267 lines
35 KiB
XML
267 lines
35 KiB
XML
<document id="EFE12A5BE1FA4E586C4926E0F464CEFB" ID-DOI="10.1080/713833844" ID-ISSN="1464-5262" ID-Zenodo-Dep="5298420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629991564990" checkinUser="felipe" docAuthor="Cumberlidge, Neil & Wranik, Wolfgang" docDate="2002" docId="046287FFA052FFEEFE4EFA1B61AAD259" docLanguage="en" docName="JNatHist.36.1.51-64.pdf" docOrigin="Journal of Natural History 36 (1)" docSource="http://www.tandfonline.com/doi/abs/10.1080/713833844" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Socotra pseudocardisoma Cumberlidge & Wranik 2002, n. sp." docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="F85BFF87A056FFE5FFC4FFF66253D138" masterDocTitle="A new genus and new species of freshwater crab (Potamoidea, Potamidae) from Socotra Island, Yemen" masterLastPageNumber="64" masterPageNumber="51" pageNumber="53" updateTime="1699133984655" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="D8D2130C9335B8E2BDAB18049FBD3D5F">
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<mods:title id="D20BAFC261A4F3BC86A2DE57DC224F29">A new genus and new species of freshwater crab (Potamoidea, Potamidae) from Socotra Island, Yemen</mods:title>
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</mods:titleInfo>
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<mods:name id="7A07574419B4BF89B28CAD136260D441" type="personal">
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<mods:roleTerm id="4C93E5D62E335C75B04A01F284473B52">Author</mods:roleTerm>
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<mods:namePart id="AFB2CC5A9518B13F59A1C8893AEC8055">Cumberlidge, Neil</mods:namePart>
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<mods:name id="F496798940240E3B9BBFA44FC1737BB1" type="personal">
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<mods:roleTerm id="FE07B2851F0E7DC52EEC1DBFF4A5C87A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E83501FD24B55E6694FF2261A7D3E0DE">Wranik, Wolfgang</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B2C1A2E1024BC61247137F3F31BAC152">text</mods:typeOfResource>
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<mods:title id="1505B2B39BCFA538526ADFA374D8645F">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="97756523A4BEA9DB33697A268975C1F9">
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<mods:date id="5EA9979D6F255D8D6C48B9A90155E05C">2002</mods:date>
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<mods:detail id="DB9BE5F0CBB647C9EBB1298DB4539F0E" type="pubDate">
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<mods:number id="16FCA3594F46A3FE5E67A2795053282F">2002-01-31</mods:number>
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</mods:detail>
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<mods:detail id="1FDE4B06BA3CCF2228829DC29B39BDFC" type="volume">
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<mods:number id="055E9A489B3C5DF239868F4EA0E372E1">36</mods:number>
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</mods:detail>
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<mods:detail id="DDF88088B5E9FCB560DBF001C99FE813" type="issue">
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<mods:number id="2EA3CF2737E1C1904B41051C2AE3FAD4">1</mods:number>
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<mods:start id="E6D00667078D1A4771055D79677170C9">51</mods:start>
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<mods:location id="0180242F01AB12AC40EE5C70A2748473">
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<mods:url id="2E009C6E2AD44E6ABA3FC0BCCD56B21E">http://www.tandfonline.com/doi/abs/10.1080/713833844</mods:url>
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<mods:classification id="360C95FB87155C8B6D9B1831423AEB83">journal article</mods:classification>
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<mods:identifier id="07A4AC6A89C6461AD9B464250580EDEE" type="DOI">10.1080/713833844</mods:identifier>
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<mods:identifier id="0C5AA8E2D049D04609FD73AA722E9043" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="C632B69243B417519B2947EBFEA682AE" type="Zenodo-Dep">5298420</mods:identifier>
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<treatment id="046287FFA052FFEEFE4EFA1B61AAD259" LSID="urn:lsid:plazi:treatment:046287FFA052FFEEFE4EFA1B61AAD259" httpUri="http://treatment.plazi.org/id/046287FFA052FFEEFE4EFA1B61AAD259" lastPageId="11" lastPageNumber="60" pageId="4" pageNumber="53">
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<subSubSection id="C4D16562A052FFE1FE4EFA1B60D0D71D" pageId="4" pageNumber="53" type="nomenclature">
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<paragraph id="8C7436E9A052FFE1FE4EFA1B60BFD73C" blockId="4.[394,748,1517,1573]" box="[394,748,1517,1541]" pageId="4" pageNumber="53">
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<heading id="D73C8185A052FFE1FE4EFA1B60BFD73C" box="[394,748,1517,1541]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="53" reason="3">
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<taxonomicName id="4BCB4D6AA052FFE1FE4EFA1B60F6D73D" ID-CoL="4XYRZ" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[394,677,1517,1541]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="53" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
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<emphasis id="BEBFEAFBA052FFE1FE4EFA1B60F6D73D" bold="true" box="[394,677,1517,1541]" italics="true" pageId="4" pageNumber="53">Socotra pseudocardisoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A58C5780A052FFE1FD69FA1B60BFD73C" box="[685,748,1517,1540]" pageId="4" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8C7436E9A052FFE1FE30F9FB60D0D71D" blockId="4.[394,748,1517,1573]" box="[500,643,1549,1573]" pageId="4" pageNumber="53">(®gures 1±6)</paragraph>
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</subSubSection>
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<subSubSection id="C4D16562A052FFEEFF42F9B061AAD259" lastPageId="11" lastPageNumber="60" pageId="4" pageNumber="53" type="materials_examined">
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<paragraph id="8C7436E9A052FFE1FF42F9B0664AD766" blockId="4.[93,1049,1606,1662]" box="[134,1049,1606,1631]" pageId="4" pageNumber="53">
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<emphasis id="BEBFEAFBA052FFE1FF42F9B0630BD766" box="[134,344,1606,1630]" italics="true" pageId="4" pageNumber="53">Material examined</emphasis>
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.
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<materialsCitation id="3CA33CB4A052FFE1FEACF9B0664AD766" box="[360,1049,1606,1631]" collectionCode="NHML" pageId="4" pageNumber="53" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5370884BA052FFE1FEACF9B06381D766" box="[360,466,1606,1630]" pageId="4" pageNumber="53" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="EADAAE2CA052FFE1FE25F9B1606AD766" box="[481,569,1607,1630]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="53">NHML</collectionCode>
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reg. 1999: adult male (cw 77.2, cl 52.4, ch
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</materialsCitation>
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</paragraph>
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<paragraph id="8C7436E9A052FFE0FF99F990604BD467" blockId="4.[93,1049,1606,1662]" lastBlockId="5.[93,1049,1256,1662]" lastPageId="5" lastPageNumber="54" pageId="4" pageNumber="53">
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28.3, fw
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<quantity id="4B339B0CA052FFE1FF04F9906373D746" box="[192,288,1638,1662]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="4" pageNumber="53" unit="mm" value="14.7">14.7 mm</quantity>
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).
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<typeStatus id="5370884BA052FFE1FEFCF99063C8D746" box="[312,411,1638,1662]" pageId="4" pageNumber="53" type="paratype">Paratype</typeStatus>
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: UR reg. 1999: adult male (cw 90.5, cl 59.0, ch 33.5, fw
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<quantity id="4B339B0CA053FFE0FF99FB1E62EDD438" box="[93,190,1256,1280]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="54" unit="mm" value="15.5">15.5 mm</quantity>
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), from Diksam (12ss29¾N, 53ss59¾E),
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<quantity id="4B339B0CA053FFE0FDBCFB1E60EFD5C7" box="[632,700,1256,1280]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="54" unit="m" value="700.0">700 m</quantity>
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, coll. W. Wranik,
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<date id="F8751029A053FFE0FC66FB1E62C2D427" pageId="5" pageNumber="54" value="1998-10-04">4 October 1998</date>
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.
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<typeStatus id="5370884BA053FFE0FF65FAFE6357D418" box="[161,260,1288,1312]" pageId="5" pageNumber="54" type="paratype">Paratype</typeStatus>
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: NMU reg. 1999: adult female (cw 77.2, cl 52.4, ch 28.3, fw
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<quantity id="4B339B0CA053FFE0FC75FAF1665CD427" box="[945,1039,1287,1311]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="5" pageNumber="54" unit="mm" value="14.7">14.7 mm</quantity>
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) from Diksam,
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<quantity id="4B339B0CA053FFE0FECEFAD1631DD407" box="[266,334,1319,1343]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="54" unit="m" value="700.0">700 m</quantity>
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, coll. W. Wranik,
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<date id="F8751029A053FFE0FDEEFAD160ABD407" box="[554,760,1319,1343]" pageId="5" pageNumber="54" value="1997-11-05">5 November 1997</date>
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. Eleven other specimens of
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<taxonomicName id="4BCB4D6AA053FFE0FFBAFAB16309D467" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[126,346,1351,1375]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="pseudocardisoma">
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<emphasis id="BEBFEAFBA053FFE0FFBAFAB16309D467" box="[126,346,1351,1375]" italics="true" pageId="5" pageNumber="54">S. pseudocardisoma</emphasis>
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</taxonomicName>
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are held at UR.
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</paragraph>
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<caption id="D8B46661A053FFE0FF99FB906331D588" ID-DOI="http://doi.org/10.5281/zenodo.5298422" ID-Zenodo-Dep="5298422" httpUri="https://zenodo.org/record/5298422/files/figure.png" pageId="5" pageNumber="54" startId="5.[93,109,1126,1147]" targetBox="[167,978,109,1097]" targetPageId="5">
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<paragraph id="8C7436E9A053FFE0FF99FB906331D588" blockId="5.[93,1049,1126,1201]" pageId="5" pageNumber="54">
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FIG. 1.
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<taxonomicName id="4BCB4D6AA053FFE0FF7DFB9063ECD544" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[185,447,1126,1148]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="54" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
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||
<emphasis id="BEBFEAFBA053FFE0FF7DFB9063ECD544" box="[185,447,1126,1148]" italics="true" pageId="5" pageNumber="54">Socotra pseudocardisoma</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A58C5780A053FFE0FE0AFB90601ED544" box="[462,589,1126,1148]" pageId="5" pageNumber="54" rank="species">new species</taxonomicNameLabel>
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, from
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<taxonomicName id="4BCB4D6AA053FFE0FD60FB9060A0D544" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[676,755,1126,1148]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
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, male, holotype, carapace width 577.2 mm. (A) Cephalothorax, dorsal aspect; (B) cephalothorax, frontal aspect. Scale bar: 5.0 mm.
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</paragraph>
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</caption>
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<paragraph id="8C7436E9A053FFE2FF42FA9E602FD706" blockId="5.[93,1049,1256,1662]" lastBlockId="7.[93,1049,1478,1662]" lastPageId="7" lastPageNumber="56" pageId="5" pageNumber="54">
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<emphasis id="BEBFEAFBA053FFE0FF42FA9E62ABD447" box="[134,248,1384,1407]" italics="true" pageId="5" pageNumber="54">Diagnosis.</emphasis>
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Cephalothorax heart-shaped (cw>cl): highly arched (ch/fw 1.9), very wide (cw/fw 5.3), very long (cl/fw 3.6); front very narrow (fw/cw 0.2). Postfrontal crest low, faint, complete, ends curving forward to meet anterolateral margins at epibranchial teeth; mid-groove between epigastric crests long, ¯at, wide, forked at posterior end. Exorbital tooth small, low, blunt; epibranchial tooth reduced, almost undetectable, forming part of rim of anterolateral margin; anterolateral margin between exorbital, epibranchial teeth extremely short, lacking intermediate tooth; anterolateral margin posterior to epibranchial tooth forming a smooth raised rim. Front sharply deēxed, meeting inferior margins of antennulular fossae so that antennules almost completely enclosed within fossae. Mandibular palp threesegmented, terminal segment single, undivided. Exopod of third maxilliped distinctly widened at midpoint, base, tip narrow (i.e. not long, columnar with parallel sides). Third sternal sulcus (s3/s4) reduced to two short notches at lateral edges of sternum; anterior margin of sterno-abdominal cavity almost reaching s3/s4 junction. Episternal sulci s4/e4 to s7/e7 not visible. Gonopod 1 long, extending anteriorly as far as s4/s5. Terminal article of gonopod 1 relatively long (about one-third as long as subterminal segment), terminal article of gonopod 2 long, ¯agellum-like (almost as long as subterminal segment of gonopod 2).
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</paragraph>
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<caption id="D8B46661A050FFE3FF99FAF160B2D498" ID-DOI="http://doi.org/10.5281/zenodo.5298424" ID-Zenodo-Dep="5298424" httpUri="https://zenodo.org/record/5298424/files/figure.png" pageId="6" pageNumber="55" startId="6.[93,109,1287,1308]" targetBox="[249,898,109,1258]" targetPageId="6">
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<paragraph id="8C7436E9A050FFE3FF99FAF160B2D498" blockId="6.[93,1049,1286,1441]" pageId="6" pageNumber="55">
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FIG. 2.
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<taxonomicName id="4BCB4D6AA050FFE3FF7DFAF063ECD424" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[185,447,1286,1308]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="55" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
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<emphasis id="BEBFEAFBA050FFE3FF7DFAF063ECD424" box="[185,447,1286,1308]" italics="true" pageId="6" pageNumber="55">Socotra pseudocardisoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A58C5780A050FFE3FE0AFAF1601ED424" box="[462,589,1286,1308]" pageId="6" pageNumber="55" rank="species">new species</taxonomicNameLabel>
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, from
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<taxonomicName id="4BCB4D6AA050FFE3FD60FAF060A0D424" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[676,755,1286,1308]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
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, male, holotype, carapace width 577.2 mm. (A) Carpus of right cheliped, superior view; (B) carpus and merus of right cheliped, superior view; (C) merus of right cheliped inferior view; (D) left mandible showing details of palp; (E) abdomen and part of sternum; (F) left third maxilliped showing details of the exopod, and anterior sternum. Scale bar: 6.0 mm (A±C), 1.2 mm (D), 10.0 mm (E), 2.6 mm (F).
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</paragraph>
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||
</caption>
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||
<caption id="D8B46661A051FFE2FF99FDAF63BED39C" ID-DOI="http://doi.org/10.5281/zenodo.5298426" ID-Zenodo-Dep="5298426" httpUri="https://zenodo.org/record/5298426/files/figure.png" pageId="7" pageNumber="56" startId="7.[93,109,601,622]" targetBox="[353,792,107,572]" targetPageId="7">
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||
<paragraph id="8C7436E9A051FFE2FF99FDAF63BED39C" blockId="7.[93,1049,601,676]" pageId="7" pageNumber="56">
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FIG. 3.
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<taxonomicName id="4BCB4D6AA051FFE2FF7DFDAF63EAD357" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[185,441,601,623]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
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<emphasis id="BEBFEAFBA051FFE2FF7DFDAF63EAD357" box="[185,441,601,623]" italics="true" pageId="7" pageNumber="56">Socotra pseudocardisoma</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A58C5780A051FFE2FE07FDAF606FD357" box="[451,572,601,623]" pageId="7" pageNumber="56" rank="species">new species</taxonomicNameLabel>
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, from
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<taxonomicName id="4BCB4D6AA051FFE2FD43FDAF6085D357" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[647,726,601,623]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
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, male, type, carapace width 5 77.2 mm. (A) Details of left orbital region, frontal view; (B) details of frontal region, frontal view. Scale bar: 2.5 mm.
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</paragraph>
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</caption>
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||
<caption id="D8B46661A051FFE2FF99FAC2664BD4A1" ID-DOI="http://doi.org/10.5281/zenodo.5298428" ID-Zenodo-Dep="5298428" httpUri="https://zenodo.org/record/5298428/files/figure.png" pageId="7" pageNumber="56" startId="7.[93,109,1332,1353]" targetBox="[212,932,720,1303]" targetPageId="7">
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||
<paragraph id="8C7436E9A051FFE2FF99FAC2664BD4A1" blockId="7.[93,1049,1332,1433]" pageId="7" pageNumber="56">
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||
FIG. 4.
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<taxonomicName id="4BCB4D6AA051FFE2FF7DFAC263EAD472" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[185,441,1332,1354]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
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<emphasis id="BEBFEAFBA051FFE2FF7DFAC263EAD472" box="[185,441,1332,1354]" italics="true" pageId="7" pageNumber="56">Socotra pseudocardisoma</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A58C5780A051FFE2FE07FAC2606FD472" box="[451,572,1332,1354]" pageId="7" pageNumber="56" rank="species">new species</taxonomicNameLabel>
|
||
, from
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<taxonomicName id="4BCB4D6AA051FFE2FD43FAC26085D472" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[647,726,1332,1354]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
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, male, type, carapace width 5 77.2 mm. (A) Right gonopod 1, ventral aspect; (B) right gonopod 1, dorsal aspect; (C) right gonopod 1, dorsal aspect; (D) right gonopod 1, superior view, showing groove; (E) right gonopod 2, ventral aspect. Scale bar: 2.5 mm (A±C, E), 1.6 mm (D).
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||
</paragraph>
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</caption>
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<paragraph id="8C7436E9A051FFE2FF42F9B1664AD746" blockId="7.[93,1049,1478,1662]" pageId="7" pageNumber="56">
|
||
<emphasis id="BEBFEAFBA051FFE2FF42F9B16357D766" box="[134,260,1607,1630]" italics="true" pageId="7" pageNumber="56">Description</emphasis>
|
||
. Following description based on male
|
||
<typeStatus id="5370884BA051FFE2FD09F9B06162D766" box="[717,817,1606,1630]" pageId="7" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="EADAAE2CA051FFE2FC82F9B161CDD766" box="[838,926,1607,1630]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="7" pageNumber="56">NHML</collectionCode>
|
||
reg. 1999: adult male, cw
|
||
<quantity id="4B339B0CA051FFE2FEC2F9906335D746" box="[262,358,1638,1662]" metricMagnitude="-2" metricUnit="m" metricValue="7.72" pageId="7" pageNumber="56" unit="mm" value="77.2">77.2 mm</quantity>
|
||
). Cephalothorax heart-shape d (cw>cl): highly arched (ch/fw
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05EFFEDFDF4FC736019D2A0" blockId="8.[560,586,901,920]" box="[560,586,901,920]" pageId="8" pageNumber="57">
|
||
(
|
||
<emphasis id="BEBFEAFBA05EFFEDFDFCFC706010D2A0" box="[568,579,902,920]" italics="true" pageId="8" pageNumber="57">a</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05EFFEDFDF4FAAC6019D455" blockId="8.[560,586,1369,1389]" box="[560,586,1369,1389]" pageId="8" pageNumber="57">
|
||
(
|
||
<emphasis id="BEBFEAFBA05EFFEDFDF3FAAF6011D454" box="[567,578,1369,1388]" italics="true" pageId="8" pageNumber="57">b</emphasis>
|
||
)
|
||
</paragraph>
|
||
<caption id="D8B46661A05EFFEDFF99FA7F63ACD4EC" ID-DOI="http://doi.org/10.5281/zenodo.5298430" ID-Zenodo-Dep="5298430" httpUri="https://zenodo.org/record/5298430/files/figure.png" pageId="8" pageNumber="57" startId="8.[93,109,1417,1438]" targetBox="[94,1051,949,1360]" targetPageId="8">
|
||
<paragraph id="8C7436E9A05EFFEDFF99FA7F63ACD4EC" blockId="8.[93,1049,1417,1492]" pageId="8" pageNumber="57">
|
||
FIG. 5. (a) Sketch map showing the
|
||
<taxonomicName id="4BCB4D6AA05EFFEDFE27FA7F60E4D4A7" authority="Archipelago" authorityName="Archipelago" box="[483,695,1417,1439]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="genus">Socotra Archipelago</taxonomicName>
|
||
(source: Kopp, in
|
||
<bibRefCitation id="E85A4B18A05EFFEDFCB9FA7F665BD4A7" author="WRANIK, W." box="[893,1032,1417,1439]" pageId="8" pageNumber="57" refId="ref7228" refString="WRANIK, W., 1999, Sokotra, Mensch und Natur, Jemen Studien, 258." type="book" year="1999">Wranik, 1999</bibRefCitation>
|
||
); (b) map of
|
||
<taxonomicName id="4BCB4D6AA05EFFEDFED8FA556338D481" box="[284,363,1443,1465]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
|
||
showing highland area where
|
||
<taxonomicName id="4BCB4D6AA05EFFEDFD65FA5561CED481" authority="Cumberlidge & Wranik, 2002" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[673,925,1443,1465]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="pseudocardisoma" status="sp. nov.">
|
||
<emphasis id="BEBFEAFBA05EFFEDFD65FA5561CED481" box="[673,925,1443,1465]" italics="true" pageId="8" pageNumber="57">Socotra pseudocardisoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A58C5780A05EFFEDFC60FA52664AD481" box="[932,1049,1443,1465]" pageId="8" pageNumber="57" rank="species">new species</taxonomicNameLabel>
|
||
was collected (12ss27¾N, 53ss59¾E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C7436E9A05EFFECFF99F9F06065D578" blockId="8.[93,1049,1542,1662]" lastBlockId="9.[93,1049,873,1662]" lastPageId="9" lastPageNumber="58" pageId="8" pageNumber="57">1.9), very wide (cw/fw 5.3), very long (cl/fw 3.6); front very narrow (fw/cw 0.2). Dorsal surface of carapace texture smooth; semicircular, urogastric grooves deep, cervical grooves deep, short; transverse cardiac grooves faint; branchial grooves not visible. Postfrontal crest low, faint, complete, ends curving forward to meet anterolateral margins at epibranchial teeth; mid-groove between epigastric crests long, ¯at, wide, forked at posterior end; mid-groove extending anteriorly for short distance on to frontal region. Exorbital tooth small, low, blunt; epibranchial tooth reduced, almost undetectable, forming part of rim of anterolateral margin; anterolateral margin between exorbital, epibranchial teeth extremely short, lacking intermediate tooth; anterolateral margin posterior to epibranchial tooth forming a smooth raised rim, posterior end curving inward.</paragraph>
|
||
<caption id="D8B46661A05FFFECFF99FCF160B2D20E" ID-DOI="http://doi.org/10.5281/zenodo.5298432" ID-Zenodo-Dep="5298432" httpUri="https://zenodo.org/record/5298432/files/figure.png" pageId="9" pageNumber="58" startId="9.[93,109,775,796]" targetBox="[93,1052,106,748]" targetPageId="9">
|
||
<paragraph id="8C7436E9A05FFFECFF99FCF160B2D20E" blockId="9.[93,1049,774,822]" pageId="9" pageNumber="58">
|
||
FIG. 6. Male specimen of
|
||
<taxonomicName id="4BCB4D6AA05FFFECFE47FCF060D5D224" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[387,646,774,796]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="pseudocardisoma">
|
||
<emphasis id="BEBFEAFBA05FFFECFE47FCF060D5D224" box="[387,646,774,796]" italics="true" pageId="9" pageNumber="58">Socotra pseudocardisoma</emphasis>
|
||
</taxonomicName>
|
||
climbing on a rock in the limestone plateau type locality of
|
||
<taxonomicName id="4BCB4D6AA05FFFECFD48FCD66088D20E" box="[652,731,800,822]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C7436E9A05FFFECFF42FBBE60E0D726" blockId="9.[93,1049,873,1662]" pageId="9" pageNumber="58">Front sharply deēxed, meeting inferior margins of antennulular fossae so that antennules almost completely enclosed within fossae. Lateral parts of epistome forming upper roof of eOEerent channels overlapping large, triangular, horizontal epistomial tooth. Margins of lateral epistome, epistomial tooth smooth, distinct endostomial ridge visible on lateral epistome marking medial boundary of eOEerent channel formed where broadened endopod of ®rst maxilliped meets endostome. Mandibular palp three-segmented terminal segment single, undivided. Third maxillipeds ®lling oral ®eld, medial margins of ischia, meri vertical, meeting along entire length; ischium with deep vertical sulcus; exopod of third maxilliped distinctly widened at midpoint, base, tip narrow; exopod ¯agellum reduced, measuring only half length of exopod. First sternal sulcus (s1/s2) short, distinct; second sulcus (s2/s3) deep, running horizontally across sternum; third sternal sulcus (s3/s4) reduced to two short notches at lateral edges of sternum, in line with basal segment of third maxillipeds; anterior margin of sterno-abdominal cavity almost reaching s3/s4 junction; episternal sulci (s4/e4 to s7/e7) not visible.</paragraph>
|
||
<paragraph id="8C7436E9A05FFFEFFF42F9D061E2D0BA" blockId="9.[93,1049,873,1662]" lastBlockId="10.[93,1050,107,1662]" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58">Dactylus of right cheliped narrow (one-quarter height of palm), upper margin smooth; dactylus slightly arched, enclosing long narrow interspace; ®nger of propodus slim (one-quarter height of palm). Lower margin of propodus indented. Both ®ngers with series of small pointed teeth, interspersed with several bigger teeth. Propodus of cheliped with conspicuous rounded boss at articulation point with carpus of pereiopod 1; carpus with raised ¯at process at this junction. Medial, lateral inferior margins of merus lined by small distinct teeth; single large pointed distal meral tooth on medial margin; superior margin of merus slightly granular. First carpal tooth of cheliped large, slender, pointed, second carpal tooth small pointed granule, on raised ridge with similar granules. Walking legs (pereiopods 2±5) long, slender; P3 longest, P5 shortest; dactyli of P2±P5 with four distinct rows of short corneous spines; anterior, posterior margins of propodi of P2±P5 smooth.</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FE7C639DD3F9" blockId="10.[93,1050,107,1662]" pageId="10" pageNumber="59">Male abdomen (a3±a6, plus telson) triangular; telson triangular with almost straight sides directed inward. Gonopod 1 long, reaching sternal groove s4/s5; terminal article relatively long (about one-third as long as subterminal segment); terminal article initially straight, curving sharply outward at halfway point, then curving sharply upward, tapering to broad tip; longitudinal groove visible only on dorsal side; wide dorsal membrane between last two parts; lateral fold of terminal article of gonopod 1 low, equal to medial fold. Gonopod 2 as long as gonopod 1; terminal article of gonopod 2 long, ¯agellum-like (almost as long as subterminal segment of gonopod 2) basal half straight, distal half either straight or folding back on itself, forming a hook-shape.</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FD3C60F7D3D9" blockId="10.[93,1050,107,1662]" box="[134,676,714,738]" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42FD3C62E6D3DA" box="[134,181,714,738]" italics="true" pageId="10" pageNumber="59">Size</emphasis>
|
||
. The adult size range is up to cw
|
||
<quantity id="4B339B0CA05CFFEFFDFAFD3C60CFD3D9" box="[574,668,714,738]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" pageId="10" pageNumber="59" unit="mm" value="90.5">90.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FD1C61C0D239" blockId="10.[93,1050,107,1662]" box="[134,915,745,769]" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42FD1C634CD239" box="[134,287,745,769]" italics="true" pageId="10" pageNumber="59">
|
||
<typeStatus id="5370884BA05CFFEFFF42FD1C62EDD239" box="[134,190,746,769]" pageId="10" pageNumber="59">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Diksam, altitude
|
||
<quantity id="4B339B0CA05CFFEFFE34FD1F6067D239" box="[496,564,745,769]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="59" unit="m" value="700.0">700 m</quantity>
|
||
, limestone plateau
|
||
<quantity id="4B339B0CA05CFFEFFCD7FD1F61DCD239" box="[787,911,745,769]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="7.4" metricValueMin="6.6" pageId="10" pageNumber="59" unit="m" value="700.0" valueMax="740.0" valueMin="660.0">660±740 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FCFF61BBD219" blockId="10.[93,1050,107,1662]" box="[134,1000,777,802]" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42FCFF6342D219" box="[134,273,777,801]" italics="true" pageId="10" pageNumber="59">Distribution.</emphasis>
|
||
The species is known only from the
|
||
<typeStatus id="5370884BA05CFFEFFD79FCFC60BDD21A" box="[701,750,778,802]" pageId="10" pageNumber="59">type</typeStatus>
|
||
locality (®gures 5, 6).
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FCDF6062D2D8" blockId="10.[93,1050,107,1662]" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42FCDF628AD279" box="[134,217,809,833]" italics="true" pageId="10" pageNumber="59">Colour.</emphasis>
|
||
In living specimens the dorsal surface of the carapace is dark purple with a broad light brown margin (most males) or brown with a broad yellowish margin (females). In some male specimens the dark surface coloration of the carapace continues over to the pterygostomium of the carapace sidewall. The walking legs (pereiopods P2±P5) are mainly yellowish, except for the ends of the merus, carpus and propodus, which have dark patches.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FC1E60D4D5E7" blockId="10.[93,1050,107,1662]" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42FC1E62B5D538" box="[134,230,1000,1024]" italics="true" pageId="10" pageNumber="59">Ecology.</emphasis>
|
||
The Haghir massif receives Socotra’s highest rainfall totals, and as a consequence the central highlands are frequently shrouded in clouds and heavy mists. Additional sources of water at these higher altitudes include permanent springs and the mist and dew. The Haghir massif forms the most important watershed on the island and this gives rise to numerous watercourses that run to the north and south of these highlands. The upper reaches of the streams running down the northern slopes tend to ¯ow year round, but at the lower elevations these streams become sporadic and ¯ow freely only after rain.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEFFF42FB1E639BD4C6" blockId="10.[93,1050,107,1662]" pageId="10" pageNumber="59">
|
||
The following ®eld observation s of the new taxon were made.
|
||
<emphasis id="BEBFEAFBA05CFFEFFCFCFB1E62CED427" italics="true" pageId="10" pageNumber="59">
|
||
<taxonomicName id="4BCB4D6AA05CFFEFFCFCFB1E665ED438" box="[824,1037,1256,1280]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="pseudocard">Socotra pseudocard</taxonomicName>
|
||
- isoma
|
||
</emphasis>
|
||
is a semi-arid zone crab with terrestrial habits whose behavioural pattern is inūenced by seasonal changes. The
|
||
<typeStatus id="5370884BA05CFFEFFDCBFADE6013D478" box="[527,576,1320,1344]" pageId="10" pageNumber="59">type</typeStatus>
|
||
locality is in a part of the island where limestone is dominant, and where climatic erosion of the calcareous substratum has produced sinks, underground streams, caverns, hollows and crevices. These temporary aquatic freshwater habitats are located far from conventional freshwater sources (rivers, streams and lakes) and are fed mainly by rainwater. It is possible that there are year-round, more permanent, water deposits deep below the surface, which could also be accessed by these crabs.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05CFFEEFF42F9F063A2D1DB" blockId="10.[93,1050,107,1662]" lastBlockId="11.[93,1049,107,866]" lastPageId="11" lastPageNumber="60" pageId="10" pageNumber="59">
|
||
<emphasis id="BEBFEAFBA05CFFEFFF42F9F063CDD726" box="[134,414,1542,1566]" italics="true" pageId="10" pageNumber="59">
|
||
<taxonomicName id="4BCB4D6AA05CFFEFFF42F9F063DCD726" box="[134,399,1542,1566]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="59" phylum="Arthropoda" rank="species" species="pseudocardisom">Socotra pseudocardisom</taxonomicName>
|
||
a
|
||
</emphasis>
|
||
lives in crevices, and crabs are not easy to ®nd, because they spend most of the daylight hours concealed from view. Crabs appear only occasionally on the surface, and startled crabs quickly retreat back into the crevice system. These animals can move rapidly on the ground, and can climb agilely and rapidly up and down rock surfaces. During the night, crabs leave their resting-places and wander on the surface in search of food. We are unsure of the natural diet of
|
||
<taxonomicName id="4BCB4D6AA05DFFEEFF99FF5D6369D1FB" authorityName="Cumberlidge & Wranik" authorityYear="2002" box="[93,314,171,195]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="pseudocardisoma">
|
||
<emphasis id="BEBFEAFBA05DFFEEFF99FF5D6369D1FB" box="[93,314,171,195]" italics="true" pageId="11" pageNumber="60">S. pseudocardisoma</emphasis>
|
||
</taxonomicName>
|
||
in its natural habitat, but specimens kept in captivity ate plant material and dead animal material.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05DFFEEFF42FF1D6357D0DA" blockId="11.[93,1049,107,866]" pageId="11" pageNumber="60">
|
||
There are distinct seasonal diOEerences in the behaviour patterns of
|
||
<emphasis id="BEBFEAFBA05DFFEEFCB4FF1D62C5D01B" italics="true" pageId="11" pageNumber="60">S. pseudocardi - soma</emphasis>
|
||
. Crabs were observed to be active only in October and November, and it is thought likely that they are active on the surface mainly during the wet season (November to March). It is possible that crabs mostly remain in their underground retreats during the drier times of the year (April to October). People that live on the limestone plateau near to the
|
||
<typeStatus id="5370884BA05DFFEEFE1EFE7D6058D09B" box="[474,523,395,419]" pageId="11" pageNumber="60">type</typeStatus>
|
||
locality, and who occasionally use these crabs as food, also report a similar seasonal pattern of wet season activity, and dry season disappearance.
|
||
</paragraph>
|
||
<paragraph id="8C7436E9A05DFFEEFF42FE1D61AAD259" blockId="11.[93,1049,107,866]" pageId="11" pageNumber="60">
|
||
We are able to report only basic information on the reproduction and life cycle of this species. It would appear that the production of eggs and release of hatchling crabs is timed to coincide with the wettest part of the year, when presumably plant and animal populations on
|
||
<taxonomicName id="4BCB4D6AA05DFFEEFE63FDBC63ACD35A" box="[423,511,586,610]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="63" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
|
||
are at their maxima. Five free-living juvenile crabs (cw
|
||
<quantity id="4B339B0CA05DFFEEFF12FD9C6319D3B9" box="[214,330,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="11" pageNumber="60" unit="mm" value="26.5" valueMax="27.0" valueMin="26.0">26±27 mm</quantity>
|
||
) already released from the maternal abdominal brood pouch were caught by one of the authors (W. W.) in summer 1998. Indeed, juvenile crabs are regularly sighted by local people during the late summer and early autumn when the monthly rainfall totals are at their highest. In captivity, mating was observed between a hard-shelled female and a hard-shelled male who remained in the copulatory position and/or a postmating embrace for about one day, before resuming normal behaviour. The sperm was presumably stored in the spermatheca of the female for later use in the fertilization of the eggs when they are eventually laid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |