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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.196.82598" ID-Pensoft-Pub="1314-2003-196-21" ID-Pensoft-UUID="A221AFB3297756818D82E7B978F2E682" ModsDocID="1314-2003-196-21" checkinTime="1652543785802" checkinUser="pensoft" docAuthor="Nobis, Marcin, Krzempek, Marta, Nowak, Arkadiusz, Gudkova, Polina D. &amp; Klichowska, Ewelina" docDate="2022" docId="15800F4FE7BB5C25B51518301DEE7B5B" docLanguage="en" docName="PhytoKeys 196: 21-47" docOrigin="PhytoKeys 196" docPubDate="2022-05-13" docSource="http://dx.doi.org/10.3897/phytokeys.196.82598" docTitle="Stipa arenosa M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and 2022, stat. nov." docType="treatment" docVersion="1" id="A221AFB3297756818D82E7B978F2E682" lastPageNumber="21" masterDocId="A221AFB3297756818D82E7B978F2E682" masterDocTitle="Resurrection of Stipa tremula and taxonomy of the high-alpine species from the Stipa purpurea complex (Poaceae, Pooideae)" masterLastPageNumber="47" masterPageNumber="21" pageNumber="21" updateTime="1652543785802" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Resurrection of Stipa tremula and taxonomy of the high-alpine species from the Stipa purpurea complex (Poaceae, Pooideae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nobis, Marcin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1594-2418</mods:nameIdentifier>
<mods:affiliation>Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="email">m.nobis@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krzempek, Marta</mods:namePart>
<mods:affiliation>Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nowak, Arkadiusz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8638-0208</mods:nameIdentifier>
<mods:affiliation>Institute of Biology, University of Opole, 45 - 052 Opole, Poland &amp; Botanical Garden, Center for Biological Diversity Conservation, Polish Academy of Sciences, 02 - 976 Warszawa, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gudkova, Polina D.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6537-8018</mods:nameIdentifier>
<mods:affiliation>Research Laboratory ' Herbarium', National Research Tomsk State University, Lenin 36 Ave., 634050 Tomsk, Russia &amp; Institute of Biology, Altai State University, Lenin 61 Ave., 656049, Barnaul, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klichowska, Ewelina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9641-5750</mods:nameIdentifier>
<mods:affiliation>Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="email">ewelina.klichowska@uj.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>196</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>21</mods:start>
<mods:end>47</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.196.82598</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.196.82598</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-196-21</mods:identifier>
<mods:identifier type="Pensoft-UUID">A221AFB3297756818D82E7B978F2E682</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:15800F4FE7BB5C25B51518301DEE7B5B" httpUri="http://treatment.plazi.org/id/15800F4FE7BB5C25B51518301DEE7B5B" lastPageNumber="21" pageId="0" pageNumber="21">
<subSubSection pageId="0" pageNumber="21" type="nomenclature">
<paragraph pageId="0" pageNumber="21">
<taxonomicName LSID="15800F4F-E7BB-5C25-B515-18301DEE7B5B" authority="(Tzvelev) M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" authorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" authorityYear="2022" baseAuthorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="arenosa" status="stat. nov.">Stipa arenosa (Tzvelev) M. Nobis, P.D. Gudkova, Krzempek &amp; Klichowska, comb. and</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="21">stat. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="reference_group">
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea subsp. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="subSpecies" species="purpurea" status="stat. nov." subSpecies="arenosa">Stipa purpurea arenosa</taxonomicName>
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea subsp. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="subSpecies" species="purpurea" status="stat. nov." subSpecies="arenosa">Stipa purpurea subsp. arenosa</taxonomicName>
(Tzvelev) D.F. Cui, Flora Xinjiangensis 6: 307. 1996.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="basionym">
<paragraph pageId="0" pageNumber="21">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea subsp. var. var. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="variety" species="purpurea" subSpecies="var." variety="arenosa">Stipa purpurea var. arenosa</taxonomicName>
Tzvelev, Rastenia
<normalizedToken originalValue="Tsentralnoi">Tsentral'noi</normalizedToken>
Azii po materialam Botanicheskogo Instituta im. V. L. Komarova 4: 60. 1968.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="type protologue">
<paragraph pageId="0" pageNumber="21">Type protologue.</paragraph>
<paragraph pageId="0" pageNumber="21">
Tibet bor.-occid., praemontium bor. jugi
<normalizedToken originalValue="Przevalskii">Przeval'skii</normalizedToken>
, ad 5000 m alt., in steppa
<taxonomicName authorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" authorityYear="2022" baseAuthorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="arenosa">Stipa arenosa</taxonomicName>
, 24.08.1980, V. Roborovski (LE).
<emphasis bold="true" pageId="0" pageNumber="21">Type</emphasis>
: Thibet boreal.-occid., Kuen-Lun, Khr.
<normalizedToken originalValue="Przevalskogo">Przeval'skogo</normalizedToken>
, severnye peredgorya, 24 Aug 1890,
<emphasis italics="true" pageId="0" pageNumber="21">W.J. Roborowski s.n</emphasis>
. (holotype, LE 01010497!).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="description">
<paragraph pageId="0" pageNumber="21">Description.</paragraph>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Perennial plant</emphasis>
, densely tufted, with a few culms and numerous vegetative shoots; culms (14.7-)16.2-39.1(-50.0) cm tall, 1-2(-3)-noded, nodes distributed close together and only in the lowermost part of the culm.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Leaves of vegetative shoots</emphasis>
: sheaths shortly and densely pubescent;
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">ligules</emphasis>
truncate to slightly acute, on the external sheaths (0.5-)1.0-1.8(-2.3) mm long, whereas on the internal sheaths (1.0-)1.5-3.0(-4.5);
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">blades</emphasis>
convolute, green, pale green to glaucous, (4.9-)6.1-11.4(-16.0) cm long, 0.4-0.5 mm in diameter, adaxial surface covered by 0.15-0.2 mm long hairs, abaxial surface glabrous, scabrous or setulose.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Cauline leaves</emphasis>
: lower sheaths sparsely pubescent to almost glabrous, upper sheath glabrous or almost so;
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">ligules</emphasis>
acute, on the lower sheaths (0.7-)1.2-1.9 mm long, on the middle sheaths (1.6-)1.9-3.5(-4.5) and on the upper 2.0-3.45(-4.5);
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">blades</emphasis>
convolute, green or pale green, (2.5-)3.0-6.5(-7.8) cm long, adaxial surface covered with short hairs, abaxial surface glabrous, scabrous to shortly pubescent.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Panicle</emphasis>
(7.6-)8.7-14.4(-17.8) cm long, open, at base enclosed by the sheath of the uppermost leaf or rarely exerted, branches flexuous, setulose or slightly scabrous or glabrous, single or paired.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Glumes</emphasis>
subequal (the lower slightly longer than the upper), purplish, glumes (19.5-)21.0-23.5(-25.0) mm long, narrowly lanceolate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Floret</emphasis>
(lemma + callus) (11.5-)12.0-14.0(-15.0) mm long.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Callus</emphasis>
(2.0-)2.3-3.0(-3.8) mm long, densely pilose on the ventral part, with hairs (0.4-)0.6-0.9(-1.0) mm long, on dorsal part sparsely and shortly pilose with straight hairs (0.5-)0.6-0.8(-0.9) mm long; callus base 0.5-0.8 mm long and 0.15-0.25 mm wide, sharply pointed, scar narrow-elliptic.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Lemma</emphasis>
coriaceous, pale-green, purplish or brownish, covered throughout (from the bottom to top) by dense ascending to appressed hairs 0.2-0.4 mm long; top of lemma surpassed by a ring of unequal hairs 0.4-0.9 mm long.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Awn</emphasis>
(78-)95-112(-130) mm long, bigeniculate, straw-coloured to brownish;
<emphasis bold="true" pageId="0" pageNumber="21">first segment of the awn</emphasis>
(column) (7-)11-22 mm long, twisted, 0.3 mm wide at base, with (1.5-)1.6-2.0(-2.3) mm long hairs,
<emphasis bold="true" pageId="0" pageNumber="21">middle segment of the awn</emphasis>
10-12(-13) mm long, twisted, with (1.7-)1.9-2.4 mm long hairs;
<emphasis bold="true" pageId="0" pageNumber="21">terminal segment of the awn</emphasis>
(seta) arcuate or flexuous (62-)68-89(-98) mm long and, hairs longer than those on columns, (2.1-)2.3-2.5(-3.0) mm long, gradually decreasing in length towards apex.
<emphasis bold="true" italics="true" pageId="0" pageNumber="21">Palea</emphasis>
equalling lemma in length.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="habitat">
<paragraph pageId="0" pageNumber="21">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="21">High mountain steppes, semi-deserts, 3500-5000 m. a.s.l.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="distribution">
<paragraph pageId="0" pageNumber="21">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="21">
China: southern Xinjiang, Qinghai, Gansu, north-western Xizang; Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The distribution range of Stipa purpurea (purple backgrounds and points), S. tremula (blue backgrounds and points) and S. arenosa (yellow points)." figureDoi="10.3897/phytokeys.196.82598.figure3" httpUri="https://binary.pensoft.net/fig/683652" pageId="0" pageNumber="21">3</figureCitation>
(
<bibRefCitation author="Tzvelev, NN" editor="Grubov, VI" journalOrPublisher="Po materialam Botanicheskogo Instituta im. Komarova VL (Plantae Asiae Centralis, secus materies Instituti botanici nomine Komarovii VL), 4. Nauka, Leningrad" pageId="0" pageNumber="21" pagination="1 - 243" refId="B61" refString="Tzvelev, NN, 1968. Zlaki (Gramineae). In: Grubov, VI, Ed., Rastieniya Centralnoi Azii. Po materialam Botanicheskogo Instituta im. Komarova VL (Plantae Asiae Centralis, secus materies Instituti botanici nomine Komarovii VL), 4. Nauka, Leningrad: 1 - 243" title="Zlaki (Gramineae)." volumeTitle="Rastieniya Centralnoi Azii." year="1968">Tzvelev 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="selected specimen examined">
<paragraph pageId="0" pageNumber="21">Selected specimen examined.</paragraph>
<paragraph pageId="0" pageNumber="21">
<emphasis bold="true" pageId="0" pageNumber="21">China: Qinghai Province</emphasis>
: Kuen-Lun, Dolina r.
<normalizedToken originalValue="Sharagol">Sharagol'</normalizedToken>
-dzhin, yr. Paidza-Tologoi, 11000 ft alt., pesch.
<normalizedToken originalValue="step">step'</normalizedToken>
, 11 Jul 1894,
<emphasis italics="true" pageId="0" pageNumber="21">W.J. Roborowski 318</emphasis>
(LE 01010495 [label 2:
<taxonomicName class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="species" species="purpurea">Stipa purpurea</taxonomicName>
Griseb f. robusta, det. R. Roshevitz; Label 3:
<taxonomicName class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">Stipa</taxonomicName>
kozlovii m. sp. nov. inedit. =
<taxonomicName family="Poaceae" genus="S." kingdom="Plantae" lsidName="S. purpurea" order="Poales" pageId="0" pageNumber="21" rank="species" species="purpurea">S. purpurea</taxonomicName>
var.
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea var. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="variety" species="purpurea" variety="arenosa">Stipa purpurea arenosa</taxonomicName>
m. var. nova, Typus varietis!, XI. 1966, N.N. Tzvelev], LE 01010496, KRA 476855, K, - paratypes);
<emphasis bold="true" pageId="0" pageNumber="21">Xinjiang Province</emphasis>
, Ruoqiang, N of Aqqikkol, on grasslands, ca. 4200 m alt., 21 Aug 1988,
<emphasis italics="true" pageId="0" pageNumber="21">S. Wu, H. Ohba, Y. Wu, Y. Fei 2747</emphasis>
(MOIS 3744710); Xinjiang, Ruoqiang, Yueya River to Aqqikkol, in
<taxonomicName class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">Stipa</taxonomicName>
grassland on gravel-rich flat places, 21 Aug 1988,
<emphasis italics="true" pageId="0" pageNumber="21">S. Wu, H. Ohba, Y. Wu, Y. Fei 2275</emphasis>
(MOIS 5660755);
<emphasis bold="true" pageId="0" pageNumber="21">Gansu Province</emphasis>
, Subei County, alt. 3500 m, 5 Aug 1956,
<emphasis italics="true" pageId="0" pageNumber="21">B-z. Guo s.n</emphasis>
. (PE 707226);
<emphasis bold="true" pageId="0" pageNumber="21">Xizang Province</emphasis>
, 16 km northeast of Shuanghu County, alt. 5000 m, 27 Jul 1976,
<emphasis italics="true" pageId="0" pageNumber="21">Gansu Agricultural University 111</emphasis>
(PE 2029866).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21" type="notes">
<paragraph pageId="0" pageNumber="21">Note.</paragraph>
<paragraph pageId="0" pageNumber="21">
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea subsp. var. var. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="variety" species="purpurea" subSpecies="var." variety="arenosa">Stipa purpurea var. arenosa</taxonomicName>
was described, based on two collections of W.J. Roborowski from Central Asia (
<bibRefCitation author="Tzvelev, NN" editor="Grubov, VI" journalOrPublisher="Po materialam Botanicheskogo Instituta im. Komarova VL (Plantae Asiae Centralis, secus materies Instituti botanici nomine Komarovii VL), 4. Nauka, Leningrad" pageId="0" pageNumber="21" pagination="1 - 243" refId="B61" refString="Tzvelev, NN, 1968. Zlaki (Gramineae). In: Grubov, VI, Ed., Rastieniya Centralnoi Azii. Po materialam Botanicheskogo Instituta im. Komarova VL (Plantae Asiae Centralis, secus materies Instituti botanici nomine Komarovii VL), 4. Nauka, Leningrad: 1 - 243" title="Zlaki (Gramineae)." volumeTitle="Rastieniya Centralnoi Azii." year="1968">Tzvelev 1968</bibRefCitation>
). During our research in 2009-2021, in addition to the holotype, we found four sheets (paratypes) with specimens of
<taxonomicName class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="21">Stipa</emphasis>
</taxonomicName>
collected by Roborowski in 1894. The two of the paratypes preserved in LE, were labeled as the type and isotype of
<taxonomicName authorityName="Nobis &amp; Krzempek &amp; Nowak &amp; Gudkova &amp; Klichowska" authorityYear="2022" baseAuthorityName="M. Nobis, P. D. Gudkova, Krzempek &amp; Klichowska, comb. and" baseAuthorityYear="2022" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa purpurea subsp. var. var. arenosa" order="Poales" pageId="0" pageNumber="21" phylum="Tracheophyta" rank="variety" species="purpurea" subSpecies="var." variety="arenosa">Stipa purpurea var. arenosa</taxonomicName>
by N. Tzvelev in 1966.
</paragraph>
</subSubSection>
</treatment>
</document>