Fauna of the Azores (NE Atlantic Ocean)" masterLastPageNumber="231" masterPageNumber="1" pageNumber="140" updateTime="1698685128211" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="99CCCFCFEF6FDA06C747F271FA15599D">The Echinoderm Fauna of the Azores (NE Atlantic Ocean)</mods:title>
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<mods:namePart id="86C7CB03990795A80F871A47E9234638">Kroh, Andreas</mods:namePart>
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<mods:namePart id="8B1BA658005BD24C003F15457D6682A6">De, António M.</mods:namePart>
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<taxonomicName id="0EE1188704DBFF58FF33F8F5745D1765" ID-CoL="76J2V" authority="Gebruk, 2008" authorityName="$ Gebruk" authorityYear="2008" box="[151,591,1848,1875]" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="139" pageNumber="140" phylum="Echinodermata" rank="species" species="midatlantica">
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<emphasis id="FB95BF1604DBFF58FF33F8F577B71764" bold="true" box="[151,421,1848,1875]" italics="true" pageId="139" pageNumber="140">Penilpidia midatlantica</emphasis>
<bibRefCitation id="AD701EF504DBFF58FE08F8F5745D1765" author="Gebruk" box="[428,591,1848,1875]" firstAuthor="Gebruk" pageId="139" pageNumber="140" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk, 2008</bibRefCitation>
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Reports for the
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:
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<taxonomicName id="0EE1188704DBFF58FF33F856747D1783" ID-CoL="76J2V" authority="$ Gebruk, 2008: 52 - 54" authorityName="$ Gebruk" authorityPageNumber="52" authorityYear="2008" box="[151,623,1945,1972]" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="139" pageNumber="140" phylum="Echinodermata" rank="species" species="midatlantica">
<emphasis id="FB95BF1604DBFF58FF33F85677951784" box="[151,391,1947,1971]" italics="true" pageId="139" pageNumber="140">Penilpidia midatlantica</emphasis>
<superScript id="3E94CE4C04DBFF58FE29F85477861790" attach="right" box="[397,404,1945,1959]" fontSize="6" pageId="139" pageNumber="140">$</superScript>
<bibRefCitation id="AD701EF504DBFF58FE31F856747D1783" author="Gebruk" box="[405,623,1947,1972]" firstAuthor="Gebruk" pageId="139" pageNumber="140" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk, 2008: 5254</bibRefCitation>
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, figs. 46;
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Rogacheva
<emphasis id="FB95BF1604DBFF58FCF2F85175911784" box="[854,899,1947,1971]" italics="true" pageId="139" pageNumber="140">et al</emphasis>
. 2013: 610611
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.
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<typeStatus id="165ADDA604DBFF58FF33F81076C617C0" box="[151,212,2013,2039]" pageId="139" pageNumber="140">Type</typeStatus>
locality:
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,
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(
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.
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<emphasis id="FB95BF1604DCFF5FFF63FF5576EB1084" bold="true" box="[199,249,152,179]" pageId="140" pageNumber="141">See:</emphasis>
<bibRefCitation id="AD701EF504DCFF5FFEA4FF5577BB1084" author="Gebruk" box="[256,425,152,179]" firstAuthor="Gebruk" pageId="140" pageNumber="141" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk (2008)</bibRefCitation>
.
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<emphasis id="FB95BF1604DCFF5FFF63FF71774E10E0" bold="true" box="[199,348,188,215]" pageId="140" pageNumber="141">Occurrence:</emphasis>
known only from the Mid-Atlantic Ridge, from SE of the Charlie-Gibbs Fracture Zone (
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Rogacheva
<emphasis id="FB95BF1604DCFF5FFF5EFF2C773910CD" box="[250,299,224,250]" italics="true" pageId="140" pageNumber="141">et al</emphasis>
. 2013
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) south to the
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(
<bibRefCitation id="AD701EF504DCFF5FFDC3FF2D751310CC" author="Gebruk" box="[615,769,224,251]" firstAuthor="Gebruk" pageId="140" pageNumber="141" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk 2008</bibRefCitation>
).
</paragraph>
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<emphasis id="FB95BF1604DCFF5FFF63FEC877081128" bold="true" box="[199,282,261,287]" pageId="140" pageNumber="141">Depth:</emphasis>
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(
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,
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Rogacheva
<emphasis id="FB95BF1604DCFF5FFD5AFEC8753D1129" box="[766,815,260,286]" italics="true" pageId="140" pageNumber="141">et al</emphasis>
. 2013
</bibRefCitation>
);
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:
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(
<bibRefCitation id="AD701EF504DCFF5FFB2FFEC973371128" author="Gebruk" box="[1163,1317,260,287]" firstAuthor="Gebruk" pageId="140" pageNumber="141" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk 2008</bibRefCitation>
).
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<paragraph id="C95E630404DCFF5FFF63FEE575D21174" blockId="140.[151,1437,152,503]" box="[199,960,296,323]" pageId="140" pageNumber="141">
<emphasis id="FB95BF1604DCFF5FFF63FEE5773E1174" bold="true" box="[199,300,296,323]" pageId="140" pageNumber="141">Habitat:</emphasis>
a probable benthopelagic species (
<bibRefCitation id="AD701EF504DCFF5FFD17FEE575A61174" author="Rogacheva" box="[691,948,296,323]" etAl="et al." firstAuthor="Rogacheva" pageId="140" pageNumber="141" pagination="213 - 224" refId="ref198932" refString="Rogacheva, A., Gebruk, A. &amp; Alt, C. H. S. (2012) Swimming deep-sea holothurians (Echinodermata: Holothuroidea) on the northern Mid-Atlantic Ridge. Zoosymposia, 7, 213 - 224. https: // doi. org / 10.1080 / 17451000.2012.750428" type="journal article" year="2012">
Rogacheva
<emphasis id="FB95BF1604DCFF5FFC9CFEE475781175" box="[824,874,296,322]" italics="true" pageId="140" pageNumber="141">et al</emphasis>
. 2012
</bibRefCitation>
).
</paragraph>
<paragraph id="C95E630404DCFF5FFF63FE80757411C0" blockId="140.[151,1437,152,503]" pageId="140" pageNumber="141">
<emphasis id="FB95BF1604DCFF5FFF63FE80772F1150" bold="true" box="[199,317,333,359]" pageId="140" pageNumber="141">Remarks:</emphasis>
<bibRefCitation id="AD701EF504DCFF5FFEE2FE8177FC1150" author="Gebruk" box="[326,494,332,359]" firstAuthor="Gebruk" pageId="140" pageNumber="141" pagination="48 - 60" refId="ref181872" refString="Gebruk, A. V. (2008) Holothurians (Holothuroidea, Echinodermata) of the northern Mid-Atlantic Ridge collected by the G. O. Sars MAR-ECO expedition with descriptions of four new species. In: Gebruk, A. V. (Ed.) Benthic fauna of the northern Mid-Atlantic Ridge: results of the MAR-ECO expedition. Marine Biology Research, 4 (1 - 2), pp. 48 - 60. https: // doi. org / 10.1080 / 17451000701842898" type="journal article" year="2008">Gebruk (2008)</bibRefCitation>
described
<taxonomicName id="0EE1188704DCFF5FFDCFFE8175651151" authorityName="$ Gebruk" authorityYear="2008" box="[619,887,332,358]" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="140" pageNumber="141" phylum="Echinodermata" rank="species" species="midatlantica">
<emphasis id="FB95BF1604DCFF5FFDCFFE8175651151" box="[619,887,332,358]" italics="true" pageId="140" pageNumber="141">Penilpidia midatlantica</emphasis>
</taxonomicName>
based on material collected in the Mid-Atlantic Ridge, in the northern waters of the
<collectingRegion id="0B25ADE604DCFF5FFD9FFEBC749811BC" box="[571,650,369,395]" country="Portugal" name="Acores" pageId="140" pageNumber="141">Azores</collectingRegion>
EEZ (
<emphasis id="FB95BF1604DCFF5FFD7EFEBC755E11BC" box="[730,844,369,395]" italics="true" pageId="140" pageNumber="141">G.O. Sars</emphasis>
,
<collectionCode id="AFF0FBC104DCFF5FFCF8FEBC75CE11BC" box="[860,988,369,395]" pageId="140" pageNumber="141">
<emphasis id="FB95BF1604DCFF5FFCF8FEBC75CE11BC" box="[860,988,369,395]" italics="true" pageId="140" pageNumber="141">MARECO</emphasis>
</collectionCode>
cruise, sta 42/368:
<geoCoordinate id="ACD505C304DCFF5FFB64FEBC733111BC" box="[1216,1315,369,395]" degrees="42" direction="north" minutes="48" orientation="latitude" pageId="140" pageNumber="141" precision="925" value="42.8">42°48N</geoCoordinate>
,
<geoCoordinate id="ACD505C304DCFF5FFA95FEBC738A11BC" box="[1329,1432,369,395]" degrees="29" direction="west" minutes="38" orientation="longitude" pageId="140" pageNumber="141" precision="925" value="-29.633333">29°38W</geoCoordinate>
,
<quantity id="0E19CEE104DCFF5FFF33FE5977561198" box="[151,324,404,431]" metricMagnitude="0" metricUnit="m" metricValue="2.085" metricValueMax="2.107" metricValueMin="2.063" pageId="140" pageNumber="141" unit="m" value="2.085" valueMax="2.107" valueMin="2.063">
<geoCoordinate id="ACD505C304DCFF5FFF33FE5976C71199" box="[151,213,404,430]" degrees="2,063" direction="north" orientation="latitude" pageId="140" pageNumber="141" precision="55" value="2.063">2,063</geoCoordinate>
<geoCoordinate id="ACD505C304DCFF5FFF72FE5977311199" box="[214,291,404,430]" degrees="2,107" direction="west" orientation="longitude" pageId="140" pageNumber="141" precision="55" value="-2.107">2,107</geoCoordinate>
m
</quantity>
). Additionally, on reviewing the genus
<taxonomicName id="0EE1188704DCFF5FFD5EFE59757D1199" authorityName="Gebruk" authorityYear="1988" box="[762,879,404,430]" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="140" pageNumber="141" phylum="Echinodermata" rank="genus">
<emphasis id="FB95BF1604DCFF5FFD5EFE59757D1199" box="[762,879,404,430]" italics="true" pageId="140" pageNumber="141">Penilpidia</emphasis>
</taxonomicName>
with a redescription of its
<typeStatus id="165ADDA604DCFF5FFB3EFE5872D81198" box="[1178,1226,405,431]" pageId="140" pageNumber="141">type</typeStatus>
species
<taxonomicName id="0EE1188704DCFF5FFA8DFE58776811E4" authority="(Marenzeller, 1893)" baseAuthorityName="Marenzeller" baseAuthorityYear="1893" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="140" pageNumber="141" phylum="Echinodermata" rank="species" species="ludwigi">
<emphasis id="FB95BF1604DCFF5FFA8DFE5873281199" box="[1321,1338,405,430]" italics="true" pageId="140" pageNumber="141">P</emphasis>
.
<emphasis id="FB95BF1604DCFF5FFAECFE59738E1199" box="[1352,1436,404,430]" italics="true" pageId="140" pageNumber="141">ludwigi</emphasis>
(
<bibRefCitation id="AD701EF504DCFF5FFF3BFE75776011E4" author="Marenzeller" box="[159,370,440,467]" firstAuthor="Marenzeller" pageId="140" pageNumber="141" pagination="1 - 32" refId="ref189554" refString="Marenzeller, E. von (1893) Contribution a l'etude des Holothuries de l'Atlantique N'ord (Golfe de Gascogne, Iles Acores). Resultats Campagnes Scientifiques du Prince de Monaco, 6, 1 - 32. https: // doi. org / 10.5962 / bhl. title. 11304" type="journal article" year="1893">Marenzeller, 1893</bibRefCitation>
)
</taxonomicName>
,
<bibRefCitation id="AD701EF504DCFF5FFE2CFE75746011E4" author="Gebruk" box="[392,626,440,467]" etAl="et al." firstAuthor="Gebruk" pageId="140" pageNumber="141" pagination="1029 - 1036" refId="ref182065" refString="Gebruk, A. V., Rogacheva, A. V., Pawson, D. L., Hamel, J. - F., Macisaac, K. G. &amp; Mercier, A. (2013) Penilidia desbarresi sp. nov. (Echinodermata: Holothuroidea: Elasipodida) from the upper slope of Newfoundland and re-description of P. ludwigi (von Marenzeller, 1893). Marine Biology Research, 9 (10), 1029 - 1036. https: // doi. org / 10.1080 / 17451000.2013.793810" type="journal article" year="2013">
Gebruk
<emphasis id="FB95BF1604DCFF5FFE41FE74740A11E5" box="[485,536,440,466]" italics="true" pageId="140" pageNumber="141">et al</emphasis>
. (2013)
</bibRefCitation>
recently concluded that due to differences in ossicles composition and morphology,
<taxonomicName id="0EE1188704DCFF5FFE88FE1077C411C1" authorityName="$ Gebruk" authorityYear="2008" box="[300,470,476,502]" class="Holothuroidea" family="Elpidiidae" genus="Penilpidia" kingdom="Animalia" order="Elasipodida" pageId="140" pageNumber="141" phylum="Echinodermata" rank="species" species="midatlantica">
<emphasis id="FB95BF1604DCFF5FFE88FE1077C411C1" box="[300,470,476,502]" italics="true" pageId="140" pageNumber="141">P. midatlantica</emphasis>
</taxonomicName>
should be assigned to a new genus.
</paragraph>
</subSubSection>
</treatment>
</document>