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<document ID-CLB-Dataset="297105" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="c7b986d2-d221-4c24-ae0a-037b5d4d8887" ID-ISBN="0-8018-8221-4" checkinTime="1670905166440" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="04806AAD64EF9E518CEC9382B0AEB95F" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Geomyidae.859-870.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Thomomys (Megascapheus) umbrinus" docType="treatment" docVersion="3" id="B149FDE7D3909CFBF93D8FFBC8EF1D48" lastPageNumber="870" masterDocId="B149FDE7D3909CFBF93D8FFBC8EF1D48" masterDocTitle="Order Rodentia - Family Geomyidae" masterLastPageNumber="870" masterPageNumber="859" pageNumber="870" updateTime="1716775291689" updateUser="ExternalLinkService">
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<mods:title id="503525E0DE8DB484140BEB8DCEFDB1B2">Order Rodentia - Family Geomyidae</mods:title>
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<mods:namePart id="1C3C528F2F3CBE23F26F0EF5BEEA4082">Reeder, DeeAnn</mods:namePart>
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<mods:title id="FC9A9D1794CB3C4C1731DFEC7E2DD260">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<treatment id="04806AAD64EF9E518CEC9382B0AEB95F" ID-GBIF-Taxon="231543105" LSID="urn:lsid:plazi:treatment:04806AAD64EF9E518CEC9382B0AEB95F" httpUri="http://treatment.plazi.org/id/04806AAD64EF9E518CEC9382B0AEB95F" lastPageNumber="870" pageNumber="870" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=12800376">
<subSubSection id="7C5F23A01F3B5057000CC3F1EA449354" pageNumber="870" type="nomenclature">
<paragraph id="EC23AD0721E67FF19D77C865790665AA" pageNumber="870">
<taxonomicName id="2F584E62D50A4EDF9947E5A1B56D5ABF" ID-CoL="7X2SQ" authority="(Richardson 1829)" baseAuthorityName="Richardson" baseAuthorityYear="1829" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="species" species="umbrinus" subGenus="Megascapheus">
<emphasis id="C5401F48A3E2672A1BB3D18C15B6116A" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
(Richardson 1829)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="87A7EFE61DC27F3AF7A6543BD8C86EBE" pageNumber="870" type="reference_group">
<paragraph id="27C110204B46334359F22FA97265518C" pageNumber="870">
<treatmentCitation id="FE4F9943BF981A11C6E44F9BB817AACC" author="Richardson" authority="Richardson 1829" authorityName="Richardson" authorityYear="1829" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" page="202" pageNumber="870" phylum="Chordata" rank="species" species="umbrinus" title="Geomys umbrinus" volumeTitle="Quadrupeds, in: Fauna Boreali-Americana, Vol. 1" year="1829">
<taxonomicName id="FC6D4CA6D1AA1377E926D4C2E27ABDE0" authority="Richardson 1829" authorityName="Richardson" authorityYear="1829" class="Mammalia" family="Geomyidae" genus="Geomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="species" species="umbrinus">
<emphasis id="AA6450FCA46D74F7F030B5275B7C7039" italics="true" pageNumber="870">[Geomys] umbrinus</emphasis>
Richardson 1829
</taxonomicName>
,
<bibCitation id="D257820BE376F36DCA66C997EE03C382" author="Richardson" pageNumber="870" pagination="202" title="Geomys umbrinus" volumeTitle="Quadrupeds, in: Fauna Boreali-Americana, Vol. 1" year="1829">Quadrupeds, in: Fauna Boreali-Americana, Vol. 1: 202</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0F0906C7B2070845A67BC1C4CDE57A88" pageNumber="870" type="materials_examined">
<paragraph id="2874BFE11D1599E92C9A98C2B3F91E10" pageNumber="870">
<emphasis id="3D7BD0535799F608AD393BCF331A5EB8" inLineHeading="true" pageNumber="870">Type Locality:</emphasis>
<materialsCitation id="2FA79277F4774CB24318A00973AF0EE4" ID-GBIF-Occurrence="4867332333" country="Mexico" pageNumber="870" stateProvince="Veracruz de Ignacio de la Llave">
Originally given as Cadadaguios, SW Louisiana, but restricted to
<collectingCountry id="6BC2230B5D1C637A4763A3C1267BB8E4" name="Mexico">Mexico</collectingCountry>
,
<collectingRegion id="A6ABF53BC8C59BAA170FA76399B89FB1" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
(probably
<collectingRegion id="614D786E5B359D67A07995978DFC9B56" country="Mexico" name="Puebla">Puebla</collectingRegion>
), vicinity of Boca del Monte, by
<bibRefCitation id="54A8BBD4AA8128D86A712A86C6749032" author="Bailey, V. E." refId="ref17640" refString="Bailey, V. E. 1906. Identity of Thomomys umbrinus (Richardson). Proceedings of the Biological Society of Washington, 19: 3 - 6." year="1906">Bailey (1906)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5489DA784D647C3E1271CD6C8A1E8FAC" pageNumber="870" type="vernacular_names">
<paragraph id="72E247D5482240B3BA7CD1660D9FF51F" pageNumber="870">
<emphasis id="01C832C7B5B7B6ABC2CACA8403309774" inLineHeading="true" pageNumber="870">Vernacular Names:</emphasis>
<vernacularName id="5877B5CB998266F0CA1B260B26B1200C" language="eng" pageNumber="870">Southern Pocket Gopher</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="FD8E6EA5C052BD4280A2B77B1A6D6257" pageNumber="870" type="child_taxa">
<paragraph id="8CCED59B31DBE21403F989540B8A79FA" pageNumber="870">
<emphasis id="BAB174C431316C1E07202599E823B8C7" inLineHeading="true" pageNumber="870">Subspecies:</emphasis>
:
</paragraph>
<paragraph id="D566DC95B328441B4D469BE65A12E874" pageNumber="870">
Subspecies
<taxonomicName id="5706FE74E664DC883525202E22D88D59" authority="(Richardson 1829)" baseAuthorityName="Richardson" baseAuthorityYear="1829" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="umbrinus">
<emphasis id="E51FF6ABD5DAC1082856C407420C5EC5" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="79B92372E34D02D7C553752B3630C208" italics="true" pageNumber="870">umbrinus</emphasis>
(Richardson 1829)
</taxonomicName>
</paragraph>
<paragraph id="0ABC07F35477C3B7B92640286DDDDFD1" pageNumber="870">
Subspecies
<taxonomicName id="2CDC3923470E2E5AC091BD1FC489B026" authority="Dalquest 1951" authorityName="Dalquest" authorityYear="1951" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="arriagensis">
<emphasis id="D2B21A22462591BB2D6330A4E476EC49" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="1B1292E1580979C0311273EDE16F285E" italics="true" pageNumber="870">arriagensis</emphasis>
Dalquest 1951
</taxonomicName>
</paragraph>
<paragraph id="F5395AF96916EB5470F0CDB0755D1A29" pageNumber="870">
Subspecies
<taxonomicName id="EDED1CECBA437FF7CA07AE294A465629" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="atrodorsalis">
<emphasis id="29A3D0A71DD1D47C57F43E857FDFD291" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="732E712FF305E2729B4FCAF40ED460A2" italics="true" pageNumber="870">atrodorsalis</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="B7EE291E146DF402A770C6B7266472F0" pageNumber="870">
Subspecies
<taxonomicName id="07B8F0370F975DA617C189BEC29D0E85" authority="(J. A. Allen 1898)" baseAuthorityName="J. A. Allen" baseAuthorityYear="1898" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="atrovarius">
<emphasis id="E1A8495CECA135BDA54E3EC33EE54E81" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="AD5236B76D60066AE7F31A7B4E325E35" italics="true" pageNumber="870">atrovarius</emphasis>
(J. A.
<bibRefCitation id="E902281ED29EDF44A55A1137C3518E33" author="Allen, J. A." refId="ref5378" refString="Allen, J. A. 1898. Revision of the chickarees, or North American red squirrels (subgenus Tamiasciurus). Bulletin of the American Museum of Natural History, 10: 249 - 298." year="1898">Allen 1898</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="7B0881A849C49CD118FF0CD7918A4D20" pageNumber="870">
Subspecies
<taxonomicName id="38BD7671265C0757BDB8E610B79B0E1D" authority="Anderson 1972" authorityName="Anderson" authorityYear="1972" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="camargensis">
<emphasis id="4D5C8BC41BAA730BE59812C0366B2470" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="5B27F335EC9B3149C4902FB2FD3A86D9" italics="true" pageNumber="870">camargensis</emphasis>
<bibRefCitation id="6CBB18FF6A0491867E850F90CA856400" author="Anderson, S." refId="ref9154" refString="Anderson, S. 1972. Mammals of Chihuahua: Taxonomy and distribution. Bulletin of the American Museum of Natural History, 148: 149 - 410." year="1972">Anderson 1972</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1BA7B8121D664249A673461C23716855" pageNumber="870">
Subspecies
<taxonomicName id="765C27F98A66E1A826B180B1706F6E83" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="chihuahuae">
<emphasis id="6296EEADCAF44506F753E7B23B61EFC4" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="B54DF7F63279A00B2012E78753839B8F" italics="true" pageNumber="870">chihuahuae</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="2CAD69774EC23AE451118F086F054AE6" pageNumber="870">
Subspecies
<taxonomicName id="8F60FC2B36666D8F9DC5B76B2CD71F05" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="crassidens">
<emphasis id="93B94C1A281FCE5FA3C67F29C17F478D" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="8CECDEFD65E8154063733E6D9786166C" italics="true" pageNumber="870">crassidens</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="C8907EFE8B8FF886177FF99E40BDC100" pageNumber="870">
Subspecies
<taxonomicName id="EDC4E99A9DC63FCE12024F1C78511A97" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="durangi">
<emphasis id="BA057230BE000FCF9C72849F430F381A" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="6E466F31F4D91248EB6A0D2CFCA27D78" italics="true" pageNumber="870">durangi</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="B430AD42EBEA6EFA33B759707AE4F0BD" pageNumber="870">
Subspecies
<taxonomicName id="DA92DBF2BA239EC8922494A9B15736CD" authority="Goldman 1933" authorityName="Goldman" authorityYear="1933" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="emotus">
<emphasis id="90EB511C3D48E11119856FFEB1A99197" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="744CDD840F2A358D4321F2A4AA202A84" italics="true" pageNumber="870">emotus</emphasis>
Goldman 1933
</taxonomicName>
</paragraph>
<paragraph id="52A2F0BE277D3F6C39F7DCA33AD08341" pageNumber="870">
Subspecies
<taxonomicName id="6EE2A22B7AC602500A8E9B456724892D" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="enixus">
<emphasis id="425C65FF3E20EEDD994C752A6FF68BA8" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="4D2BEB1E3A863DBC9FE3EA9D6B340805" italics="true" pageNumber="870">enixus</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="EA0B8734039F28E098B6F1FDC72E62D4" pageNumber="870">
Subspecies
<taxonomicName id="ED4542EE9EC6AE8923F6BAA33EC4F181" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="eximius">
<emphasis id="8420E2D77AD33DE440A4008D54743DFE" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="EFE060897F4331FF1DD1118E87FBC2F2" italics="true" pageNumber="870">eximius</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="B968E6197A733DAEF994FBEDC73E8DFA" pageNumber="870">
Subspecies
<taxonomicName id="9746385FB4AD07FABD8376D2A677D0F7" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="extimus">
<emphasis id="C623136C0369B2C65C6A40A035376DF6" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="82CE65D8AE492F363D9215664784F196" italics="true" pageNumber="870">extimus</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="5F682E001B0F5DFE034A509943197F8E" pageNumber="870">
Subspecies
<taxonomicName id="32BB4FB3B4C30C66613BAAD8BC9F572F" authority="(Merriam 1901)" baseAuthorityName="Merriam" baseAuthorityYear="1901" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="goldmani">
<emphasis id="F1C97F36EA4C1F0D83F23C757CAB3131" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="F34937D13FF35A362305D3599ACCAF72" italics="true" pageNumber="870">goldmani</emphasis>
(Merriam 1901)
</taxonomicName>
</paragraph>
<paragraph id="123B9B02229A773CFCA106E2ECEDABDE" pageNumber="870">
Subspecies
<taxonomicName id="5B004C386D6191083E03264575C51793" authority="Mearns 1897" authorityName="Mearns" authorityYear="1897" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="intermedius">
<emphasis id="3A414848E9EA5328DC76F7404755E52E" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="BE2DBA1B4BEF76FC14F5A2A870C42E12" italics="true" pageNumber="870">intermedius</emphasis>
Mearns 1897
</taxonomicName>
</paragraph>
<paragraph id="28627227CE720899D05F1071334692BF" pageNumber="870">
Subspecies
<taxonomicName id="6A75F8C7C2FB8447A698ED613B72CA90" authority="Anderson 1972" authorityName="Anderson" authorityYear="1972" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="juntae">
<emphasis id="7DD6473C8358B568FD8620E28A97FBE3" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="696F66621A20A4EBBDCBCC6B3350BAE9" italics="true" pageNumber="870">juntae</emphasis>
<bibRefCitation id="66AA0D0A092832949B816100E80A4C4E" author="Anderson, S." refId="ref9154" refString="Anderson, S. 1972. Mammals of Chihuahua: Taxonomy and distribution. Bulletin of the American Museum of Natural History, 148: 149 - 410." year="1972">Anderson 1972</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="7285EBD567496C81F23F20C39C4DE6B0" pageNumber="870">
Subspecies
<taxonomicName id="40C605B097A5760DD13FF2A3F3DA3788" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="madrensis">
<emphasis id="B840641C15EFDDF3F67FD375C4701D5A" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="2EAD1887CC84C007AD3BB05BED4CBA2A" italics="true" pageNumber="870">madrensis</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="E6429B34854B6716A4DAE12FEA0848EB" pageNumber="870">
Subspecies
<taxonomicName id="6AD8ECA70857173C98136F92A0A84597" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="musculus">
<emphasis id="05F7AC4A6FF4B7C4F4F9DDD1B43E1E47" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="56C337C5A210E72B9AFD169F998A16A4" italics="true" pageNumber="870">musculus</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="1A01D29C0D3E79A920A2F1F2879F1A86" pageNumber="870">
Subspecies
<taxonomicName id="3965174D903F35FD4624FA50895B1A91" authority="(Merriam 1901)" baseAuthorityName="Merriam" baseAuthorityYear="1901" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="nelsoni">
<emphasis id="B887A56E5839A6A20C3A1B1B9C844BA6" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="C65ABCDF5298A746EA7B2E911ADAA698" italics="true" pageNumber="870">nelsoni</emphasis>
(Merriam 1901)
</taxonomicName>
</paragraph>
<paragraph id="11C484FC66A67B351BD0383BBF9DE289" pageNumber="870">
Subspecies
<taxonomicName id="23A0B0378CBFF9E911E21454340700C0" authority="Dalquest 1951" authorityName="Dalquest" authorityYear="1951" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="newmani">
<emphasis id="FBDE5D5B90BC321A9DA9574D8DA58B51" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="3B7E954803AD4B96EF83BA8B5D04059A" italics="true" pageNumber="870">newmani</emphasis>
Dalquest 1951
</taxonomicName>
</paragraph>
<paragraph id="DCFA24496AE4F5D93354F9313B32C908" pageNumber="870">
Subspecies
<taxonomicName id="6B6ED9B48F09C83AC17E362845718547" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="parviceps">
<emphasis id="8CA0B6FED249B57CE23E40BE39AED19E" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="BC2BE34B567FD2DF73FBE8B01ED26361" italics="true" pageNumber="870">parviceps</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="8552FCE28D5AC953B4CC42004A74A73F" pageNumber="870">
Subspecies
<taxonomicName id="590C80A7E27345891A334B30BFCCE244" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="potosinus">
<emphasis id="FF7A7445412294D114F8E6B0D1106BF6" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="B5E29B3A0EC109A6FE9058B8E50DC7AB" italics="true" pageNumber="870">potosinus</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="CE8B8DFEBC8E927749721CE075A32CAC" pageNumber="870">
Subspecies
<taxonomicName id="8C2CA528B0DB5FC827AC27C0FD9A510E" authority="Hall and Villa 1948" authorityName="Hall and Villa" authorityYear="1948" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="pullus">
<emphasis id="19737D09DA9776D8BE4844D86869A071" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="4E3CFD94D28B496B5A6CCEB896804B10" italics="true" pageNumber="870">pullus</emphasis>
Hall and Villa 1948
</taxonomicName>
</paragraph>
<paragraph id="47485F127C4957AC020F1325D7689926" pageNumber="870">
Subspecies
<taxonomicName id="D0DDBDD538D328811CFBC311346F5216" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="sonoriensis">
<emphasis id="B93DE2F95BB303FEA08C5BE56740226E" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="64796E30833A5CE5DFE312EA40FAC4E1" italics="true" pageNumber="870">sonoriensis</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="2ADFD243B8BBFA6443ED7C40C01C8BFB" pageNumber="870">
Subspecies
<taxonomicName id="9EDCCDFF9B5F8C828CCBE5E0419D5167" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="supernus">
<emphasis id="D18A228520D5AA58C57857117FCDDC2E" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="C4142867AF54093AFE2B7E7E87536DFC" italics="true" pageNumber="870">supernus</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
<paragraph id="943073C5D29EED9816CA19D122046808" pageNumber="870">
Subspecies
<taxonomicName id="EA09E217E17EA384FACA8DB701084FB9" authority="Nelson and Goldman 1934" authorityName="Nelson and Goldman" authorityYear="1934" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" pageNumber="870" phylum="Chordata" rank="subSpecies" species="umbrinus" subGenus="Megascapheus" subSpecies="zacatecae">
<emphasis id="F45A20C57E58E6A12C903EA5AFB58D70" italics="true" pageNumber="870">Thomomys (Megascapheus) umbrinus</emphasis>
subsp.
<emphasis id="0854503F9D69BC378608B55A3869F8AE" italics="true" pageNumber="870">zacatecae</emphasis>
Nelson and Goldman 1934
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8172464E6454B14053CA80AE4697CCD1" pageNumber="870" type="distribution">
<paragraph id="A7C9A32D0812FC2D4835841C062328DD" pageNumber="870">
<emphasis id="EF4F66661232922E9A93B784C9D1BF07" inLineHeading="true" pageNumber="870">Distribution:</emphasis>
SC
<collectingRegion id="B64F704DFC9AD49A9F784CAFB95034E9" country="United States of America" name="Arizona">Arizona</collectingRegion>
and SW
<collectingRegion id="2ABE95CCB1DBE6964E93777559AAB939" country="United States of America" name="New Mexico">New Mexico</collectingRegion>
(
<collectingCountry id="815E53C1B2269E21DB74DCDFD5BF64DF" name="United States of America">USA</collectingCountry>
) south to
<collectingRegion id="D468AA44C636701FF9E1C240B890CFD1" country="Mexico" name="Puebla">Puebla</collectingRegion>
and
<collectingRegion id="3DA9DE7C5EB81D6DCFD50DE62E685B93" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
(
<collectingCountry id="C9CA4966A0F022FAAEB8A2B9E0865279" name="Mexico">Mexico</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="B0E0E0946FF3DF8F35820CEF8F249D85" pageNumber="870" type="conservation">
<paragraph id="29078C7D400E92CD3567CEEB4CC0A4A0" pageNumber="870">
<emphasis id="764699D8CA56A3C9FBF73E9DC683B120" inLineHeading="true" pageNumber="870">Conservation:</emphasis>
<collectionCode id="489AE493BE35189986CEE1E31335CC75">IUCN</collectionCode>
Lowered Risk (nt) as
<emphasis id="F90017918E94B5EB09F34FB808EC8B76" italics="true" pageNumber="870">
<collectionCode id="304B0F42A19EC3E3CAC967442DD442DF" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. u. emotus
</emphasis>
, otherwise Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="E74EF643E23E511204961FF99B7A9D2F" pageNumber="870" type="discussion">
<paragraph id="1F4F8F4126EB32A5C8652EE9F01C4799" pageNumber="870">
<emphasis id="42CEF7763DAA25332F4B69EE93E577AB" inLineHeading="true" pageNumber="870">Discussion:</emphasis>
Subgenus
<taxonomicName id="0CF1F8C0ACD01386721BA49F68F1294B" authorityName="Elliot" authorityYear="1903" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="subGenus" subGenus="Megascapheus">
<emphasis id="EF4B28D87D9362563C42D7F82B8ABC8C" italics="true" pageNumber="870">Megascapheus</emphasis>
</taxonomicName>
. Hybridization with
<taxonomicName id="10AA166F95CF19CBF8BF4DB2C1B34D92" baseAuthorityName="Eydoux and Gervais" baseAuthorityYear="1836" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bottae">
<emphasis id="75B8F8501E437FE5B2E70F50546FB752" italics="true" pageNumber="870">bottae</emphasis>
</taxonomicName>
reported in Patagonia Mtns of S Arizona by
<bibRefCitation id="8CBDE7ADE86464BFF2EC31A502878D10" author="Patton, J. L. &amp; R. E. Dingman" refId="ref275505" refString="Patton, J. L., and R. E. Dingman. 1968. Chromosome studies of pocket gophers, genus Thomomys. I. The specific status of Thomomys umbrinus (Richardson) in Arizona. Journal of Mammalogy, 49: 1 - 13." year="1968">Patton and Dingman (1968)</bibRefCitation>
,
<bibRefCitation id="4FAB629F84FD7B101AB6E7DBF05D4FF1" author="Patton, J. L." refId="ref274980" refString="Patton, J. L. 1973. An analysis of natural hybridization between the pocket gophers Thomomys bottae and Thomomys umbrinus, in Arizona. Journal of Mammalogy, 54: 561 - 584." year="1973">Patton (1973)</bibRefCitation>
, and
<bibRefCitation id="FF76F1D99B7F6F1F85F4367D011F89C6" author="Hoffmeister, D. F." refId="ref155598" refString="Hoffmeister, D. F. 1969. The species problem in the Thomomys bottae-Thomomys umbrinus complex of gophers in Arizona. Miscellaneous Publications, Museum of Natural History, University of Kansas, 51: 75 - 91." year="1969">Hoffmeister (1969</bibRefCitation>
,
<bibRefCitation id="4166DC31DF3E12C4220D729ABF8831ED" author="Hoffmeister, D. F." refId="ref155684" refString="Hoffmeister, D. F. 1986. Mammals of Arizona. University of Arizona Press, Tucson., 602 pp." year="1986">1986</bibRefCitation>
). Considered a species separate from
<taxonomicName id="CD269BE423F5B09825BE9DABC73F82FE" baseAuthorityName="Eydoux and Gervais" baseAuthorityYear="1836" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bottae">
<emphasis id="CD78E148B91A4E9CBDC1A75A194233D9" italics="true" pageNumber="870">bottae</emphasis>
</taxonomicName>
by these authors and by
<bibRefCitation id="503EA60C3129B1F7472476AC6E268B8B" author="Anderson, S." refId="ref9015" refString="Anderson, S. 1966. Taxonomy of gophers, especially Thomomys in Chihuahua, Mexico. Systematic Zoology, 15: 189 - 198." year="1966">Anderson (1966</bibRefCitation>
,
<bibRefCitation id="07CA6ECD7A52AEB7BBF5D49C0744AC2E" author="Anderson, S." refId="ref9154" refString="Anderson, S. 1972. Mammals of Chihuahua: Taxonomy and distribution. Bulletin of the American Museum of Natural History, 148: 149 - 410." year="1972">1972</bibRefCitation>
).
<bibRefCitation id="0D0C8CEDB8C35DDCA1A0371956C9BC4B" author="Hall, E. R." refId="ref137310" refString="Hall, E. R. 1981. The mammals of North America. Second ed. John Wiley and Sons, New York, 1: 1 - 600 + 90, 2: 601 - 1181 + 90." year="1981">Hall (1981:469)</bibRefCitation>
included
<emphasis id="9E82EB413E64DDC12A1A874FC7639BC2" italics="true" pageNumber="870">baileyi</emphasis>
,
<taxonomicName id="BF12AE3824482494AA2EBC9EE9010D4C" baseAuthorityName="Eydoux and Gervais" baseAuthorityYear="1836" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bottae">
<emphasis id="26A4511F7DC20C77ECA84A3598A3BDFA" italics="true" pageNumber="870">bottae</emphasis>
</taxonomicName>
, and
<taxonomicName id="423C7274A644580D9E7440CCFA8FFE92" baseAuthorityName="Bachman" baseAuthorityYear="1839" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="townsendii">
<emphasis id="6E4A1D172B8EAEC11672EFAED3171668" italics="true" pageNumber="870">townsendii</emphasis>
</taxonomicName>
in this species based on his view of the hybridization data in
<bibRefCitation id="7CA687E43F8E911E78019A0A1B1AA781" author="Hoffmeister, D. F." refId="ref155598" refString="Hoffmeister, D. F. 1969. The species problem in the Thomomys bottae-Thomomys umbrinus complex of gophers in Arizona. Miscellaneous Publications, Museum of Natural History, University of Kansas, 51: 75 - 91." year="1969">Hoffmeister (1969)</bibRefCitation>
,
<bibRefCitation id="69498A4DA535C7C10F9A7130F84113C1" author="Patton, J. L." refId="ref274980" refString="Patton, J. L. 1973. An analysis of natural hybridization between the pocket gophers Thomomys bottae and Thomomys umbrinus, in Arizona. Journal of Mammalogy, 54: 561 - 584." year="1973">Patton (1973)</bibRefCitation>
, and
<bibRefCitation id="1C3E89EF5DC6F4A62A36EB356ECB7FA3" author="Thaeler, C. S., Jr." refId="ref350523" refString="Thaeler, C. S., Jr. 1968 b. An analysis of three hybrid populations of pocket gophers (genus Thomomys). Evolution, 22: 543 - 555." year="1968">
Thaeler (1968
<emphasis id="B9529F671EFE516E1DAAB84FB700DA8F" italics="true" pageNumber="870">b</emphasis>
)
</bibRefCitation>
; he reported intergradation between
<taxonomicName id="F2E5C86CE8B3F310D02F53E20EE007BB" baseAuthorityName="Richardson" baseAuthorityYear="1829" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="umbrinus">
<emphasis id="60C1813E5F3E155B625F5BA6D10B1149" italics="true" pageNumber="870">umbrinus</emphasis>
</taxonomicName>
and
<taxonomicName id="C0F26915CEB7C99C787FF375F9286D89" baseAuthorityName="Eydoux and Gervais" baseAuthorityYear="1836" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="bottae">
<emphasis id="4A69388577DBEFDB2352BD240D6BE4F1" italics="true" pageNumber="870">bottae</emphasis>
</taxonomicName>
in
<collectingRegion id="4747A17E8011977F7C3605836518FEAB" country="Mexico" name="Nuevo Leon">Nuevo Leon</collectingRegion>
(
<collectingCountry id="B594F71396A8E80D7716D5628F66B62D" name="Mexico">Mexico</collectingCountry>
), which has not been verified by genetic analyses. Hafner et al. (1987) examined phylogenetic relationships among geographic units of
<taxonomicName id="B8E702EBAA21CFCCA3825DE207C78B2F" baseAuthorityName="Richardson" baseAuthorityYear="1829" class="Mammalia" family="Geomyidae" genus="Thomomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="umbrinus">
<emphasis id="03D9CC717721DB2AAB3DE47C36993096" italics="true" pageNumber="870">umbrinus</emphasis>
</taxonomicName>
. Revised, in part, by
<bibRefCitation id="7A65AE9D885625B2BFBB3EBBF6395997" author="Nelson, E. W. &amp; E. A. Goldman" refId="ref257669" refString="Nelson, E. W., and E. A. Goldman. 1934 b. Pocket gophers of the genus Thomomys of the Mexican mainland and bordering territory. Journal of Mammalogy, 15: 105 - 124." year="1934">
Nelson and Goldman (1934
<emphasis id="1E1F2E02EB4C27385B69454160A00213" italics="true" pageNumber="870">b</emphasis>
)
</bibRefCitation>
,
<bibRefCitation id="2824F40209BD1DD558DD5633D0E9BAB4" author="Anderson, S." refId="ref9154" refString="Anderson, S. 1972. Mammals of Chihuahua: Taxonomy and distribution. Bulletin of the American Museum of Natural History, 148: 149 - 410." year="1972">Anderson (1972</bibRefCitation>
; for
<collectingRegion id="08E97DF5028081E2C3964E31FEEC06FF" country="Mexico" name="Chihuahua">Chihuahua</collectingRegion>
), and Castro-Campillo and Ramírez-Pulido (2000; for C
<collectingCountry id="F801488165A64A1F42223F02A12C7344" name="Mexico">Mexico</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>