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<mods:title id="62FEEA8C3DBB7AA2B1878FE568CA65F3">Taxonomic notes on Trigonidium and related groups with one new species from the Philippines (Orthoptera: Trigonidiidae; Trigonidiinae)</mods:title>
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<emphasis id="B93BEAF8FFBAFFD7BDFBEA43FC65FBEC" bold="true" box="[151,263,1041,1067]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
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<emphasis id="B93BEAF8FFBAFFD7BDFBEA0AFC76FBA9" box="[151,276,1112,1134]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
(
<emphasis id="B93BEAF8FFBAFFD7BC4FEA0AFCE3FBA9" box="[291,385,1112,1134]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
)
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:
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<bibRefCitation id="EFDE4B1BFFBAFFD7BCFBEA0AFF62FBA9" author="Stal, C." box="[407,512,1112,1134]" pageId="2" pageNumber="575" pagination="33 - 58" refId="ref7831" refString="Stal, C. (1877) Orthoptera nova ex Insulis Philippinis descripsit. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlinger, 34 (10), 33 - 58." type="journal article" year="1877">Stål, 1877</bibRefCitation>
: 48
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<bibRefCitation id="EFDE4B1BFFBAFFD7BF5DEA0AFFB5FBA9" author="Gorochov, A. V." box="[561,727,1112,1134]" pageId="2" pageNumber="575" pagination="5 - 17" refId="ref7133" refString="Gorochov, A. V. (1987) To the cricket fauna of the subfamilies Euscyrtinae, Trigonidiinae and Oecanthinae (Orthoptera, Gryllidae) from Eastern Indochina. In: Medvedev, L. N. (ed.) Entomofauna Vietnama [Entomofauna of Vietnam]: 5 - 17. Moscow: Nauka. (In Russian)." type="book chapter" year="1987">Gorochov, 1987</bibRefCitation>
: 10
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<bibRefCitation id="EFDE4B1BFFBAFFD7BE64EA0AFE14FBA9" author="Otte, D." box="[776,886,1112,1134]" pageId="2" pageNumber="575" refId="ref7761" refString="Otte, D. (1994) The Crickets of Hawaii: Origin, Systematics and Evolution. The Orthopterists' Society, The Academy of Natural Sciences of Philadelphia. 396 pp." type="book" year="1994">Otte, 1994</bibRefCitation>
: 46
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;
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<bibRefCitation id="EFDE4B1BFFBAFFD7BECBEA0AF92EFBA9" author="Sugimoto, M." box="[935,1100,1112,1134]" pageId="2" pageNumber="575" pagination="81 - 87" refId="ref7887" refString="Sugimoto, M. (2001) Taxonomic study of Trigonidiinae (Orthoptera: Trigonidiidae) of the Ryukyus, SW Japan, with descriptions of two new species. Tettigonia: Memoirs of the Orthopterological Society of Japan, 3, 81 - 87." type="journal article" year="2001">Sugimoto, 2001</bibRefCitation>
: 81
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87.
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<emphasis id="B93BEAF8FFBAFFD7BDFBEA24FD97FB4B" box="[151,245,1142,1164]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
:
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<bibRefCitation id="EFDE4B1BFFBAFFD7BC6EEA24FC1CFB4B" author="Kirby, W. F." box="[258,382,1142,1164]" pageId="2" pageNumber="575" refId="ref7635" refString="Kirby, W. F. (1906) A Synonymic Catalogue of Orthoptera. Volume II. British Museum (Nature History), London. 562 pp." type="book" year="1906">Kirby, 1906</bibRefCitation>
: 78
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;
<treatmentCitation id="0AEE10FBFFBAFFD7BCC3EA24FF1EFB4B" author="Chopard, L." box="[431,636,1142,1164]" page="467" pageId="2" pageNumber="575" year="1951">
<bibRefCitation id="EFDE4B1BFFBAFFD7BCC3EA24FF2BFB4B" author="Chopard, L." box="[431,585,1142,1164]" pageId="2" pageNumber="575" pagination="397 - 533" refId="ref6839" refString="Chopard, L. (1951) A revision of the Australian Grylloidea. Records of the South Australian Museum, 9, 397 - 533." type="journal article" year="1951">Chopard, 1951</bibRefCitation>
: 467
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;
<treatmentCitation id="0AEE10FBFFBAFFD7BFEAEA24FE31FB4B" author="Chopard, L." box="[646,851,1142,1164]" page="331" pageId="2" pageNumber="575" year="1968">
<bibRefCitation id="EFDE4B1BFFBAFFD7BFEAEA24FE42FB4B" author="Chopard, L." box="[646,800,1142,1164]" pageId="2" pageNumber="575" pagination="215 - 500" refId="ref6866" refString="Chopard, L. (1968) Family Gryllidae: Subfamilies Mogoplistinae, Myrecophilinae, Scleropterinae, Cachoplistinae, Pteroplistinae, Pentacentrinae, Phalangopsinae, Trigonidiinae, Eneopterinae; Family Oecanthidae, Gryllotalpidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Vol. 12. Uitgeverij Dr. W. Junk N. V. ' s, Gravenhage, pp. 215 - 500." type="book chapter" year="1968">Chopard, 1968</bibRefCitation>
: 331
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;
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<bibRefCitation id="EFDE4B1BFFBAFFD7BE31EA24FE9AFB4B" author="Chopard, L." box="[861,1016,1142,1164]" pageId="2" pageNumber="575" refId="ref6939" refString="Chopard, L. (1969) Grylloidea. In: Sewell, R. B. S. (Ed.), The fauna of India and the adjacent countries. Orthoptera. Vol. 2. Zoological Survey of India and Baptist Mission Press, Calcutta, pp. i-xviii + 1 - 421 + 1." type="journal volume" year="1969">Chopard, 1969</bibRefCitation>
: 295
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;
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: 185
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;
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: 111
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;
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<bibRefCitation id="EFDE4B1BFFBAFFD7BCE0EAC7FC9EFB6C" author="Kim, T. W." box="[396,508,1173,1195]" pageId="2" pageNumber="575" pagination="74 - 83" refId="ref7577" refString="Kim, T. W. (2013) A taxonomic review of the sword-tailed cricket subfamily Trigonidiinae (Orthoptera: Ensifera: Gryllidae) from Korea. Journal of Animal Systematics, Evolution and Diversity, 29 (1), 74 - 83. https: // doi. org / 10.5635 / ASED. 2013.29.1.74" type="journal article" year="2013">Kim, 2013</bibRefCitation>
: 75
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.
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<taxonomicName id="4C4F4D69FFBAFFD7BDFBEAE6FC86FB0D" ID-CoL="8P2D7" authority=": Ichikawa, 2001: 45" authorityName=": Ichikawa" authorityPageNumber="45" authorityYear="2001" box="[151,484,1204,1226]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BDFBEAE6FC76FB0D" box="[151,276,1204,1226]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
:
<treatmentCitation id="0AEE10FBFFBAFFD7BC4DEAE6FC86FB0D" author="Ichikawa, A." box="[289,484,1204,1226]" page="45" pageId="2" pageNumber="575" year="2001">
<bibRefCitation id="EFDE4B1BFFBAFFD7BC4DEAE6FCA3FB0D" author="Ichikawa, A." box="[289,449,1204,1226]" pageId="2" pageNumber="575" pagination="45 - 58" refId="ref7483" refString="Ichikawa, A. (2001) New species of Japanese crickets (Orthoptera; Grylloidea) with notes on certain taxa. Tettigonia: Memoirs of the Orthopterological Society of Japan, 3, 45 - 58." type="journal article" year="2001">Ichikawa, 2001</bibRefCitation>
: 45
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58
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<paragraph id="8BF036EAFFBAFFD7BDFBEAA4FED8FAF5" blockId="2.[151,1437,1270,1871]" pageId="2" pageNumber="575">
The literature treated
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<emphasis id="B93BEAF8FFBAFFD7BCE0EAA4FC96FAC8" box="[396,500,1270,1295]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
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vastly different, either as a distinct genus or a subgenus of
<taxonomicName id="4C4F4D69FFBAFFD7B9F6EAA4F846FAC8" ID-CoL="8P2D7" authorityName=": Ichikawa" authorityYear="2001" box="[1178,1316,1270,1295]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7B9F6EAA4F846FAC8" box="[1178,1316,1270,1295]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
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, and there seems to be no explanation in most literature on the status of this taxon.
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<paragraph id="8BF036EAFFBAFFD7BDABEB6CF87BF994" blockId="2.[151,1437,1270,1871]" pageId="2" pageNumber="575">
The
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species
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<emphasis id="B93BEAF8FFBAFFD7BCE7EB6CFFD3FA90" box="[395,689,1342,1367]" italics="true" pageId="2" pageNumber="575">Trigonidium cicindeloides</emphasis>
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is very similar to
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<emphasis id="B93BEAF8FFBAFFD7BEECEB6CFE8BFA90" box="[896,1001,1342,1367]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
<bibRefCitation id="EFDE4B1BFFBAFFD7BE9CEB6CF900FA90" author="Stal, C." box="[1008,1122,1342,1367]" pageId="2" pageNumber="575" pagination="33 - 58" refId="ref7831" refString="Stal, C. (1877) Orthoptera nova ex Insulis Philippinis descripsit. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlinger, 34 (10), 33 - 58." type="journal article" year="1877">
Stål,
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</taxonomicName>
in that the tympana on fore tibia is generally absent (but not always in
<taxonomicName id="4C4F4D69FFBAFFD7BF1BEB30FFBDFABC" authorityName="Stal" authorityYear="1877" box="[631,735,1378,1403]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BF1BEB30FFBDFABC" box="[631,735,1378,1403]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
</taxonomicName>
), the tegmen is not pubescent but differs by exhibiting a range of yellow and black colouration among the congeners (instead of mostly black). The absence of metanotal glands in males, and tegmen rounded in lateral view (not always) (instead of flat) are also characters.
<taxonomicName id="4C4F4D69FFBAFFD7B9B5EBF8F801FA04" authorityName=": Ichikawa" authorityYear="2001" box="[1241,1379,1450,1475]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7B9B5EBF8F801FA04" box="[1241,1379,1450,1475]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C4F4D69FFBAFFD7BC40EB9CFCF6FA20" authorityName="Stal" authorityYear="1877" box="[300,404,1486,1511]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BC40EB9CFCF6FA20" box="[300,404,1486,1511]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
</taxonomicName>
in the male genitalia (
<figureCitation id="13742A6FFFBAFFD7BFCFEB9CFF8AFA20" box="[675,744,1486,1511]" captionStart="FIGURE 1" captionStartId="3.[151,250,1703,1725]" captionTargetBox="[167,1416,430,1673]" captionTargetId="figure@3.[167,1416,430,1673]" captionTargetPageId="3" captionText="FIGURE 1. Live habitus of Metioche (Metioche) pallipes (Stål, 1861) (A) and Trigonidium cicindeloides Rambur, 1838 (C); male genitalia of Metioche (Metioche) pallipes (B). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2589060/files/figure.png" pageId="2" pageNumber="575">Fig. 1</figureCitation>
). Until a molecular phylogeny can reveal the phylogenetic relationships of
<taxonomicName id="4C4F4D69FFBAFFD7BC38EBA0FCBCF9CC" authorityName=": Ichikawa" authorityYear="2001" box="[340,478,1522,1547]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BC38EBA0FCBCF9CC" box="[340,478,1522,1547]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D69FFBAFFD7BF72EBA0FFE4F9CC" authorityName="Stal" authorityYear="1877" box="[542,646,1522,1547]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BF72EBA0FFE4F9CC" box="[542,646,1522,1547]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
</taxonomicName>
, we consider these taxa as separate genera, despite the similarities mentioned above, owing to
<taxonomicName id="4C4F4D69FFBAFFD7BCB1E844FF27F9E8" authorityName="Stal" authorityYear="1877" box="[477,581,1558,1583]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7BCB1E844FF27F9E8" box="[477,581,1558,1583]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
</taxonomicName>
lacking in the diagnostic characters of
<taxonomicName id="4C4F4D69FFBAFFD7B979E844F9C2F9E8" authorityName=": Ichikawa" authorityYear="2001" box="[1045,1184,1558,1583]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7B979E844F9C2F9E8" box="[1045,1184,1558,1583]" italics="true" pageId="2" pageNumber="575">Trigonidium</emphasis>
</taxonomicName>
: i.e., distal epiphallic lobes small and immovable in relation to the rest of the epiphallus, rachis with apical spine-like lobule.
</paragraph>
<paragraph id="8BF036EAFFBAFFD7BDABE80CFEC4F888" blockId="2.[151,1437,1270,1871]" pageId="2" pageNumber="575">
There are currently 41 species: 1 from Atlantic Oceanic Islands, 7
<collectingCountry id="F358767AFFBAFFD7BEBCE80CF955F9B0" box="[976,1079,1630,1655]" name="Australia" pageId="2" pageNumber="575">Australia</collectingCountry>
and New
<collectingCountry id="F358767AFFBAFFD7B9DDE80CF867F9B0" box="[1201,1285,1630,1655]" name="Guinea-Bissau" pageId="2" pageNumber="575">Guinea</collectingCountry>
, 1
<collectingCountry id="F358767AFFBAFFD7B847E80CF8E2F9B0" box="[1323,1408,1630,1655]" name="Bolivia" pageId="2" pageNumber="575">Bolivia</collectingCountry>
, 1 Borneo, 1
<collectingCountry id="F358767AFFBAFFD7BC63E8D0FC37F95C" box="[271,341,1666,1691]" name="China" pageId="2" pageNumber="575">China</collectingCountry>
, 2
<collectingCountry id="F358767AFFBAFFD7BC14E8D0FCD6F95C" box="[376,436,1666,1691]" name="India" pageId="2" pageNumber="575">India</collectingCountry>
, 4 Indian Oceanic Islands, 2
<collectingCountry id="F358767AFFBAFFD7BE6DE8D1FE26F95D" box="[769,836,1667,1690]" name="Japan" pageId="2" pageNumber="575">Japan</collectingCountry>
, 2 Java, 4
<collectingCountry id="F358767AFFBAFFD7BED3E8D0F92AF95C" box="[959,1096,1666,1691]" name="Madagascar" pageId="2" pageNumber="575">Madagascar</collectingCountry>
, 1 Malay Peninsula, 3 Pacific Oceanic Islands, 1
<collectingCountry id="F358767AFFBAFFD7BC19E8F5FCC9F979" box="[373,427,1703,1726]" name="Peru" pageId="2" pageNumber="575">Peru</collectingCountry>
, 1
<collectingCountry id="F358767AFFBAFFD7BCBDE8F4FF30F978" box="[465,594,1702,1727]" name="Philippines" pageId="2" pageNumber="575">Philippines</collectingCountry>
, 1
<collectingCountry id="F358767AFFBAFFD7BF14E8F4FFAFF978" box="[632,717,1702,1727]" name="Taiwan" pageId="2" pageNumber="575">Taiwan</collectingCountry>
, 3 central Africa, 3 eastern Africa, and 4 western Africa. It is likely that most species need closer examination to verify their generic statuses and it is likely that
<taxonomicName id="4C4F4D69FFBAFFD7B87BE898F81DF924" authorityName="Stal" authorityYear="1877" box="[1303,1407,1738,1763]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF8FFBAFFD7B87BE898F81DF924" box="[1303,1407,1738,1763]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
</taxonomicName>
is currently not monophyletic.
<taxonomicName id="4C4F4D69FFBAFFD7BCB5E8BCFED0F8C0" authority="(Stal, 1861)" baseAuthorityName="Stal" baseAuthorityYear="1861" box="[473,946,1774,1799]" class="Insecta" family="Trigonidiidae" genus="Metioche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="pallipes" subGenus="Metioche">
<emphasis id="B93BEAF8FFBAFFD7BCB5E8BCFF23F8C0" box="[473,577,1774,1799]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
(
<emphasis id="B93BEAF8FFBAFFD7BF3FE8BCFFD9F8C0" box="[595,699,1774,1799]" italics="true" pageId="2" pageNumber="575">Metioche</emphasis>
)
<emphasis id="B93BEAF8FFBAFFD7BFA0E8BCFE47F8C0" box="[716,805,1774,1799]" italics="true" pageId="2" pageNumber="575">pallipes</emphasis>
(Stål, 1861)
</taxonomicName>
is synoptic with
<taxonomicName id="4C4F4D69FFBAFFD7B919E8BCF8FEF8C0" authorityName="Rambur" authorityYear="1838" box="[1141,1436,1774,1799]" class="Insecta" family="Trigonidiidae" genus="Trigonidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="cicindeloides">
<emphasis id="B93BEAF8FFBAFFD7B919E8BCF8FEF8C0" box="[1141,1436,1774,1799]" italics="true" pageId="2" pageNumber="575">Trigonidium cicindeloides</emphasis>
</taxonomicName>
in the formers
<typeStatus id="54F48848FFBAFFD7BC26E941FC18F8EC" box="[330,378,1811,1835]" pageId="2" pageNumber="575">type</typeStatus>
locality,
<collectingCountry id="F358767AFFBAFFD7BC89E940FF35F8EC" box="[485,599,1810,1835]" name="Singapore" pageId="2" pageNumber="575">Singapore</collectingCountry>
(Tan, 2012) (
<figureCitation id="13742A6FFFBAFFD7BF99E940FE5BF8EC" box="[757,825,1810,1835]" captionStart="FIGURE 1" captionStartId="3.[151,250,1703,1725]" captionTargetBox="[167,1416,430,1673]" captionTargetId="figure@3.[167,1416,430,1673]" captionTargetPageId="3" captionText="FIGURE 1. Live habitus of Metioche (Metioche) pallipes (Stål, 1861) (A) and Trigonidium cicindeloides Rambur, 1838 (C); male genitalia of Metioche (Metioche) pallipes (B). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2589060/files/figure.png" pageId="2" pageNumber="575">Fig. 1</figureCitation>
). Specifically, the two species co-occur in the grassy areas of Bidadari which is currently cleared for housing development.
</paragraph>
</subSubSection>
</treatment>
</document>