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<document id="2826AFED3624678E1CF0BCF4334A5B1E" ID-DOI="10.5281/zenodo.212330" ID-GBIF-Dataset="3c0fa1a5-f60b-4952-8548-50dcc16730a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="212330" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460493992240" checkinUser="plazi" docAuthor="Sun, Ning, Li, Bin &amp; Tu, Lihong" docDate="2012" docId="412787F77202FFBCFF2758900A7D8C83" docLanguage="en" docName="zt03358p054.pdf" docOrigin="Zootaxa 3358" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ternatus malleatus Sun, Li &amp; Tu, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="BD1EFF8F7210FFAFFFB05D1409688D77" masterDocTitle="Te rn atu s, a new spider genus from China with a cladistic analysis and comments on its phylogenetic placement (Araneae: Linyphiidae)" masterLastPageNumber="54" masterPageNumber="28" pageNumber="46" updateTime="1698310617636" updateUser="plazi">
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<mods:title id="E60346101040E6FB0A96CA42938F4C2C">Te rn atu s, a new spider genus from China with a cladistic analysis and comments on its phylogenetic placement (Araneae: Linyphiidae)</mods:title>
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<mods:namePart id="0E198A596E14AFF552FF5A9A877C8F24">Sun, Ning</mods:namePart>
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<mods:namePart id="46C1693B7B3083C309D0392EF40E3FE3">Li, Bin</mods:namePart>
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<mods:namePart id="590FF04A24ADA8C377D14258BEB0D325">Tu, Lihong</mods:namePart>
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<mods:date id="380BB83BFFCA8B33C089257EA49BB478">2012</mods:date>
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<treatment id="412787F77202FFBCFF2758900A7D8C83" ID-DOI="http://doi.org/10.5281/zenodo.6180847" ID-GBIF-Taxon="119520317" ID-Zenodo-Dep="6180847" LSID="urn:lsid:plazi:treatment:412787F77202FFBCFF2758900A7D8C83" httpUri="http://treatment.plazi.org/id/412787F77202FFBCFF2758900A7D8C83" lastPageId="19" lastPageNumber="47" pageId="18" pageNumber="46">
<subSubSection id="8194656A7202FFBDFF27589008B288B7" pageId="18" pageNumber="46" type="nomenclature">
<paragraph id="C93136E17202FFBDFF2758900B6C88EB" blockId="18.[151,516,1411,1472]" box="[151,516,1411,1436]" pageId="18" pageNumber="46">
<heading id="9279818D7202FFBDFF2758900B6C88EB" bold="true" box="[151,516,1411,1436]" fontSize="10" level="2" pageId="18" pageNumber="46" reason="2">
<emphasis id="FBFAEAF37202FFBDFF2758900B6C88EB" bold="true" box="[151,516,1411,1436]" pageId="18" pageNumber="46">
<taxonomicName id="0E8E4D627202FFBDFF275890081C88EB" ID-CoL="55H93" box="[151,372,1411,1436]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="malleatus" status="sp. nov.">
<emphasis id="FBFAEAF37202FFBDFF275890081C88EB" bold="true" box="[151,372,1411,1436]" italics="true" pageId="18" pageNumber="46">Ternatus malleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887202FFBDFECB58900B6C88EB" box="[379,516,1411,1436]" pageId="18" pageNumber="46" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C93136E17202FFBDFF2758BC08B288B7" blockId="18.[151,516,1411,1472]" box="[151,474,1448,1472]" pageId="18" pageNumber="46">
<figureCitation id="51B52A647202FFBDFF2758BC098B88B7" box="[151,227,1448,1472]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1834]" captionTargetBox="[151,1435,493,1795]" captionTargetId="figure@2.[151,1436,492,1803]" captionTargetPageId="2" captionText="FIGURE 1. Ternatus malleatus n. sp. A, male, dorsal; B, female, dorsal; C, male, lateral; D, female epigynum, ventral; E, male palp, retrolateral; F, male palp, prolateral." httpUri="https://zenodo.org/record/212331/files/figure.png" pageId="18" pageNumber="46">Figs. 1</figureCitation>
,
<figureCitation id="51B52A647202FFBDFF4158BC086988B7" box="[241,257,1448,1472]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1962]" captionTargetBox="[211,1375,183,1901]" captionTargetId="figure@4.[211,1379,177,1913]" captionTargetPageId="4" captionText="FIGURE 3. Ternatus malleatus n. sp. A, male prosoma; B, male palp, prolateral; C, male palp, retrolateral; D, tibia of right palp, retrolateral; E, male palp with embolic division removed, ventral; F, female tracheal system; G, embolic division, anterior; H, male chelicera, ventral." httpUri="https://zenodo.org/record/212333/files/figure.png" pageId="18" pageNumber="46">3</figureCitation>
,
<figureCitation id="51B52A647202FFBDFEBD58BC087288B7" box="[269,282,1448,1472]" captionStart="FIGURE 5" captionStartId="6.[151,250,1398,1421]" captionTargetBox="[158,1435,184,1397]" captionTargetId="figure@6.[151,1436,177,1400]" captionTargetPageId="6" captionText="FIGURE 5. Female epigynum. A C, Ternatus malleatus n. sp. A, ventral; B, ventral, cleared, arrow indicates lateral loop of copulatory groove; C, dorsal, cleared. D F, Ternatus siculus n. sp. D, ventral; E, ventral, cleared; F, dorsal, cleared." httpUri="https://zenodo.org/record/212335/files/figure.png" pageId="18" pageNumber="46">5</figureCitation>
AC, 67, 8AD
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBDFF2758E408688BEF" pageId="18" pageNumber="46" type="etymology">
<paragraph id="C93136E17202FFBDFF2758E40A4B8B5B" blockId="18.[151,1436,1520,1976]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF2758E408768B7F" bold="true" box="[151,286,1520,1544]" pageId="18" pageNumber="46">Etymology.</emphasis>
The species name comes from the Latin word
<taxonomicName id="0E8E4D627202FFBDFC9C58E40AF18B7F" box="[812,921,1520,1544]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="malleatus">
<emphasis id="FBFAEAF37202FFBDFC9C58E40AF18B7F" box="[812,921,1520,1544]" italics="true" pageId="18" pageNumber="46">malleatus</emphasis>
</taxonomicName>
(“wrought or beaten with a hammer”), in reference to the shape of the male prolateral tibial apophysis.
</paragraph>
<paragraph id="C93136E17202FFBDFF795B2C08688BEF" blockId="18.[151,1436,1520,1976]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795B2C087B8B27" bold="true" box="[201,275,1592,1616]" pageId="18" pageNumber="46">
<typeStatus id="163588437202FFBDFF795B2C08658B27" box="[201,269,1592,1616]" pageId="18" pageNumber="46">Types</typeStatus>
.
</emphasis>
Male
<typeStatus id="163588437202FFBDFEEE5B2C08AC8B27" box="[350,452,1592,1616]" pageId="18" pageNumber="46" type="holotype">holotype</typeStatus>
: Xiaojialu Village, Hongtan Station, Huaping National Nature Reserve, Guangxi Prov.,
<geoCoordinate id="ACBA50267202FFBDFF275B48084A8B03" box="[151,290,1628,1652]" direction="north" orientation="latitude" pageId="18" pageNumber="46" precision="1" value="25.60931">N25.60931°</geoCoordinate>
,
<geoCoordinate id="ACBA50267202FFBDFE9E5B4808AC8B03" box="[302,452,1628,1652]" direction="east" orientation="longitude" pageId="18" pageNumber="46" precision="1" value="109.95181">E109.95181°</geoCoordinate>
, Alt.
<quantity id="0E769B047202FFBDFDB35B480B2D8B03" box="[515,581,1628,1652]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="46" unit="m" value="850.0">850m</quantity>
, Coll. Lihong Tu,
<date id="BD3010217202FFBDFCA55B480ACC8B03" box="[789,932,1628,1652]" pageId="18" pageNumber="46" value="2007-11-03">3 Nov. 2007</date>
.
<typeStatus id="163588437202FFBDFC1F5B490D488B02" box="[943,1056,1629,1653]" pageId="18" pageNumber="46" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="DF88FD687202FFBDFB9B5B480DED8B03" box="[1067,1157,1628,1652]" pageId="18" pageNumber="46" type="male">2 males</specimenCount>
,
<specimenCount id="DF88FD687202FFBDFB215B480C698B03" box="[1169,1281,1628,1652]" pageId="18" pageNumber="46" type="female">4 females</specimenCount>
, same data as
<typeStatus id="163588437202FFBDFF275B9409958BEF" box="[151,253,1664,1688]" pageId="18" pageNumber="46" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBDFF795BB7082E8A50" pageId="18" pageNumber="46" type="diagnosis">
<paragraph id="C93136E17202FFBDFF795BB7082E8A50" blockId="18.[151,1436,1520,1976]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795BB7082B8BCB" bold="true" box="[201,323,1699,1724]" pageId="18" pageNumber="46">Diagnosis.</emphasis>
Males of the two new species can be easily distinguished by the prolateral tibial apophysis stout and dentate, strongly sclerotized in
<taxonomicName id="0E8E4D627202FFBDFD985BDD0BD98B97" box="[552,689,1736,1760]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="malleatus">
<emphasis id="FBFAEAF37202FFBDFD985BDD0B5E8B97" box="[552,566,1737,1760]" italics="true" pageId="18" pageNumber="46">T</emphasis>
.
<emphasis id="FBFAEAF37202FFBDFDF45BDC0BD98B97" box="[580,689,1736,1760]" italics="true" pageId="18" pageNumber="46">malleatus</emphasis>
</taxonomicName>
(
<figureCitation id="51B52A647202FFBDFD725BDC0A6E8B97" box="[706,774,1736,1760]" captionStart="FIGURE 7" captionStartId="8.[151,250,1927,1950]" captionTargetBox="[165,1424,193,1896]" captionTargetId="figure@8.[151,1436,193,1906]" captionTargetPageId="8" captionText="FIGURE 7. Ternatus malleatus n. sp., male palp. A, prolateral; B, retrolateral; C, anterior; D, palpal tibia, dorsal. E H, embolic division with distal suprategular apophysis. E, ventral, arrow indicates embolic socket; F, anterior, arrow indicates embolic socket; G, anteroectal; H, distal end of embolus, arrow indicates hook on embolus apex." httpUri="https://zenodo.org/record/212337/files/figure.png" pageId="18" pageNumber="46">Fig. 7</figureCitation>
D), but thumb-like without dentils in
<taxonomicName id="0E8E4D627202FFBDFB1C5BDD0C7E8B97" box="[1196,1302,1736,1760]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="siculus">
<emphasis id="FBFAEAF37202FFBDFB1C5BDD0DD18B97" box="[1196,1209,1737,1760]" italics="true" pageId="18" pageNumber="46">T</emphasis>
.
<emphasis id="FBFAEAF37202FFBDFB785BDC0C7E8B97" box="[1224,1302,1736,1760]" italics="true" pageId="18" pageNumber="46">siculus</emphasis>
</taxonomicName>
(
<figureCitation id="51B52A647202FFBDFA965BDC0C108B97" box="[1318,1400,1736,1760]" captionStart="FIGURE 10" captionStartId="11.[151,250,1927,1950]" captionTargetBox="[156,1432,193,1905]" captionTargetId="figure@11.[151,1436,193,1906]" captionTargetPageId="11" captionText="FIGURE 10. Ternatus siculus n. sp., male palp. A, prolateral; B, ventral; C, retrolateral; D, palpal tibial, dorsal; E H, embolic division. E, ventral; F, anterior; G, anteroectal; H, distal end of embolus, arrow indicates hook on embolus apex." httpUri="https://zenodo.org/record/212341/files/figure.png" pageId="18" pageNumber="46">Fig. 10</figureCitation>
D). Females are distinguished by having the median plate longer than wide in
<taxonomicName id="0E8E4D627202FFBDFC555BF90D078A73" box="[997,1135,1772,1796]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="malleatus">
<emphasis id="FBFAEAF37202FFBDFC555BF90A9A8A73" box="[997,1010,1773,1796]" italics="true" pageId="18" pageNumber="46">T</emphasis>
.
<emphasis id="FBFAEAF37202FFBDFBB25BF80D078A73" box="[1026,1135,1772,1796]" italics="true" pageId="18" pageNumber="46">malleatus</emphasis>
</taxonomicName>
, and wider than long in
<taxonomicName id="0E8E4D627202FFBDFA375BF9098D8A5F" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="species" species="siculus">
<emphasis id="FBFAEAF37202FFBDFA375BF90CFC8A73" box="[1415,1428,1773,1796]" italics="true" pageId="18" pageNumber="46">T</emphasis>
.
<emphasis id="FBFAEAF37202FFBDFF275A04098D8A5F" box="[151,229,1808,1832]" italics="true" pageId="18" pageNumber="46">siculus</emphasis>
</taxonomicName>
(
<figureCitation id="51B52A647202FFBDFF445A1B08508A50" box="[244,312,1807,1831]" captionStart="FIGURE 8" captionStartId="9.[151,250,1927,1950]" captionTargetBox="[160,1426,193,1902]" captionTargetId="figure@9.[151,1436,193,1906]" captionTargetPageId="9" captionText="FIGURE 8. Female epigynum. A H, Ternatus malleatus n. sp., A, ventral, arrow indicates opened copulatory groove; B, dorsal; C, inner, arrow indicates lateral loop of copulatory groove; D, caudal, arrow indicates opened copulatory groove. E F, Ternatus siculus n. sp. E, caudal, arrow indicates opened copulatory groove; F, dorsal; G, inner; H, ventral." httpUri="https://zenodo.org/record/212338/files/figure.png" pageId="18" pageNumber="46">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBCFF795A2708248CB8" lastPageId="19" lastPageNumber="47" pageId="18" pageNumber="46" type="description">
<paragraph id="C93136E17202FFBDFF795A2708338A3B" blockId="18.[151,1436,1520,1976]" box="[201,347,1843,1868]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795A2708338A3B" bold="true" box="[201,347,1843,1868]" pageId="18" pageNumber="46">Description.</emphasis>
</paragraph>
<paragraph id="C93136E17202FFBDFF795A430C268ACF" blockId="18.[151,1436,1520,1976]" pageId="18" pageNumber="46">
Male (
<typeStatus id="163588437202FFBDFEA25A4308128A18" box="[274,378,1879,1903]" pageId="18" pageNumber="46" type="holotype">holotype</typeStatus>
). Total length 2.49. Carapace 1.09 long, 0.69 wide. Abdomen 1.42 long, 0.87 wide. Chelicera with six promarginal and five retromarginal teeth (
<figureCitation id="51B52A647202FFBDFD7A5A680A7F8AE3" box="[714,791,1916,1940]" captionStart="FIGURE 3" captionStartId="4.[151,250,1939,1962]" captionTargetBox="[211,1375,183,1901]" captionTargetId="figure@4.[211,1379,177,1913]" captionTargetPageId="4" captionText="FIGURE 3. Ternatus malleatus n. sp. A, male prosoma; B, male palp, prolateral; C, male palp, retrolateral; D, tibia of right palp, retrolateral; E, male palp with embolic division removed, ventral; F, female tracheal system; G, embolic division, anterior; H, male chelicera, ventral." httpUri="https://zenodo.org/record/212333/files/figure.png" pageId="18" pageNumber="46">Figs. 3</figureCitation>
H, 6E). Length of legs: I 4.71 (1.40 + 1.52 + 1.06 + 0.73); II 4.32 (1.47 + 1.36 + 0.87 + 0.62); III 3.29 (1.19 +0.97 + 0.70 + 0.43); IV 4.41 (1.49 + 1.42 + 0.93 + 0.57).
</paragraph>
<paragraph id="C93136E17203FFBCFF795D830A388D80" blockId="19.[151,1437,151,500]" pageId="19" pageNumber="47">
Female (
<typeStatus id="163588437203FFBCFE9B5D8308FB8DD8" box="[299,403,151,175]" pageId="19" pageNumber="47" type="paratype">paratype</typeStatus>
): Total length 2.51. Carapace 1.22 long, 0.81 wide. Abdomen 1.45 long, 0.89 wide. Length of legs: I 5.39 (1.59 + 1.77 + 1.22 + 0.81); II 4.72 (1.49 + 1.40 + 1.09 + 0.74); III 3.56 (1.12 +1.23 + 0.70 + 0.51); IV 4.76 (1.61 + 1.53 + 1.00 + 0.62). Tm I 0.32. Tm IV absent.
</paragraph>
<paragraph id="C93136E17203FFBCFF795C110AB08C6A" blockId="19.[151,1437,151,500]" box="[201,984,260,285]" pageId="19" pageNumber="47">For other somatic and genital characters see description for the genus.</paragraph>
<paragraph id="C93136E17203FFBCFF795C3308568C37" blockId="19.[151,1437,151,500]" box="[201,318,295,320]" pageId="19" pageNumber="47">
<emphasis id="FBFAEAF37203FFBCFF795C3308568C37" bold="true" box="[201,318,295,320]" pageId="19" pageNumber="47">Variation.</emphasis>
</paragraph>
<paragraph id="C93136E17203FFBCFF795C5808248CF0" blockId="19.[151,1437,151,500]" pageId="19" pageNumber="47">Males (n = 3). Total length 2.352.60. Carapace 0.961.22 long, 0.640.72 wide. Abdomen 1.141.52 long, 0.830.91 wide.</paragraph>
<paragraph id="C93136E17203FFBCFF795C8008248CB8" blockId="19.[151,1437,151,500]" pageId="19" pageNumber="47">Females (n = 4). Total length 2.392.63. Carapace 0.931.28 long, 0.790.98 wide. Abdomen 1.171.52 long, 0.760.91 wide.</paragraph>
</subSubSection>
<subSubSection id="8194656A7203FFBCFF795CCF0A7D8C83" box="[201,789,475,500]" pageId="19" pageNumber="47" type="distribution">
<paragraph id="C93136E17203FFBCFF795CCF0A7D8C83" blockId="19.[151,1437,151,500]" box="[201,789,475,500]" pageId="19" pageNumber="47">
<emphasis id="FBFAEAF37203FFBCFF795CCF080A8C83" bold="true" box="[201,354,475,500]" pageId="19" pageNumber="47">Distribution.</emphasis>
<collectingCountry id="B19976717203FFBCFED95CC808C48C83" box="[361,428,476,500]" name="China" pageId="19" pageNumber="47">China</collectingCountry>
(Guangxi) (see map in
<figureCitation id="51B52A647203FFBCFD055CC80A6F8C83" box="[693,775,476,500]" captionStart="FIGURE 16" captionStartId="19.[151,250,1366,1389]" captionTargetBox="[185,1370,564,1327]" captionTargetId="figure@19.[151,1436,539,1342]" captionTargetPageId="19" captionText="FIGURE 16. Collecting localities in China of Ternatus malleatus n. sp. (triangle: Huaping, Guangxi Province) and Ternatus siculus n. sp. (circle: Zhangjiajie, Hunan Province)." httpUri="https://zenodo.org/record/212347/files/figure.png" pageId="19" pageNumber="47">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>