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<document id="9DB578B70B1F4A03B75C10467014C02F" ID-DOI="10.5281/zenodo.212330" ID-GBIF-Dataset="3c0fa1a5-f60b-4952-8548-50dcc16730a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="212330" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460493992240" checkinUser="plazi" docAuthor="Sun, Ning, Li, Bin &amp; Tu, Lihong" docDate="2012" docId="412787F77201FFBDFF275B4108B78847" docLanguage="en" docName="zt03358p054.pdf" docOrigin="Zootaxa 3358" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ternatus Sun, Li &amp; Tu, 2012, new genus" docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="BD1EFF8F7210FFAFFFB05D1409688D77" masterDocTitle="Te rn atu s, a new spider genus from China with a cladistic analysis and comments on its phylogenetic placement (Araneae: Linyphiidae)" masterLastPageNumber="54" masterPageNumber="28" pageNumber="45" updateTime="1698310617636" updateUser="plazi">
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<mods:title id="CDE2B5A7231038147BDE8593AA8284C0">Te rn atu s, a new spider genus from China with a cladistic analysis and comments on its phylogenetic placement (Araneae: Linyphiidae)</mods:title>
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<mods:namePart id="8ED0B11A5FB6BCBA0C1502851FAE1600">Sun, Ning</mods:namePart>
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<mods:namePart id="157447DDD489464E48DA5F534E6D4069">Li, Bin</mods:namePart>
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<mods:namePart id="ECF8575070E0C8CD5EEA7AD4A6A8FCCA">Tu, Lihong</mods:namePart>
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<treatment id="412787F77201FFBDFF275B4108B78847" ID-DOI="http://doi.org/10.5281/zenodo.6180845" ID-GBIF-Taxon="119520315" ID-Zenodo-Dep="6180845" LSID="urn:lsid:plazi:treatment:412787F77201FFBDFF275B4108B78847" httpUri="http://treatment.plazi.org/id/412787F77201FFBDFF275B4108B78847" lastPageId="18" lastPageNumber="46" pageId="17" pageNumber="45">
<subSubSection id="8194656A7201FFBEFF275B4108E18B18" box="[151,393,1621,1647]" pageId="17" pageNumber="45" type="nomenclature">
<paragraph id="C93136E17201FFBEFF275B4108E18B18" blockId="17.[151,393,1621,1647]" box="[151,393,1621,1647]" pageId="17" pageNumber="45">
<heading id="9279818D7201FFBEFF275B4108E18B18" bold="true" box="[151,393,1621,1647]" fontSize="11" level="1" pageId="17" pageNumber="45" reason="1">
<emphasis id="FBFAEAF37201FFBEFF275B4108E18B18" bold="true" box="[151,393,1621,1647]" pageId="17" pageNumber="45">
<taxonomicName id="0E8E4D627201FFBEFF275B4109978B18" ID-CoL="63V8T" ID-ENA="1663313" box="[151,255,1621,1647]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="45" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBFAEAF37201FFBEFF275B4109978B18" bold="true" box="[151,255,1621,1647]" italics="true" pageId="17" pageNumber="45">Ternatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887201FFBEFEB85B4108E18B18" box="[264,393,1621,1647]" pageId="17" pageNumber="45" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="8194656A7201FFBEFF275B890D738B81" pageId="17" pageNumber="45" type="materials_examined">
<paragraph id="C93136E17201FFBEFF275B890B0E8BC6" blockId="17.[151,614,1691,1714]" box="[151,614,1691,1714]" pageId="17" pageNumber="45">
<typeStatus id="163588437201FFBEFF275B8909A08BC5" box="[151,200,1693,1714]" pageId="17" pageNumber="45">Type</typeStatus>
species:
<taxonomicName id="0E8E4D627201FFBEFE965B89088F8BC5" box="[294,487,1692,1714]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="45" phylum="Arthropoda" rank="species" species="malleatus" status="sp. nov.">
<emphasis id="FBFAEAF37201FFBEFE965B89088F8BC5" box="[294,487,1692,1714]" italics="true" pageId="17" pageNumber="45">Ternatus malleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887201FFBEFE5C5B890B0E8BC6" box="[492,614,1691,1714]" pageId="17" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="C93136E17201FFBEFF275BCA0D738B81" blockId="17.[151,1436,1758,1998]" box="[151,1051,1758,1783]" pageId="17" pageNumber="45">
<emphasis id="FBFAEAF37201FFBEFF275BCA085E8B80" bold="true" box="[151,310,1758,1783]" pageId="17" pageNumber="45">Composition.</emphasis>
Two species,
<taxonomicName id="0E8E4D627201FFBEFE645BF40BC28B81" box="[468,682,1758,1783]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="45" phylum="Arthropoda" rank="species" species="malleatus" status="sp. nov.">
<emphasis id="FBFAEAF37201FFBEFE645BF40BC28B81" box="[468,682,1758,1783]" italics="true" pageId="17" pageNumber="45">Ternatus malleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887201FFBEFD005BCB0B848B81" box="[688,748,1759,1782]" pageId="17" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="0E8E4D627201FFBEFC935BF40AB18B81" box="[803,985,1758,1783]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="45" phylum="Arthropoda" rank="species" species="siculus" status="sp. nov.">
<emphasis id="FBFAEAF37201FFBEFC935BF40AB18B81" box="[803,985,1758,1783]" italics="true" pageId="17" pageNumber="45">Ternatus siculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887201FFBEFC6F5BCB0D738B81" box="[991,1051,1759,1782]" pageId="17" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8194656A7201FFBEFF795A170A6C8A49" pageId="17" pageNumber="45" type="etymology">
<paragraph id="C93136E17201FFBEFF795A170A6C8A49" blockId="17.[151,1436,1758,1998]" pageId="17" pageNumber="45">
<emphasis id="FBFAEAF37201FFBEFF795A1708278A6C" bold="true" box="[201,335,1795,1819]" pageId="17" pageNumber="45">Etymology.</emphasis>
The generic name, masculine in gender, is derived from the Latin
<emphasis id="FBFAEAF37201FFBEFB815A100DE88A6C" box="[1073,1152,1796,1819]" italics="true" pageId="17" pageNumber="45">ternate</emphasis>
(“consisting of three”) in reference to the three tibial apophyses of the male palp.
</paragraph>
</subSubSection>
<subSubSection id="8194656A7201FFBEFF795A5E0DF08AB9" pageId="17" pageNumber="45" type="diagnosis">
<paragraph id="C93136E17201FFBEFF795A5E0DF08AB9" blockId="17.[151,1436,1758,1998]" pageId="17" pageNumber="45">
<emphasis id="FBFAEAF37201FFBEFF795A5E082B8A14" bold="true" box="[201,323,1866,1891]" pageId="17" pageNumber="45">Diagnosis.</emphasis>
Males of
<taxonomicName id="0E8E4D627201FFBEFE045A580B7D8A14" box="[436,533,1868,1891]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37201FFBEFE045A580B7D8A14" box="[436,533,1868,1891]" italics="true" pageId="17" pageNumber="45">Ternatus</emphasis>
</taxonomicName>
species can be distinguished from other erigonines by three well developed palpal tibial apophyses, of which the retrolateral one is furnished with modified macrosetae (
<figureCitation id="51B52A647201FFBEFB1F5A7A0D948AF1" box="[1199,1276,1902,1926]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1834]" captionTargetBox="[151,1435,493,1795]" captionTargetId="figure@2.[151,1436,492,1803]" captionTargetPageId="2" captionText="FIGURE 1. Ternatus malleatus n. sp. A, male, dorsal; B, female, dorsal; C, male, lateral; D, female epigynum, ventral; E, male palp, retrolateral; F, male palp, prolateral." httpUri="https://zenodo.org/record/212331/files/figure.png" pageId="17" pageNumber="45">Figs. 1</figureCitation>
E, 2E, 7AB, 10AD). Females are diagnosed by having the copulatory grooves forming lateral loops before entering the spermathecae (
<figureCitation id="51B52A647201FFBEFEA05AA208348AB9" box="[272,348,1974,1998]" captionStart="FIGURE 5" captionStartId="6.[151,250,1398,1421]" captionTargetBox="[158,1435,184,1397]" captionTargetId="figure@6.[151,1436,177,1400]" captionTargetPageId="6" captionText="FIGURE 5. Female epigynum. A C, Ternatus malleatus n. sp. A, ventral; B, ventral, cleared, arrow indicates lateral loop of copulatory groove; C, dorsal, cleared. D F, Ternatus siculus n. sp. D, ventral; E, ventral, cleared; F, dorsal, cleared." httpUri="https://zenodo.org/record/212335/files/figure.png" pageId="17" pageNumber="45">Figs. 5</figureCitation>
BC, 5EF, 8C, 8G), with their distal part opened (
<figureCitation id="51B52A647201FFBEFC285AA20A8B8AB9" box="[920,995,1974,1998]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1834]" captionTargetBox="[151,1435,493,1795]" captionTargetId="figure@2.[151,1436,492,1803]" captionTargetPageId="2" captionText="FIGURE 1. Ternatus malleatus n. sp. A, male, dorsal; B, female, dorsal; C, male, lateral; D, female epigynum, ventral; E, male palp, retrolateral; F, male palp, prolateral." httpUri="https://zenodo.org/record/212331/files/figure.png" pageId="17" pageNumber="45">Figs. 1</figureCitation>
D, 2D, 8A, 8E).
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBDFF795D830B6F8E9B" pageId="18" pageNumber="46" type="description">
<paragraph id="C93136E17202FFBDFF795D830C0E8CDB" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795D8308328DC7" bold="true" box="[201,346,151,176]" pageId="18" pageNumber="46">Description.</emphasis>
Small, male total length 2.352.60, female 2.382.67. Carapace bright orange to red, abdomen black. Male cephalic region raised with lateral sulci on sides, several setae arise from cephalic pit (
<figureCitation id="51B52A647202FFBDFB535DA80C588DA3" box="[1251,1328,188,212]" captionStart="FIGURE 6" captionStartId="7.[151,250,1932,1955]" captionTargetBox="[160,1422,195,1902]" captionTargetId="figure@7.[151,1436,195,1909]" captionTargetPageId="7" captionText="FIGURE 6. Ternatus malleatus n. sp. A G, prosoma. A, male, lateral; B, female, lateral; C, male, dorsal, arrow indicates sulci; D, detail of A; E, female, frontal; F, detail of E (area in the box), arrow indicates dentations on chelicera; G, female chelicerae, ectal; H, male posterior lateral spinneret." httpUri="https://zenodo.org/record/212336/files/figure.png" pageId="18" pageNumber="46">Figs. 6</figureCitation>
A, 6CD, 9A). AMEs smaller than other eyes; AER recurved, AME-AME shorter than AMEd, AME-ALE longer, PER straight, PME-PLE equal to PMEd, PME-PME shorter, ALE and PLE juxtaposed (
<figureCitation id="51B52A647202FFBDFB895C100DE98C6B" box="[1081,1153,260,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1834]" captionTargetBox="[151,1435,493,1795]" captionTargetId="figure@2.[151,1436,492,1803]" captionTargetPageId="2" captionText="FIGURE 1. Ternatus malleatus n. sp. A, male, dorsal; B, female, dorsal; C, male, lateral; D, female epigynum, ventral; E, male palp, retrolateral; F, male palp, prolateral." httpUri="https://zenodo.org/record/212331/files/figure.png" pageId="18" pageNumber="46">Figs. 1</figureCitation>
AC, 2AC). Chelicerae with mesal surface dentate (
<figureCitation id="51B52A647202FFBDFE605C330B768C48" box="[464,542,295,319]" captionStart="FIGURE 6" captionStartId="7.[151,250,1932,1955]" captionTargetBox="[160,1422,195,1902]" captionTargetId="figure@7.[151,1436,195,1909]" captionTargetPageId="7" captionText="FIGURE 6. Ternatus malleatus n. sp. A G, prosoma. A, male, lateral; B, female, lateral; C, male, dorsal, arrow indicates sulci; D, detail of A; E, female, frontal; F, detail of E (area in the box), arrow indicates dentations on chelicera; G, female chelicerae, ectal; H, male posterior lateral spinneret." httpUri="https://zenodo.org/record/212336/files/figure.png" pageId="18" pageNumber="46">Figs. 6</figureCitation>
EF). Macrosetae on legs absent. Tm I 0.320.34, Tm IV absent. Female palpal claw absent. Tracheal system desmitracheate. Spinnerets (
<figureCitation id="51B52A647202FFBDFC855C580AEE8C13" box="[821,902,332,356]" captionStart="FIGURE 6" captionStartId="7.[151,250,1932,1955]" captionTargetBox="[160,1422,195,1902]" captionTargetId="figure@7.[151,1436,195,1909]" captionTargetPageId="7" captionText="FIGURE 6. Ternatus malleatus n. sp. A G, prosoma. A, male, lateral; B, female, lateral; C, male, dorsal, arrow indicates sulci; D, detail of A; E, female, frontal; F, detail of E (area in the box), arrow indicates dentations on chelicera; G, female chelicerae, ectal; H, male posterior lateral spinneret." httpUri="https://zenodo.org/record/212336/files/figure.png" pageId="18" pageNumber="46">Figs. 6</figureCitation>
H, 9H): ALS with one major ampullate gland spigot and one nubbin; in female PLS basal part of mesal cylindrical gland spigot larger than distal one; in male PLS flagelliform and aggregate gland spigots (“araneoid triplet”) present. Epiandrous fusules absent in males.
</paragraph>
<paragraph id="C93136E17202FFBDFF795CA30B518FF3" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
Male palp (
<figureCitation id="51B52A647202FFBDFEFB5CA308FD8CB8" box="[331,405,439,463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1811,1834]" captionTargetBox="[151,1435,493,1795]" captionTargetId="figure@2.[151,1436,492,1803]" captionTargetPageId="2" captionText="FIGURE 1. Ternatus malleatus n. sp. A, male, dorsal; B, female, dorsal; C, male, lateral; D, female epigynum, ventral; E, male palp, retrolateral; F, male palp, prolateral." httpUri="https://zenodo.org/record/212331/files/figure.png" pageId="18" pageNumber="46">Figs. 1</figureCitation>
EF, 2EF, 3BE, 4BE, 7, 10). Palpal tibia with three apophyses: one distal, one retrolateral, and one prolateral; the retrolateral one furnished with long modified macrosetae. Paracymbium L-shaped. Tegular sac short. Distal suprategular apophysis bowl-shaped. Embolic membrane with long thread-like projections. Embolic division: radix with half rounded anterior radical process and tailpiece; embolus continuous with radix, long with basal socket, the wide basal part forming a spiral circle anterior to palpal bulb, embolic tip with hook covering the opening (
<figureCitation id="51B52A647202FFBDFE625F780B7D8FF3" box="[466,533,620,644]" captionStart="FIGURE 7" captionStartId="8.[151,250,1927,1950]" captionTargetBox="[165,1424,193,1896]" captionTargetId="figure@8.[151,1436,193,1906]" captionTargetPageId="8" captionText="FIGURE 7. Ternatus malleatus n. sp., male palp. A, prolateral; B, retrolateral; C, anterior; D, palpal tibia, dorsal. E H, embolic division with distal suprategular apophysis. E, ventral, arrow indicates embolic socket; F, anterior, arrow indicates embolic socket; G, anteroectal; H, distal end of embolus, arrow indicates hook on embolus apex." httpUri="https://zenodo.org/record/212337/files/figure.png" pageId="18" pageNumber="46">Fig. 7</figureCitation>
H).
</paragraph>
<paragraph id="C93136E17202FFBDFF795F9B0A2E8E40" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
Epigynum (
<figureCitation id="51B52A647202FFBDFEFD5F9B08F48FD0" box="[333,412,655,679]" captionStart="FIGURE 5" captionStartId="6.[151,250,1398,1421]" captionTargetBox="[158,1435,184,1397]" captionTargetId="figure@6.[151,1436,177,1400]" captionTargetPageId="6" captionText="FIGURE 5. Female epigynum. A C, Ternatus malleatus n. sp. A, ventral; B, ventral, cleared, arrow indicates lateral loop of copulatory groove; C, dorsal, cleared. D F, Ternatus siculus n. sp. D, ventral; E, ventral, cleared; F, dorsal, cleared." httpUri="https://zenodo.org/record/212335/files/figure.png" pageId="18" pageNumber="46">Figs. 5</figureCitation>
,
<figureCitation id="51B52A647202FFBDFE195F9B08D18FD0" box="[425,441,655,679]" captionStart="FIGURE 8" captionStartId="9.[151,250,1927,1950]" captionTargetBox="[160,1426,193,1902]" captionTargetId="figure@9.[151,1436,193,1906]" captionTargetPageId="9" captionText="FIGURE 8. Female epigynum. A H, Ternatus malleatus n. sp., A, ventral, arrow indicates opened copulatory groove; B, dorsal; C, inner, arrow indicates lateral loop of copulatory groove; D, caudal, arrow indicates opened copulatory groove. E F, Ternatus siculus n. sp. E, caudal, arrow indicates opened copulatory groove; F, dorsal; G, inner; H, ventral." httpUri="https://zenodo.org/record/212338/files/figure.png" pageId="18" pageNumber="46">8</figureCitation>
). Dorsal plate concave resulting in centrally hollowed epigynum with plump lateral lobe at each side. Slits of epigynal grooves extending from dorsal to ventral surface with distal parts opened; no distinct trails show the location of spermathecae and boundary between fertilization grooves and copulatory grooves on the epigynal surface. Copulatory grooves forming two lateral loops before entering spermathecae. Spermathecae divided into two chambers, only one of them well developed.
</paragraph>
<paragraph id="C93136E17202FFBDFF795E500C698ED3" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795E50083F8E2B" bold="true" box="[201,343,836,860]" pageId="18" pageNumber="46">Monophyly.</emphasis>
<taxonomicName id="0E8E4D627202FFBDFEED5E5108D68E2B" box="[349,446,837,860]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFEED5E5108D68E2B" box="[349,446,837,860]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
monophyly is unambiguously supported by the following six synapomorphies: modified setae on retrolateral tibial apophysis, distal hook on embolic tip, presence of embolic socket, distal part of copulatory groove opened, copulatory grooves forming lateral loops and presence of distal tibial apophysis.
</paragraph>
<paragraph id="C93136E17202FFBDFF795EA40B6F8E9B" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795EA408EF8EBF" bold="true" box="[201,391,943,968]" pageId="18" pageNumber="46">Natural history.</emphasis>
<taxonomicName id="0E8E4D627202FFBDFE3E5EA508878EBF" box="[398,495,945,968]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFE3E5EA508878EBF" box="[398,495,945,968]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
species build small webs at the base of grasses, above the ground. They also can be found in the leaf-litter by sifting.
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBDFF795EE30D628978" box="[201,1034,1015,1040]" pageId="18" pageNumber="46" type="distribution">
<paragraph id="C93136E17202FFBDFF795EE30D628978" blockId="18.[151,1437,151,1328]" box="[201,1034,1015,1040]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF795EE3080A8967" bold="true" box="[201,354,1015,1040]" pageId="18" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="B19976717202FFBDFED95EE308C48978" box="[361,428,1015,1039]" name="China" pageId="18" pageNumber="46">China</collectingCountry>
(Guangxi and Hunan provinces) (see map in
<figureCitation id="51B52A647202FFBDFC1A5EE30A948978" box="[938,1020,1015,1039]" captionStart="FIGURE 16" captionStartId="19.[151,250,1366,1389]" captionTargetBox="[185,1370,564,1327]" captionTargetId="figure@19.[151,1436,539,1342]" captionTargetPageId="19" captionText="FIGURE 16. Collecting localities in China of Ternatus malleatus n. sp. (triangle: Huaping, Guangxi Province) and Ternatus siculus n. sp. (circle: Zhangjiajie, Hunan Province)." httpUri="https://zenodo.org/record/212347/files/figure.png" pageId="18" pageNumber="46">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8194656A7202FFBDFF79590808B78847" pageId="18" pageNumber="46" type="discussion">
<paragraph id="C93136E17202FFBDFF79590808B78847" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="46">
<emphasis id="FBFAEAF37202FFBDFF79590808558943" bold="true" box="[201,317,1052,1076]" pageId="18" pageNumber="46">Remarks.</emphasis>
<taxonomicName id="0E8E4D627202FFBDFEF4590908CD8943" box="[324,421,1053,1076]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FBFAEAF37202FFBDFEF4590908CD8943" box="[324,421,1053,1076]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0C957887202FFBDFE1C5909089F8942" box="[428,503,1053,1077]" pageId="18" pageNumber="46" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="0E8E4D627202FFBDFD9E59090BF68943" box="[558,670,1053,1076]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFD9E59090BF68943" box="[558,670,1053,1076]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
, are sister groups. They can be distinguished by the three palpal tibial apophyses in adult males: distal tibial apophysis in
<taxonomicName id="0E8E4D627202FFBDFD4F59550A08892F" box="[767,864,1089,1112]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFD4F59550A08892F" box="[767,864,1089,1112]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
arising from the anterior membranous area of palpal tibia, absent in
<taxonomicName id="0E8E4D627202FFBDFECA59710882890B" box="[378,490,1125,1148]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFECA59710882890B" box="[378,490,1125,1148]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
; dorsal tibial apophysis arising from tibial dorsal surface in
<taxonomicName id="0E8E4D627202FFBDFB0259710C49890B" box="[1202,1313,1125,1148]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFB0259710C49890B" box="[1202,1313,1125,1148]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
, absent in
<taxonomicName id="0E8E4D627202FFBDFF27599D099089D7" box="[151,248,1161,1184]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFF27599D099089D7" box="[151,248,1161,1184]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
; retrolateral tibial apophysis well developed and furnished with long modified macrosetae in
<taxonomicName id="0E8E4D627202FFBDFA84599D0CFB89D7" box="[1332,1427,1161,1184]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFA84599D0CFB89D7" box="[1332,1427,1161,1184]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
, small in
<taxonomicName id="0E8E4D627202FFBDFF4659B9080E89B3" box="[246,358,1197,1220]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFF4659B9080E89B3" box="[246,358,1197,1220]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
; prolateral tibial apophysis long, extending distally and tightly covering over cymbium in
<taxonomicName id="0E8E4D627202FFBDFAE459B909A1899F" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFAE459B909A1899F" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
, free in
<taxonomicName id="0E8E4D627202FFBDFE9859C508E1899F" box="[296,393,1233,1256]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFE9859C508E1899F" box="[296,393,1233,1256]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
. Females of the two genera are different in the distal parts of copulatory grooves: opened in
<taxonomicName id="0E8E4D627202FFBDFF2759E10990887B" box="[151,248,1269,1292]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFF2759E10990887B" box="[151,248,1269,1292]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
, duct-like with slits opening on the epigynal surface in
<taxonomicName id="0E8E4D627202FFBDFCD159E10AB8887B" box="[865,976,1269,1292]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFCD159E10AB8887B" box="[865,976,1269,1292]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
, and by the tract tracings: lateral loops in
<taxonomicName id="0E8E4D627202FFBDFF27580D09908847" box="[151,248,1305,1328]" class="Arachnida" family="Linyphiidae" genus="Ternatus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFF27580D09908847" box="[151,248,1305,1328]" italics="true" pageId="18" pageNumber="46">Ternatus</emphasis>
</taxonomicName>
, spiral in
<taxonomicName id="0E8E4D627202FFBDFED8580C08B08858" box="[360,472,1304,1327]" class="Arachnida" family="Linyphiidae" genus="Gonatium" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="FBFAEAF37202FFBDFED8580C08B08858" box="[360,472,1304,1327]" italics="true" pageId="18" pageNumber="46">Gonatium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>