511 lines
61 KiB
XML
511 lines
61 KiB
XML
<document id="C40C9E6C3828C844C1FA1586FD523174" ID-DOI="10.11646/zootaxa.4094.1.1" ID-GBIF-Dataset="60d01d07-7735-4a32-9292-82ead909e804" ID-ISSN="1175-5326" ID-Zenodo-Dep="266544" ID-ZooBank="E5F1E766-687D-4B00-974B-8D7939DC66A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459242848466" checkinUser="plazi" docAuthor="David G. Herbert & Adnan Mousalli" docDate="2016" docId="03E687FCE025D76CFF1A11FA0FFA9CC0" docLanguage="en" docName="zootaxa.4094.1.pdf.imf" docOrigin="Zootaxa 4094 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Nata erugata Herbert & Mousalli, 2016, sp. nov." docType="treatment" docVersion="11" lastPageNumber="44" masterDocId="FFDFFF84E00DD747FF8D116D0B0A9828" masterDocTitle="Revision of the dwarf cannibal snails (Nata s. l.) of southern Africa — Nata s. s. and Natella (Mollusca: Gastropoda: Rhytididae), with description of three new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="41" updateTime="1698186665173" updateUser="plazi">
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<mods:mods id="FE6E19D19980C7F9EA2EF6233C8E2CAD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2B82A3BDD4ACC6C648CA88BC5D70CD95">
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<mods:title id="7733300663832C2022979B2132B7E09E">Revision of the dwarf cannibal snails (Nata s. l.) of southern Africa — Nata s. s. and Natella (Mollusca: Gastropoda: Rhytididae), with description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="3F9286DCBA17760B3337B6309FAC93DA" type="personal">
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<mods:role id="258EEB0DD79990BBD2DEBF92DF0990EF">
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<mods:roleTerm id="5C72B349366F379681B7DBF50F15F457">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF402888393EBCEAA7AEB3061129B355">David G. Herbert</mods:namePart>
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</mods:name>
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<mods:name id="BC436BD97DB81FFFB4906AA38A39F07D" type="personal">
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<mods:role id="D8745E31F167EB6991C72ADA391D34BC">
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<mods:roleTerm id="E586850E1FE3EFB98421C0CEE6B48122">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A6BAAEF8F50990EB316A0291EB19B405">Adnan Mousalli</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C9D3AB3A1934073E7ABFEA7CA281CA25">text</mods:typeOfResource>
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<mods:titleInfo id="51F2AD5E0218F79BFD226D9FB7EDCCC3">
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<mods:title id="4E15E933AD7337C6322DAD0D303065D2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="543C85CF236C905C8E6D6C95A94F8985">
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<mods:date id="415A924B265639CF5783F5CCA3BCAC49">2016</mods:date>
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<mods:detail id="325A19A674468BEB5CC3A7AE1968C2D9" type="volume">
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<mods:number id="68C4E2E07FA8E5226740BFAA783BB9F8">4094</mods:number>
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</mods:detail>
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<mods:detail id="9B53E7EAC61D3B3F637226B0857DD148" type="issue">
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<mods:number id="631510E438C91A7B24A6D6F56603CA36">1</mods:number>
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<mods:end id="FF67E2F8AD0D340395098E2CBBBBFB08">67</mods:end>
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</mods:part>
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<mods:classification id="7EC28F4BED6BC43F229145D8446E2989">journal article</mods:classification>
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<mods:identifier id="2E8188D9653E6534DE0E85868BB1B703" type="DOI">10.11646/zootaxa.4094.1.1</mods:identifier>
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<mods:identifier id="BE021905EBAA06CF68C75F3A6949A658" type="GBIF-Dataset">60d01d07-7735-4a32-9292-82ead909e804</mods:identifier>
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<mods:identifier id="22DB5810D232FCDB470CD9B7935A3D5A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E5ED2198A07CFE80803F130A2AED76FF" type="Zenodo-Dep">266544</mods:identifier>
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<mods:identifier id="12A9F2D758423D4DCEFC5559559413FD" type="ZooBank">E5F1E766-687D-4B00-974B-8D7939DC66A0</mods:identifier>
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</mods:mods>
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<treatment id="03E687FCE025D76CFF1A11FA0FFA9CC0" ID-DOI="http://doi.org/10.5281/zenodo.5615856" ID-GBIF-Taxon="119319683" ID-Zenodo-Dep="5615856" LSID="urn:lsid:plazi:treatment:03E687FCE025D76CFF1A11FA0FFA9CC0" httpUri="http://treatment.plazi.org/id/03E687FCE025D76CFF1A11FA0FFA9CC0" lastPageId="43" lastPageNumber="44" pageId="40" pageNumber="41">
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<subSubSection id="C3556561E025D76FFF1A11FA0A9C98FB" pageId="40" pageNumber="41" type="nomenclature">
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<paragraph id="8BF036EAE025D76FFF1A11FA0A90989A" blockId="40.[151,410,151,211]" box="[151,410,151,178]" pageId="40" pageNumber="41">
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<heading id="D0B88186E025D76FFF1A11FA0A90989A" bold="true" box="[151,410,151,178]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
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<emphasis id="B93BEAF8E025D76FFF1A11FA0A90989A" bold="true" box="[151,410,151,178]" pageId="40" pageNumber="41">
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<taxonomicName id="4C4F4D69E025D76FFF1A11FA0A3F9899" box="[151,309,151,177]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
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<emphasis id="B93BEAF8E025D76FFF1A11FA0A3F9899" bold="true" box="[151,309,151,177]" italics="true" pageId="40" pageNumber="41">Nata erugata</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A2085783E025D76FFEB111F50A90989A" box="[316,410,152,178]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BF036EAE025D76FFF1A11D70A9C98FB" blockId="40.[151,410,151,211]" box="[151,406,186,211]" pageId="40" pageNumber="41">
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||
<figureCitation id="13742A6FE025D76FFF1A11D70A0B98FB" box="[151,257,186,211]" captionStart="FIGURE 4" captionStartId="9.[151,250,1791,1813]" captionTargetBox="[184,1395,199,1756]" captionTargetId="figure@9.[184,1396,193,1762]" captionTargetPageId="9" captionText="FIGURE 4. Living animals of Nata and Natella. A, Nata aequiplicata sp. nov., holotype, Stinkhoutkloof, E. Cape, shell diameter 6.5 mm (NMSA W 9248 / T 3066). B, Nata dumeticola, West Coast National Park, W. Cape, shell diameter 10.8 mm (NMSA W 3224). C, Nata erugata sp. nov., Tullach-Mohr, Nelspruit area, Mpumalanga, shell diameter 20 mm (MV ex NMSA W 3301). D, Nata tarachodes, Grootbos Private Nat. Res., Gansbaai, W. Cape, shell diameter 7.7 mm (NMSA W 5658). E, Nata watsoni sp. nov., Hankey, E. Cape, shell diameter 7.5 mm (paratype NMSA W 6524 / T 3921). F, Nata watsoni sp. nov. Camdeboo National Park, Graaff-Reinet, E. Cape, shell diameter 13.0 mm (paratype, NMSA W 6091 / T 2303). G, Nata vernicosa, Vernon Crookes Nat. Res., KwaZulu-Natal, shell diameter 14.0 mm (NMSA W 8020). H, Natella viridescens, Ithala Game Res., KwaZulu-Natal, shell diameter 3.7 mm (NMSA W 7263)." httpUri="https://zenodo.org/record/266548/files/figure.png" pageId="40" pageNumber="41">Figures 4</figureCitation>
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C, 6C, 29–32
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3556561E025D76FFF1A106F0AFA997E" pageId="40" pageNumber="41" type="description">
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||
<paragraph id="8BF036EAE025D76FFF1A106F09AC9930" blockId="40.[151,1436,258,342]" box="[151,678,258,280]" pageId="40" pageNumber="41">
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||
<treatmentCitationGroup id="AB5F11C4E025D76FFF1A106F09AC9930" box="[151,678,258,280]" pageId="40" pageNumber="41">
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||
<taxonomicName id="4C4F4D69E025D76FFF1A106F0A4C9930" box="[151,326,258,280]" class="Gastropoda" family="Rhytididae" genus="Natalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="caffrula">
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<emphasis id="B93BEAF8E025D76FFF1A106F0A4C9930" box="[151,326,258,280]" italics="true" pageId="40" pageNumber="41">Natalina caffrula</emphasis>
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</taxonomicName>
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||
—
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||
<treatmentCitation id="0AEE10FBE025D76FFEEC106E0AFD9930" author="Haas" box="[353,503,258,280]" page="18" pageId="40" pageNumber="41" year="1936">
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||
<bibRefCitation id="EFDE4B1BE025D76FFEEC106E0ADE9930" author="Haas" box="[353,468,258,280]" pageId="40" pageNumber="57" refString="Haas, F. (1936) Binnen-Mollusken aus Inner-Afrika, hauptsachlich gesammelt von Dr. F. Haas wahrend der Schomburgk- Expedition in den Jahren 1931 / 32. Abhandlungen Senckenbergischen Naturforschenden Gesellschaft, 431, 1 - 156." type="journal article" year="1936">Haas, 1936</bibRefCitation>
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||
: 18
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||
</treatmentCitation>
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||
(in part, Mkuzi).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BF036EAE025D76FFF1A104F0AFA997E" blockId="40.[151,1436,258,342]" pageId="40" pageNumber="41">
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||
<treatmentCitationGroup id="AB5F11C4E025D76FFF1A104F0AFA997E" pageId="40" pageNumber="41">
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||
<taxonomicName id="4C4F4D69E025D76FFF1A104F0A3C991F" box="[151,310,290,311]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="vernicosa">
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<emphasis id="B93BEAF8E025D76FFF1A104F0A3C991F" box="[151,310,290,311]" italics="true" pageId="40" pageNumber="41">Nata vernicosa</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="0AEE10FBE025D76FFEDC104F092D991F" author="Bruggen" box="[337,551,289,311]" page="376" pageId="40" pageNumber="41" year="1966">
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||
<bibRefCitation id="EFDE4B1BE025D76FFEDC104F0AE5991F" author="Bruggen" box="[337,495,289,311]" pageId="40" pageNumber="56" refString="Bruggen, A. C. van (1966) The terrestrial Mollusca of the Kruger National Park: a contribution to the malacology of the Eastern Transvaal. Annals of the Natal Museum, 18, 315 - 399." type="journal article" year="1966">Bruggen, 1966</bibRefCitation>
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: 376
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||
</treatmentCitation>
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||
, fig. 54.
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<treatmentCitation id="0AEE10FBE025D76FFD1D104F0851991F" author="Bruggen" box="[656,859,289,311]" page="37" pageId="40" pageNumber="41" year="1969">
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||
<bibRefCitation id="EFDE4B1BE025D76FFD1D104F0825991F" author="Bruggen" box="[656,815,289,311]" pageId="40" pageNumber="56" refString="Bruggen, A. C. van (1969) Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in southern Africa. Zoologische Verhandelingen, 103, 1 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
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: 37
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</treatmentCitation>
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||
.
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<treatmentCitation id="0AEE10FBE025D76FFCE7104F0F40991F" author="Bruggen" box="[874,1098,289,311]" page="468" pageId="40" pageNumber="41" year="1970">
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||
<bibRefCitation id="EFDE4B1BE025D76FFCE7104F0F1C991F" author="Bruggen" box="[874,1046,289,311]" pageId="40" pageNumber="56" refString="Bruggen, A. C. van (1970 a) Non-marine Mollusca. Chapter 18. In: Hanstrom, B., Brinck, P. & Rudebeck, G. (Eds.), South African Animal Life. Results of the Lund University Expedition in 1950 - 1951. Vol XIV. Statens Naturvetenskapliga Forskningsrad, Stockholm, pp. 445 - 476." type="book chapter" year="1970" yearSuffix="a">Bruggen, 1970a</bibRefCitation>
|
||
: 468
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||
</treatmentCitation>
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||
(in part).
|
||
<treatmentCitation id="0AEE10FBE025D76FFB4D104F0E93991F" author="Bruggen" box="[1216,1433,289,311]" page="59" pageId="40" pageNumber="41" year="1970">
|
||
<bibRefCitation id="EFDE4B1BE025D76FFB4D104F0E66991F" author="Bruggen" box="[1216,1388,289,311]" pageId="40" pageNumber="56" refString="Bruggen, A. C. van (1970 b) The terrestrial Mollusca of the Mkuzi and Ndumu Game Reserves, Zululand. Lammergeyer, 11, 58 - 59." type="journal article" year="1970" yearSuffix="b">Bruggen, 1970b</bibRefCitation>
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||
: 59
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</treatmentCitation>
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||
.
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||
<treatmentCitation id="0AEE10FBE025D76FFF4A102C0A9B997E" author="Bruggen" box="[199,401,320,342]" page="286" pageId="40" pageNumber="41" year="1985">
|
||
<bibRefCitation id="EFDE4B1BE025D76FFF4A102C0A6B997E" author="Bruggen" box="[199,353,320,342]" pageId="40" pageNumber="56" refString="Bruggen, A. C. van (1985) The terrestrial molluscs of Lesotho (southern Africa), a first contribution, with detailed notes on Archachatina machachensis (Mollusca, Gastropoda). Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, Series C, 88, 267 - 296." type="journal article" year="1985">Bruggen, 1985</bibRefCitation>
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||
: 286
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||
</treatmentCitation>
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||
(in part).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3556561E025D76FFF1A10EF0FAF99B3" box="[151,1189,386,411]" pageId="40" pageNumber="41" type="etymology">
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||
<paragraph id="8BF036EAE025D76FFF1A10EF0FAF99B3" blockId="40.[151,1437,386,1851]" box="[151,1189,386,411]" pageId="40" pageNumber="41">
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||
<emphasis id="B93BEAF8E025D76FFF1A10EF0A1499B3" bold="true" box="[151,286,386,411]" pageId="40" pageNumber="41">Etymology.</emphasis>
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From
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<emphasis id="B93BEAF8E025D76FFEE510E90AA399B3" box="[360,425,388,411]" italics="true" pageId="40" pageNumber="41">erugo</emphasis>
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(L.)—to clear of wrinkles,
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<emphasis id="B93BEAF8E025D76FFD5310E9084A99B3" box="[734,832,388,411]" italics="true" pageId="40" pageNumber="41">erugatus</emphasis>
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||
(L.)–without wrinkles, smooth.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3556561E025D76FFF4A10CB08189CD3" pageId="40" pageNumber="41" type="materials_examined">
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||
<paragraph id="8BF036EAE025D76FFF4A10CB0A419A2F" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">
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||
<emphasis id="B93BEAF8E025D76FFF4A10CB0A889997" bold="true" box="[199,386,422,447]" pageId="40" pageNumber="41">
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||
<typeStatus id="54F48848E025D76FFF4A10CB0A089997" box="[199,258,422,447]" pageId="40" pageNumber="41">Type</typeStatus>
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||
material.
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||
</emphasis>
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||
<typeStatus id="54F48848E025D76FFE1810CB0AF79997" box="[405,509,422,447]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
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||
(
|
||
<figureCitation id="13742A6FE025D76FFD9710CB09739997" box="[538,633,422,447]" captionStart="FIGURE 30" captionStartId="41.[151,250,1167,1189]" captionTargetBox="[155,1431,204,1137]" captionTargetId="figure@41.[151,1436,193,1145]" captionTargetPageId="41" captionText="FIGURE 30. Nata erugata sp. nov. A – C, holotype, Monk’s Cowl Forest, KwaZulu-Natal, diameter 15.5 mm (NMSA W 5599 / T 3906). D – J, additional specimens from throughout the range of the species [D, Majuba, KwaZulu-Natal, diameter 14.5 mm (paratype, NMSA V 5932 / T 3909). E, Pongola Bush, KwaZulu-Natal, diameter 17.0 mm (paratype, NMSA V 776 / T 3915). F, Phinda Game Reserve, KwaZulu-Natal, diameter 16.2 mm (W 1334). G, Balloch, Barkly East, E. Cape, diameter 16.8 mm (paratype, NMSA W 4836 / T 3914). H, Mapaya Hill, Zastron, Free State, diameter 13.2 mm (paratype, W 6573 / T 3907). I, Mariepskop, Mpumalanga, diameter 17.1 mm (W 9364). J, Sand River area, Kruger National Park, diameter 15.6 mm (A 9759)]." httpUri="https://zenodo.org/record/266574/files/figure.png" pageId="40" pageNumber="41">Fig. 30</figureCitation>
|
||
A–C):
|
||
<collectingCountry id="F358767AE025D76FFD5F10CA08A49997" box="[722,942,423,447]" name="South Africa" pageId="40" pageNumber="41">SOUTH AFRICA</collectingCountry>
|
||
: KWAZULU-NATAL: Monk’s Cowl area (
|
||
<geoCoordinate id="EE7B502DE025D76FFF1210A70A2299CB" box="[159,296,458,483]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-29.05479">29.05479°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFEBB10A60ACA99CB" box="[310,448,459,483]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="29.38287">29.38287°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFE5B10A7093B99CA" box="[470,561,458,483]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="40" pageNumber="41" unit="m" value="1625.0">1625 m</quantity>
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||
, indigenous forest, in leaf-litter, F78, M. Hamer
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||
<emphasis id="B93BEAF8E025D76FFBE210A10FA899CB" box="[1135,1186,458,483]" italics="true" pageId="40" pageNumber="41">et al</emphasis>
|
||
.,
|
||
<date id="FFF1102AE025D76FFB3110A70E3D99CB" box="[1212,1335,458,483]" pageId="40" pageNumber="41" value="2006-03-16">16/iii/2006</date>
|
||
(
|
||
<collectionCode id="ED5EAE2FE025D76FFAC410A60E9699CB" LSID="urn:lsid:biocol.org:col:34985" box="[1353,1436,459,483]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="40" pageNumber="41">NMSA</collectionCode>
|
||
W5599/T3906).
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE025D76FFF4A137E09AE9C6F" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">
|
||
<typeStatus id="54F48848E025D76FFF4A137E0A3E9A03" box="[199,308,531,555]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
|
||
(all
|
||
<collectionCode id="ED5EAE2FE025D76FFEE7137E0AB19A03" LSID="urn:lsid:biocol.org:col:34985" box="[362,443,531,555]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="40" pageNumber="41">NMSA</collectionCode>
|
||
unless indicated otherwise):
|
||
<collectingCountry id="F358767AE025D76FFC8A137E08D29A03" box="[775,984,531,555]" name="South Africa" pageId="40" pageNumber="41">SOUTH AFRICA</collectingCountry>
|
||
: KWAZULU-NATAL: Pongola Bush Nat. Res. (
|
||
<geoCoordinate id="EE7B502DE025D76FFF5A135A0A4E9A67" box="[215,324,567,591]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="55" value="-27.323">27.323°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFEC2135A0AC09A67" box="[335,458,567,591]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="30.4817">30.4817°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFE50135A093E9A66" box="[477,564,567,591]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="40" pageNumber="41" unit="m" value="1300.0">1300 m</quantity>
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||
, montane
|
||
<taxonomicName id="4C4F4D69E025D76FFD25135B08259A67" box="[680,815,566,591]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93BEAF8E025D76FFD25135B08259A67" box="[680,815,566,591]" italics="true" pageId="40" pageNumber="41">Podocarpus</emphasis>
|
||
</taxonomicName>
|
||
forest, in leaf-litter and under logs, D. Herbert & NPB,
|
||
<date id="FFF1102AE025D76FFF1A13370A089A5B" box="[151,258,602,627]" pageId="40" pageNumber="41" value="1995-01-19">19/i/1995</date>
|
||
(V776/T3915, one adult and three juvenile specimens;
|
||
<collectionCode id="ED5EAE2FE025D76FFCFF133608A79A5A" box="[882,941,603,626]" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="40" pageNumber="41">ELM</collectionCode>
|
||
D18003/T031, one specimen); Majuba area (
|
||
<geoCoordinate id="EE7B502DE025D76FFF1213120A069ABF" box="[159,268,639,663]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="55" value="-27.477">27.477°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFE9B13130A899ABF" box="[278,387,638,663]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="55" value="29.685">29.685°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFE1B13120AE69ABE" box="[406,492,639,663]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="40" pageNumber="41" unit="m" value="1700.0">1700 m</quantity>
|
||
, montane
|
||
<taxonomicName id="4C4F4D69E025D76FFDD3131309EF9ABF" box="[606,741,638,663]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93BEAF8E025D76FFDD3131309EF9ABF" box="[606,741,638,663]" italics="true" pageId="40" pageNumber="41">Podocarpus</emphasis>
|
||
</taxonomicName>
|
||
forest, in leaf-litter, D. Herbert,
|
||
<date id="FFF1102AE025D76FFBC313130FB39ABF" box="[1102,1209,638,663]" pageId="40" pageNumber="41" value="1998-01-28">28/i/1998</date>
|
||
(V5932/T3909, one adult and three juvenile specimens);
|
||
<collectingCountry id="F358767AE025D76FFDB913CE095F9A92" box="[564,597,675,698]" name="South Georgia and the South Sandwich Islands" pageId="40" pageNumber="41">De</collectingCountry>
|
||
Beers farm, approx.
|
||
<quantity id="4CB79B0FE025D76FFCCF13CF08809A93" box="[834,906,674,699]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="40" pageNumber="41" unit="km" value="15.0">15 km</quantity>
|
||
NE of Van Reenen (
|
||
<geoCoordinate id="EE7B502DE025D76FFBFF13CE0FF69A93" box="[1138,1276,675,699]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-28.30403">28.30403°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFA8A13CF0E9B9A93" box="[1287,1425,674,699]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="29.5024">29.50240°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFF1A13AA0BFB9AF6" box="[151,241,711,735]" metricMagnitude="3" metricUnit="m" metricValue="1.841" pageId="40" pageNumber="41" unit="m" value="1841.0">1841 m</quantity>
|
||
, afrotemperate forest, in leaf-litter, A. Moussalli, D. Stuart-Fox & M. Cunningham,
|
||
<date id="FFF1102AE025D76FFB4613AB0E249AF7" box="[1227,1326,710,735]" pageId="40" pageNumber="41" value="2006-10-09">9/x/2006</date>
|
||
(W4688/ T3910, one specimen); Van Reenen (
|
||
<geoCoordinate id="EE7B502DE025D76FFDBB138709A89B2B" box="[566,674,746,771]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="55" value="-28.405">28.405°S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE7B502DE025D76FFD23138608169B2B" box="[686,796,747,771]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="55" value="29.39">29.390°E</geoCoordinate>
|
||
),
|
||
<date id="FFF1102AE025D76FFCA21387088D9B2B" box="[815,903,746,771]" bridgedPair="/" pageId="40" pageNumber="41" value="1918-03">iii/1918</date>
|
||
(A7069/T3911, five specimens;
|
||
<collectionCode id="ED5EAE2FE025D76FFB7513860E399B2A" box="[1272,1331,747,770]" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="40" pageNumber="41">ELM</collectionCode>
|
||
D18002/ T030, one specimen); Kranskop (
|
||
<geoCoordinate id="EE7B502DE025D76FFDBC126309979B0F" box="[561,669,782,807]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="55" value="-28.967">28.967°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFD3C126308149B0F" box="[689,798,782,807]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="55" value="30.863">30.863°E</geoCoordinate>
|
||
), Falcon coll’n,
|
||
<date id="FFF1102AE025D76FFC6212630F3D9B0F" box="[1007,1079,782,807]" bridgedPair="/" pageId="40" pageNumber="41" value="1922-01">i/1922</date>
|
||
(
|
||
<collectionCode id="ED5EAE2FE025D76FFBC312620FAB9B0F" LSID="urn:lsid:biocol.org:col:34985" box="[1102,1185,783,807]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="40" pageNumber="41">NMSA</collectionCode>
|
||
A7070/T3912, three specimens).
|
||
<collectionCode id="ED5EAE2FE025D76FFEA9125E0AA79B63" box="[292,429,819,843]" pageId="40" pageNumber="41">FREE STATE</collectionCode>
|
||
: Platberg Nat. Res. (
|
||
<geoCoordinate id="EE7B502DE025D76FFD14125F08289B63" box="[665,802,818,843]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-28.27631">28.27631°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFCA3125E08B29B63" box="[814,952,819,843]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="29.20772">29.20772°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFC46125E0F299B62" box="[971,1059,819,843]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="40" pageNumber="41" unit="m" value="2100.0">2100 m</quantity>
|
||
, montane shrubland/grassland, in leaf-litter, A. Moussalli, D. Stuart-Fox & M. Cunningham,
|
||
<date id="FFF1102AE025D76FFCCC123B08CB9B47" box="[833,961,854,879]" pageId="40" pageNumber="41" value="2006-12-11">11/xii/2006</date>
|
||
(W5621/T3908, two specimens); Golden Gate Highlands National Park (
|
||
<geoCoordinate id="EE7B502DE025D76FFD8E121609869BBB" box="[515,652,891,915]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-28.4943">28.49430°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFD171217082E9BBB" box="[666,804,890,915]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="28.61868">28.61868°E</geoCoordinate>
|
||
),
|
||
<quantity id="4CB79B0FE025D76FFCB71216089E9BBA" box="[826,916,891,915]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="40" pageNumber="41" unit="m" value="2000.0">2000 m</quantity>
|
||
, gully thicket/grassland, buried in shrubs, A. Moussalli & D. Stuart-Fox,
|
||
<date id="FFF1102AE025D76FFE5512F309539B9F" box="[472,601,926,951]" pageId="40" pageNumber="41" value="2005-12-28">28/xii/2005</date>
|
||
(W4243/T3913, one specimen); Zastron area, Mapaya Hill (
|
||
<geoCoordinate id="EE7B502DE025D76FFA8212F20E939B9F" box="[1295,1433,927,951]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-30.27439">30.27439°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFF1A12AE0A2B9BF3" box="[151,289,963,987]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="27.21902">27.21902°E</geoCoordinate>
|
||
),
|
||
<emphasis id="B93BEAF8E025D76FFEBA12A90A4C9BF3" box="[311,326,964,987]" italics="true" pageId="40" pageNumber="41">~</emphasis>
|
||
<quantity id="4CB79B0FE025D76FFECB12AF0AAA9BF2" box="[326,416,962,987]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="40" pageNumber="41" unit="m" value="1860.0">1860 m</quantity>
|
||
, in soil amongst boulders in narrow, shaded cleft above sandstone cliff, st'n 08-059, D. Herbert & L. Davis,
|
||
<date id="FFF1102AE025D76FFE07128B090E9BD7" box="[394,516,998,1023]" pageId="40" pageNumber="41" value="2008-11-22">22/xi/2008</date>
|
||
(W6573/T3907, three specimens). E.
|
||
<collectionCode id="ED5EAE2FE025D76FFC33128A08F19BD7" box="[958,1019,999,1023]" pageId="40" pageNumber="41">CAPE</collectionCode>
|
||
: Witteberge, Balloch farm, north of Barkly East (
|
||
<geoCoordinate id="EE7B502DE025D76FFEA415670AB89C0B" box="[297,434,1034,1059]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-30.71255">30.71255°S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE7B502DE025D76FFE311567094C9C0B" box="[444,582,1034,1059]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="27.69507">27.69507°E</geoCoordinate>
|
||
),
|
||
<collectingCountry id="F358767AE025D76FFDD4156709B99C0B" box="[601,691,1034,1059]" name="Lesotho" pageId="40" pageNumber="41">Lesotho</collectingCountry>
|
||
highland grassland, under rock, D. Stuart-Fox,
|
||
<date id="FFF1102AE025D76FFB4A15670E3B9C0B" box="[1223,1329,1034,1059]" pageId="40" pageNumber="41" value="2005-04-01">1/iv/2005</date>
|
||
(W4836/ T3914, one adult and two juvenile specimens).
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE025D76FFF4A153F09FD9C43" blockId="40.[151,1437,386,1851]" box="[199,759,1106,1131]" pageId="40" pageNumber="41">
|
||
<emphasis id="B93BEAF8E025D76FFF4A153F093C9C43" bold="true" box="[199,566,1106,1131]" pageId="40" pageNumber="41">Additional material examined:</emphasis>
|
||
See Appendix 4.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE025D76FFF4A151B08189CD3" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">
|
||
<emphasis id="B93BEAF8E025D76FFF4A151B0A7A9CA7" bold="true" box="[199,368,1142,1167]" pageId="40" pageNumber="41">Identification.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="4C4F4D69E025D76FFD80151A09B39CA6" box="[525,697,1143,1166]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E025D76FFD80151A09B39CA6" box="[525,697,1143,1166]" italics="true" pageId="40" pageNumber="41">Nata vernicosa</emphasis>
|
||
</taxonomicName>
|
||
in that the apical surface is almost completely devoid of regular, close-set, axial riblets; although axial sculpture remains, it is present only in the form of weak, irregularly spaced growth-lines. More similar to
|
||
<taxonomicName id="4C4F4D69E025D76FFE7E15D209809CFE" box="[499,650,1215,1238]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E025D76FFE7E15D209809CFE" box="[499,650,1215,1238]" italics="true" pageId="40" pageNumber="41">Nata watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E025D76FFD1915D309FA9CFF" box="[660,752,1214,1239]" pageId="40" pageNumber="41" rank="species">
|
||
<emphasis id="B93BEAF8E025D76FFD1915D309FA9CFF" bold="true" box="[660,752,1214,1239]" pageId="40" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but that species has very different radula morphology and occurs in a different region and biome (Albany Thicket).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E025D76EFF4A146B0E2C9FBD" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="description">
|
||
<paragraph id="8BF036EAE025D76FFF4A146B096A9EAF" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">
|
||
<emphasis id="B93BEAF8E025D76FFF4A146B0A739D37" bold="true" box="[199,377,1286,1311]" pageId="40" pageNumber="41">
|
||
Shell (
|
||
<figureCitation id="13742A6FE025D76FFE9F146B0A609D37" box="[274,362,1286,1311]" captionStart="FIGURE 30" captionStartId="41.[151,250,1167,1189]" captionTargetBox="[155,1431,204,1137]" captionTargetId="figure@41.[151,1436,193,1145]" captionTargetPageId="41" captionText="FIGURE 30. Nata erugata sp. nov. A – C, holotype, Monk’s Cowl Forest, KwaZulu-Natal, diameter 15.5 mm (NMSA W 5599 / T 3906). D – J, additional specimens from throughout the range of the species [D, Majuba, KwaZulu-Natal, diameter 14.5 mm (paratype, NMSA V 5932 / T 3909). E, Pongola Bush, KwaZulu-Natal, diameter 17.0 mm (paratype, NMSA V 776 / T 3915). F, Phinda Game Reserve, KwaZulu-Natal, diameter 16.2 mm (W 1334). G, Balloch, Barkly East, E. Cape, diameter 16.8 mm (paratype, NMSA W 4836 / T 3914). H, Mapaya Hill, Zastron, Free State, diameter 13.2 mm (paratype, W 6573 / T 3907). I, Mariepskop, Mpumalanga, diameter 17.1 mm (W 9364). J, Sand River area, Kruger National Park, diameter 15.6 mm (A 9759)]." httpUri="https://zenodo.org/record/266574/files/figure.png" pageId="40" pageNumber="41">Fig. 30</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Lenticular, occasionally discoidal; whorls comparatively shallow and spire often low; largest specimens comprising 3.25–3.5 teleoconch whorls; periphery at or more usually slightly below mid-whorl; suture above periphery, at most weakly descendant prior to aperture. Protoconch diameter 1.0–
|
||
<quantity id="4CB79B0FE025D76FFBF214220FDD9D4E" box="[1151,1239,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="40" pageNumber="41" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, comprising ±1.0 whorl, smooth and glossy, junction with teleoconch usually rather weakly defined. Apical surface of teleoconch smooth and glossy, lacking regular axial riblets and sculptured only by weak growth irregularities; these strongest below suture, weakest at periphery and on base; no significant spiral sculpture, but some specimens with faint traces of vermiform wrinkles in places. Base and umbilical margin evenly rounded; umbilicus of moderate width, steep-sided; growth-lines slightly stronger within umbilicus. Aperture roundly or distinctly ovate, often obliquely descendant (
|
||
<figureCitation id="13742A6FE025D76FFEA1174B0A839E17" box="[300,393,1574,1599]" captionStart="FIGURE 30" captionStartId="41.[151,250,1167,1189]" captionTargetBox="[155,1431,204,1137]" captionTargetId="figure@41.[151,1436,193,1145]" captionTargetPageId="41" captionText="FIGURE 30. Nata erugata sp. nov. A – C, holotype, Monk’s Cowl Forest, KwaZulu-Natal, diameter 15.5 mm (NMSA W 5599 / T 3906). D – J, additional specimens from throughout the range of the species [D, Majuba, KwaZulu-Natal, diameter 14.5 mm (paratype, NMSA V 5932 / T 3909). E, Pongola Bush, KwaZulu-Natal, diameter 17.0 mm (paratype, NMSA V 776 / T 3915). F, Phinda Game Reserve, KwaZulu-Natal, diameter 16.2 mm (W 1334). G, Balloch, Barkly East, E. Cape, diameter 16.8 mm (paratype, NMSA W 4836 / T 3914). H, Mapaya Hill, Zastron, Free State, diameter 13.2 mm (paratype, W 6573 / T 3907). I, Mariepskop, Mpumalanga, diameter 17.1 mm (W 9364). J, Sand River area, Kruger National Park, diameter 15.6 mm (A 9759)]." httpUri="https://zenodo.org/record/266574/files/figure.png" pageId="40" pageNumber="41">Fig. 30</figureCitation>
|
||
J); peristome interrupted by bulging parietal region; outer lip thin with membranous periostracal fringe (commonly lost); outer lip shallowly sinuous when undamaged; upper part of columella lip extending forward and slightly reflected.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE025D76FFF4A17FF0A439EDB" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">Shell translucent milky-white when fresh; periostracum glossy, pale yellowish-brown to honey-gold, commonly with some axial variation in intensity, producing unevenly spaced, slightly darker axial banding, but this generally weak.</paragraph>
|
||
<paragraph id="8BF036EAE025D76FFF4A179309489F13" blockId="40.[151,1437,386,1851]" pageId="40" pageNumber="41">
|
||
Dimensions:
|
||
<typeStatus id="54F48848E025D76FFED017930AC29F3F" box="[349,456,1790,1815]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
|
||
, diameter
|
||
<quantity id="4CB79B0FE025D76FFDCC179309A39F3E" box="[577,681,1790,1815]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="40" pageNumber="41" unit="mm" value="15.5">15.5 mm</quantity>
|
||
, height
|
||
<quantity id="4CB79B0FE025D76FFC8B1792086B9F3E" box="[774,865,1791,1815]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="40" pageNumber="41" unit="mm" value="8.1">8.1 mm</quantity>
|
||
; largest specimen, diameter 21.0 mm, height
|
||
<quantity id="4CB79B0FE025D76FFAE117920BCC9F12" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="40" pageNumber="41" unit="mm" value="11.4">11.4 mm</quantity>
|
||
; H:D of adults 0.46–0.60 [N=34].
|
||
</paragraph>
|
||
<caption id="DF306662E024D76EFF1A15E20BF69D75" httpUri="https://zenodo.org/record/266574/files/figure.png" pageId="41" pageNumber="42" targetBox="[155,1431,204,1137]" targetPageId="41">
|
||
<paragraph id="8BF036EAE024D76EFF1A15E20BF69D75" blockId="41.[151,1436,1167,1373]" pageId="41" pageNumber="42">
|
||
<emphasis id="B93BEAF8E024D76EFF1A15E20A159C8D" bold="true" box="[151,287,1167,1189]" pageId="41" pageNumber="42">FIGURE 30.</emphasis>
|
||
<taxonomicName id="4C4F4D69E024D76EFEA815FD0AA09C8D" box="[293,426,1168,1189]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E024D76EFEA815FD0AA09C8D" box="[293,426,1168,1189]" italics="true" pageId="41" pageNumber="42">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93BEAF8E024D76EFE3F15E2094A9C8D" bold="true" box="[434,576,1167,1189]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="A2085783E024D76EFE3F15E209089C8D" box="[434,514,1167,1189]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A–C,
|
||
</emphasis>
|
||
holotype, Monk’s Cowl Forest, KwaZulu-Natal, diameter 15.5 mm (NMSA W5599/ T3906).
|
||
<emphasis id="B93BEAF8E024D76EFF6215C00A289CEB" bold="true" box="[239,290,1197,1219]" pageId="41" pageNumber="42">D–J,</emphasis>
|
||
additional specimens from throughout the range of the species [
|
||
<emphasis id="B93BEAF8E024D76EFC3515C008DB9CEB" bold="true" box="[952,977,1197,1219]" pageId="41" pageNumber="42">D,</emphasis>
|
||
Majuba, KwaZulu-Natal, diameter 14.5 mm (paratype, NMSA V5932/T3909).
|
||
<emphasis id="B93BEAF8E024D76EFE7115A1091E9CCA" bold="true" box="[508,532,1228,1250]" pageId="41" pageNumber="42">E,</emphasis>
|
||
Pongola Bush, KwaZulu-Natal, diameter 17.0 mm (paratype, NMSA V776/T3915).
|
||
<emphasis id="B93BEAF8E024D76EFA0515A10E939CCA" bold="true" box="[1416,1433,1228,1250]" pageId="41" pageNumber="42">F,</emphasis>
|
||
Phinda Game Reserve, KwaZulu-Natal, diameter 16.2 mm (W1334).
|
||
<emphasis id="B93BEAF8E024D76EFCFC1586088F9D29" bold="true" box="[881,901,1259,1281]" pageId="41" pageNumber="42">G,</emphasis>
|
||
Balloch, Barkly East, E. Cape, diameter 16.8 mm (paratype, NMSA W4836/T3914).
|
||
<emphasis id="B93BEAF8E024D76EFD871464092E9D37" bold="true" box="[522,548,1289,1311]" pageId="41" pageNumber="42">H,</emphasis>
|
||
Mapaya Hill, Zastron, Free State, diameter 13.2 mm (paratype, W6573/T3907).
|
||
<emphasis id="B93BEAF8E024D76EFA0114640E919D37" bold="true" box="[1420,1435,1289,1311]" pageId="41" pageNumber="42">I,</emphasis>
|
||
Mariepskop, Mpumalanga, diameter 17.1 mm (W9364).
|
||
<emphasis id="B93BEAF8E024D76EFD71144508049D16" bold="true" box="[764,782,1320,1342]" pageId="41" pageNumber="42">J,</emphasis>
|
||
Sand River area, Kruger National Park, diameter 15.6 mm (A9759)].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036EAE024D76EFF4A14E90BD99E5D" blockId="41.[151,1436,1412,1941]" pageId="41" pageNumber="42">
|
||
<emphasis id="B93BEAF8E024D76EFF4A14E909159DB5" bold="true" box="[199,543,1412,1437]" pageId="41" pageNumber="42">
|
||
Living animal (
|
||
<figureCitation id="13742A6FE024D76EFE0F14E90AC79DB5" box="[386,461,1412,1437]" captionStart="FIGURE 4" captionStartId="9.[151,250,1791,1813]" captionTargetBox="[184,1395,199,1756]" captionTargetId="figure@9.[184,1396,193,1762]" captionTargetPageId="9" captionText="FIGURE 4. Living animals of Nata and Natella. A, Nata aequiplicata sp. nov., holotype, Stinkhoutkloof, E. Cape, shell diameter 6.5 mm (NMSA W 9248 / T 3066). B, Nata dumeticola, West Coast National Park, W. Cape, shell diameter 10.8 mm (NMSA W 3224). C, Nata erugata sp. nov., Tullach-Mohr, Nelspruit area, Mpumalanga, shell diameter 20 mm (MV ex NMSA W 3301). D, Nata tarachodes, Grootbos Private Nat. Res., Gansbaai, W. Cape, shell diameter 7.7 mm (NMSA W 5658). E, Nata watsoni sp. nov., Hankey, E. Cape, shell diameter 7.5 mm (paratype NMSA W 6524 / T 3921). F, Nata watsoni sp. nov. Camdeboo National Park, Graaff-Reinet, E. Cape, shell diameter 13.0 mm (paratype, NMSA W 6091 / T 2303). G, Nata vernicosa, Vernon Crookes Nat. Res., KwaZulu-Natal, shell diameter 14.0 mm (NMSA W 8020). H, Natella viridescens, Ithala Game Res., KwaZulu-Natal, shell diameter 3.7 mm (NMSA W 7263)." httpUri="https://zenodo.org/record/266548/files/figure.png" pageId="41" pageNumber="42">Figs 4</figureCitation>
|
||
C, 31):
|
||
</emphasis>
|
||
Body generally pale; head and neck greyish-white to cream, dorsal region of neck sometimes pale grey or pale bluish-grey; optic tentacles a slightly darker shade of grey; foot paler beneath shell; posterior portion of foot relatively short; some individuals with microscopic orange pigment granules in skin, particularly on hind end of foot; skin texture moderately well defined; mantle edge pale, also with minute orange granules in some individuals; lining of mantle cavity with little or no dark pigmentation, except for a diffuse blackish mark just inside pneumostome, visible through shell; kidney trigonal, pale buff, not conspicuous through shell.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE024D76EFF4A17ED0AFC9F2C" blockId="41.[151,1436,1412,1941]" pageId="41" pageNumber="42">
|
||
<emphasis id="B93BEAF8E024D76EFF4A17ED0AC79EB1" bold="true" box="[199,461,1664,1689]" pageId="41" pageNumber="42">
|
||
Radula (
|
||
<figureCitation id="13742A6FE024D76EFEA217ED0A719EB1" box="[303,379,1664,1689]" captionStart="FIGURE 6" captionStartId="12.[151,250,1492,1514]" captionTargetBox="[154,1432,339,1464]" captionTargetId="figure@12.[151,1436,338,1471]" captionTargetPageId="12" captionText="FIGURE 6. Radulae of Nata and Natella, automontage light micrographs of half-row portions of stained radula mounts. A, Nata aequiplicata sp. nov., type locality (paratype, NMSA W 9249 / T 3067). B, Nata dumeticola, Jacobsbaai, W. Cape (NMSA V 7760). C, Nata erugata sp. nov., Golden Gate Highlands National Park, Free State (paratype, NMSA W 4243 / T 3913). D, Nata tarachodes, Oubos, W. Cape (NMSA W 5671). E, Nata watsoni sp. nov., Graaff-Reinet, E. Cape (paratype, NMSA W 6091 / T 2303). F, Natella viridescens, Ithala Game Res., KwaZulu-Natal (NMSA V 2530). Scale bars: A = 100 µm; B = 200 µm; C – E = 250 µm; F = 50 µm." httpUri="https://zenodo.org/record/266550/files/figure.png" pageId="41" pageNumber="42">Figs 6</figureCitation>
|
||
C, 32):
|
||
</emphasis>
|
||
Very similar to that of
|
||
<taxonomicName id="4C4F4D69E024D76EFD5317EC08819EB0" box="[734,907,1665,1688]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E024D76EFD5317EC08819EB0" box="[734,907,1665,1688]" italics="true" pageId="41" pageNumber="42">Nata vernicosa</emphasis>
|
||
</taxonomicName>
|
||
, but usually with fewer lateral teeth. Formula 0+(7–8)+(3–4); length up to 12.0 mm, with 24–33 V-shaped rows of teeth; teeth progressively larger from anterior to posterior of radula; rachidian absent; largest tooth (number 7 or 8) taken to be outermost lateral. Tooth morphology as in
|
||
<taxonomicName id="4C4F4D69E024D76EFEEF17800AFA9F2C" box="[354,496,1773,1796]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E024D76EFEEF17800AFA9F2C" box="[354,496,1773,1796]" italics="true" pageId="41" pageNumber="42">N. vernicosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE024D76EFF4A167D0E2C9FBD" blockId="41.[151,1436,1412,1941]" pageId="41" pageNumber="42">
|
||
<emphasis id="B93BEAF8E024D76EFF4A167D0A8E9F01" bold="true" box="[199,388,1808,1833]" pageId="41" pageNumber="42">Distal genitalia:</emphasis>
|
||
<collectingCountry id="F358767AE024D76EFE06167C0AA09F01" box="[395,426,1809,1833]" name="American Samoa" pageId="41" pageNumber="42">As</collectingCountry>
|
||
in
|
||
<taxonomicName id="4C4F4D69E024D76EFE43167C09739F00" box="[462,633,1809,1832]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E024D76EFE43167C09739F00" box="[462,633,1809,1832]" italics="true" pageId="41" pageNumber="42">Nata vernicosa</emphasis>
|
||
</taxonomicName>
|
||
. Epiphallus enters penis between one-third and two-thirds length of the latter from its base; internally, lining of penis forms a raised rim around epiphallus pore; basal portion of bursa copulatrix duct swollen (particularly so in specimens from Hogsback), tapering to mid region and evenly slender thereafter, but constricted prior to bursa copulatrix which is small, subovate to reniform and thin walled.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF306662E027D76DFF1A13F70E5D9AE7" httpUri="https://zenodo.org/record/266575/files/figure.png" pageId="42" pageNumber="43" targetBox="[152,1435,200,640]" targetPageId="42">
|
||
<paragraph id="8BF036EAE027D76DFF1A13F70E5D9AE7" blockId="42.[151,1436,666,719]" pageId="42" pageNumber="43">
|
||
<emphasis id="B93BEAF8E027D76DFF1A13F70A2B9A98" bold="true" box="[151,289,666,688]" pageId="42" pageNumber="43">FIGURE 31.</emphasis>
|
||
<taxonomicName id="4C4F4D69E027D76DFEA413F60ABB9A98" box="[297,433,667,688]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E027D76DFEA413F60ABB9A98" box="[297,433,667,688]" italics="true" pageId="42" pageNumber="43">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E027D76DFE3413F709019A98" box="[441,523,666,688]" pageId="42" pageNumber="43" rank="species">
|
||
<emphasis id="B93BEAF8E027D76DFE3413F709019A98" bold="true" box="[441,523,666,688]" pageId="42" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, living animals.
|
||
<emphasis id="B93BEAF8E027D76DFD3413F709D89A98" bold="true" box="[697,722,666,688]" pageId="42" pageNumber="43">A,</emphasis>
|
||
Mapaya Hill, Zastron, Free State, shell diameter 13.2 mm (paratype, NMSA W6573/T3907).
|
||
<emphasis id="B93BEAF8E027D76DFE0213D40AAF9AE7" bold="true" box="[399,421,697,719]" pageId="42" pageNumber="43">B,</emphasis>
|
||
Tullach-Mohr, Nelspruit area, Mpumalanga, shell diameter 20 mm (MV, ex NMSA W3301).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF306662E027D76DFF1A164F0FC89F7E" httpUri="https://zenodo.org/record/266576/files/figure.png" pageId="42" pageNumber="43" targetBox="[156,1433,771,1797]" targetPageId="42">
|
||
<paragraph id="8BF036EAE027D76DFF1A164F0FC89F7E" blockId="42.[151,1436,1826,1878]" pageId="42" pageNumber="43">
|
||
<emphasis id="B93BEAF8E027D76DFF1A164F0A2E9F10" bold="true" box="[151,292,1826,1848]" pageId="42" pageNumber="43">FIGURE 32.</emphasis>
|
||
<taxonomicName id="4C4F4D69E027D76DFEA2164E0AB09F10" box="[303,442,1827,1848]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="42" pageNumber="43" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E027D76DFEA2164E0AB09F10" box="[303,442,1827,1848]" italics="true" pageId="42" pageNumber="43">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E027D76DFE49164F09139F10" box="[452,537,1826,1848]" pageId="42" pageNumber="43" rank="species">
|
||
<emphasis id="B93BEAF8E027D76DFE49164F09139F10" bold="true" box="[452,537,1826,1848]" pageId="42" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, radula.
|
||
<emphasis id="B93BEAF8E027D76DFDF1164F099E9F10" bold="true" box="[636,660,1826,1848]" pageId="42" pageNumber="43">A,</emphasis>
|
||
Sudwala, Mpumalanga (NMSA W4473).
|
||
<emphasis id="B93BEAF8E027D76DFBD5164F0F659F10" bold="true" box="[1112,1135,1826,1848]" pageId="42" pageNumber="43">B,</emphasis>
|
||
Hogsback E. Cape (NMSA W6654).
|
||
<emphasis id="B93BEAF8E027D76DFF7B162D0A249F7E" bold="true" box="[246,302,1856,1878]" pageId="42" pageNumber="43">C, D,</emphasis>
|
||
Mapaya Hill, Zastron, Free State (paratype, NMSA W6573/T3907). Scale bars = 250 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3556561E027D76CFF4A16EE088C9985" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="distribution">
|
||
<paragraph id="8BF036EAE027D76CFF4A16EE08049934" blockId="42.[151,1436,1923,2020]" lastBlockId="43.[151,1437,151,1257]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
|
||
<emphasis id="B93BEAF8E027D76DFF4A16EE0ADF9FB4" bold="true" box="[199,469,1923,1948]" pageId="42" pageNumber="43">
|
||
Distribution (
|
||
<figureCitation id="13742A6FE027D76DFEE116EE0ACC9FB4" box="[364,454,1923,1948]" captionStart="FIGURE 29" captionStartId="39.[151,250,1922,1944]" captionTargetBox="[244,1345,1114,1906]" captionTargetId="figure@39.[244,1346,1114,1906]" captionTargetPageId="39" captionText="FIGURE 29. Comparison of distributions of Nata vernicosa (triangles) and Nata erugata sp. nov. (circles). Green symbols indicate localities for specimens used in DNA analyses (Moussalli & Herbert 2016); contour at 1000 m. The isolated population of Nata vernicosa in the Cape Town is considered to be derived from translocated individuals." httpUri="https://zenodo.org/record/266573/files/figure.png" pageId="42" pageNumber="43">Fig. 29</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Endemic to eastern
|
||
<collectingCountry id="F358767AE027D76DFD4416EE08579FB4" box="[713,861,1923,1948]" name="South Africa" pageId="42" pageNumber="43">South Africa</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="F358767AE027D76DFB8A16EE0F949FB4" box="[1031,1182,1923,1948]" name="Mozambique" pageId="42" pageNumber="43">Mozambique</collectingCountry>
|
||
, but relatively widely distributed within this region, ranging from Hogsback in the south west, through western
|
||
<collectingCountry id="F358767AE027D76DFB2B16CA0E099FE8" box="[1190,1283,1959,1984]" name="Lesotho" pageId="42" pageNumber="43">Lesotho</collectingCountry>
|
||
, eastern Free State and Gauteng, into western and northern KwaZulu-Natal, the Lowveld of Mpumalanga and southern
|
||
<collectingCountry id="F358767AE026D76CFF1A11FA0A259898" box="[151,303,151,176]" name="Mozambique" pageId="43" pageNumber="44">Mozambique</collectingCountry>
|
||
. Throughout the southern and western parts of its range the species is largely restricted to altitudes over
|
||
<quantity id="4CB79B0FE026D76CFF5F11D10A2098FC" box="[210,298,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="m" value="1000.0">1000 m</quantity>
|
||
and extending to
|
||
<quantity id="4CB79B0FE026D76CFE7111D1095E98FC" box="[508,596,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="43" pageNumber="44" unit="m" value="2200.0">2200 m</quantity>
|
||
along the KwaZulu-Natal escarpment edge. However, the populations in north-eastern KwaZulu-Natal (Zululand), the Mpumalanga Lowveld (southern Kruger National Park) and
|
||
<collectingCountry id="F358767AE026D76CFF1A10690A219935" box="[151,299,260,285]" name="Mozambique" pageId="43" pageNumber="44">Mozambique</collectingCountry>
|
||
occur at much lower altitudes (
|
||
<quantity id="4CB79B0FE026D76CFD01106908089934" box="[652,770,260,284]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.0" metricValueMin="0.3" pageId="43" pageNumber="44" unit="m" value="165.0" valueMax="300.0" valueMin="30.0">30–300 m</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE026D76CFF4A104A088C9985" blockId="43.[151,1437,151,1257]" pageId="43" pageNumber="44">
|
||
<emphasis id="B93BEAF8E026D76CFF4A104A0A209968" bold="true" box="[199,298,295,320]" pageId="43" pageNumber="44">Habitat.</emphasis>
|
||
At altitudes above
|
||
<quantity id="4CB79B0FE026D76CFD9F104509619917" box="[530,619,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="m" value="1000.0">1000 m</quantity>
|
||
the species occurs primarily in afrotemperate forest and more thickly vegetated habitats such as
|
||
<taxonomicName id="4C4F4D69E026D76CFE3210210934994D" box="[447,574,332,357]" class="Magnoliopsida" family="Rosaceae" genus="Leucosidea" kingdom="Plantae" order="Rosales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93BEAF8E026D76CFE3210210934994D" box="[447,574,332,357]" italics="true" pageId="43" pageNumber="44">Leucosidea</emphasis>
|
||
</taxonomicName>
|
||
scrub. However, specimens have also been found in sheltering microhabitats in more open vegetation, even montane grassland. At lower altitudes in Zululand and the Kruger National Park it occurs in a variety of vegetation
|
||
<typeStatus id="54F48848E026D76CFD8B10F8094B9985" box="[518,577,405,429]" pageId="43" pageNumber="44">types</typeStatus>
|
||
within the Savannah Biome.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E026D76CFF4A10DA0FFA9CC0" pageId="43" pageNumber="44" type="discussion">
|
||
<paragraph id="8BF036EAE026D76CFF4A10DA093A9B10" blockId="43.[151,1437,151,1257]" pageId="43" pageNumber="44">
|
||
<emphasis id="B93BEAF8E026D76CFF4A10DA0A3199F8" bold="true" box="[199,315,439,464]" pageId="43" pageNumber="44">Remarks.</emphasis>
|
||
When discussing Pfeiffer’s ‘
|
||
<taxonomicName id="4C4F4D69E026D76CFDF010D509FF99F8" authority="Connolly" authorityName="Connolly" authorityYear="1939" box="[637,757,440,464]" pageId="43" pageNumber="44" rank="variety" variety="minor">
|
||
var.
|
||
<emphasis id="B93BEAF8E026D76CFD3F10D409FF99F8" box="[690,757,441,464]" italics="true" pageId="43" pageNumber="44">minor</emphasis>
|
||
</taxonomicName>
|
||
’,
|
||
<bibRefCitation id="EFDE4B1BE026D76CFC8310DA08C299F8" author="Connolly" box="[782,968,439,464]" pageId="43" pageNumber="57" refString="Connolly, M. (1939) A monographic survey of South African non-marine Mollusca. Annals of the South African Museum, 33, 1 - 660." type="journal article" year="1939">Connolly (1939)</bibRefCitation>
|
||
noted that although Pfeiffer’s description ‘
|
||
<emphasis id="B93BEAF8E026D76CFF2D10B1091799DD" box="[160,541,476,501]" italics="true" pageId="43" pageNumber="44">costulis superficiei obsoletioribus</emphasis>
|
||
’ (Pfeiffer 1853: 95) may apply to an immature example of
|
||
<taxonomicName id="4C4F4D69E026D76CFB4B10B00E9F99DD" box="[1222,1429,476,501]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="liparoxantha">
|
||
<emphasis id="B93BEAF8E026D76CFB4B10B00E9F99DD" box="[1222,1429,476,501]" italics="true" pageId="43" pageNumber="44">Nata liparoxantha</emphasis>
|
||
</taxonomicName>
|
||
, ‘…there extends from the Cape Province to Delagoa a hitherto undetermined race, which may represent his variety. It differs from
|
||
<taxonomicName id="4C4F4D69E026D76CFEC413480ABC9A14" box="[329,438,549,572]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E026D76CFEC413480ABC9A14" box="[329,438,549,572]" italics="true" pageId="43" pageNumber="44">vernicosa</emphasis>
|
||
</taxonomicName>
|
||
solely in being very much smoother, the sculpture consisting of fine growth striolae instead of costulae’ (
|
||
<bibRefCitation id="EFDE4B1BE026D76CFE04132A097E9A48" author="Connolly" box="[393,628,583,608]" pageId="43" pageNumber="57" refString="Connolly, M. (1939) A monographic survey of South African non-marine Mollusca. Annals of the South African Museum, 33, 1 - 660." type="journal article" year="1939">Connolly 1939: 100</bibRefCitation>
|
||
). We have likewise been able to discern such a ‘race’, but we do not consider that it can be considered conspecific with Pfeiffer’s
|
||
<taxonomicName id="4C4F4D69E026D76CFCC7130008B79AAC" box="[842,957,621,644]" pageId="43" pageNumber="44" rank="variety" variety="minor">
|
||
var.
|
||
<emphasis id="B93BEAF8E026D76CFCF6130008B79AAC" box="[891,957,621,644]" italics="true" pageId="43" pageNumber="44">minor</emphasis>
|
||
</taxonomicName>
|
||
. Examination of the
|
||
<typeStatus id="54F48848E026D76CFB2613010E189AAD" box="[1195,1298,620,645]" pageId="43" pageNumber="44" type="lectotype">lectotype</typeStatus>
|
||
of the latter at the SMNH (see above regarding its authenticity) reveals that although the axial sculpture is weak, distinct and regularly spaced axial riblets are nonetheless present on the adapical surface and the specimen is simply a weakly sculptured example of the variable
|
||
<taxonomicName id="4C4F4D69E026D76CFDB713B409C79AD8" box="[570,717,729,752]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E026D76CFDB713B409C79AD8" box="[570,717,729,752]" italics="true" pageId="43" pageNumber="44">N. vernicosa</emphasis>
|
||
</taxonomicName>
|
||
. Since all the other names currently considered to be junior synonyms of
|
||
<taxonomicName id="4C4F4D69E026D76CFEA013900AB19B3C" box="[301,443,765,788]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E026D76CFEA013900AB19B3C" box="[301,443,765,788]" italics="true" pageId="43" pageNumber="44">N. vernicosa</emphasis>
|
||
</taxonomicName>
|
||
likewise relate to material with distinct axial riblets, we are therefore obliged to propose a new name for the present material.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE026D76CFF4A122909129BC4" blockId="43.[151,1437,151,1257]" pageId="43" pageNumber="44">
|
||
Throughout much of its distribution
|
||
<taxonomicName id="4C4F4D69E026D76CFDEB122809F69B74" box="[614,764,837,860]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E026D76CFDEB122809F69B74" box="[614,764,837,860]" italics="true" pageId="43" pageNumber="44">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E026D76CFC89122908559B75" box="[772,863,836,861]" pageId="43" pageNumber="44" rank="species">
|
||
<emphasis id="B93BEAF8E026D76CFC89122908559B75" bold="true" box="[772,863,836,861]" pageId="43" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the only species of
|
||
<taxonomicName id="4C4F4D69E026D76CFBED12280F9D9B74" box="[1120,1175,837,860]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93BEAF8E026D76CFBED12280F9D9B74" box="[1120,1175,837,860]" italics="true" pageId="43" pageNumber="44">Nata</emphasis>
|
||
</taxonomicName>
|
||
to be found, but at the lower limits of its altitudinal range, particularly in E. Cape and KwaZulu-Natal, there is some geographical overlap with
|
||
<taxonomicName id="4C4F4D69E026D76CFF5D12E00A709B8C" box="[208,378,909,932]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E026D76CFF5D12E00A709B8C" box="[208,378,909,932]" italics="true" pageId="43" pageNumber="44">Nata vernicosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13742A6FE026D76CFE0412E10AD49B8C" box="[393,478,908,933]" captionStart="FIGURE 29" captionStartId="39.[151,250,1922,1944]" captionTargetBox="[244,1345,1114,1906]" captionTargetId="figure@39.[244,1346,1114,1906]" captionTargetPageId="39" captionText="FIGURE 29. Comparison of distributions of Nata vernicosa (triangles) and Nata erugata sp. nov. (circles). Green symbols indicate localities for specimens used in DNA analyses (Moussalli & Herbert 2016); contour at 1000 m. The isolated population of Nata vernicosa in the Cape Town is considered to be derived from translocated individuals." httpUri="https://zenodo.org/record/266573/files/figure.png" pageId="43" pageNumber="44">Fig. 29</figureCitation>
|
||
). However, further studies are needed to establish whether and to what extent the two species co-occur. There is certainly evidence that this may be the case at some localities (Hogsback, Umthatha area, Karkloof and Pilgrim’s Rest area).
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE026D76CFF4A129A0E299C88" blockId="43.[151,1437,151,1257]" pageId="43" pageNumber="44">
|
||
The occurrence of populations of what is largely a higher altitude species in the hotter more low-lying regions of northern KwaZulu-Natal, the Kruger National Park and southern
|
||
<collectingCountry id="F358767AE026D76CFC2315710F489C1D" box="[942,1090,1052,1077]" name="Mozambique" pageId="43" pageNumber="44">Mozambique</collectingCountry>
|
||
(
|
||
<figureCitation id="13742A6FE026D76CFBD815710FA19C1C" box="[1109,1195,1052,1077]" captionStart="FIGURE 29" captionStartId="39.[151,250,1922,1944]" captionTargetBox="[244,1345,1114,1906]" captionTargetId="figure@39.[244,1346,1114,1906]" captionTargetPageId="39" captionText="FIGURE 29. Comparison of distributions of Nata vernicosa (triangles) and Nata erugata sp. nov. (circles). Green symbols indicate localities for specimens used in DNA analyses (Moussalli & Herbert 2016); contour at 1000 m. The isolated population of Nata vernicosa in the Cape Town is considered to be derived from translocated individuals." httpUri="https://zenodo.org/record/266573/files/figure.png" pageId="43" pageNumber="44">Fig. 29</figureCitation>
|
||
), at altitudes of well below
|
||
<quantity id="4CB79B0FE026D76CFF6F152D0A329C7F" box="[226,312,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="m" value="1000.0">1000 m</quantity>
|
||
, is anomalous and hard to explain. It is certainly suggestive of there being a further cryptic lineage in what we group under one species. For this reason, we have not included any specimens from these localities in the
|
||
<typeStatus id="54F48848E026D76CFF1A15E40BCD9C89" box="[151,199,1161,1185]" pageId="43" pageNumber="44">type</typeStatus>
|
||
material for this species. This is an issue needing to be studied in more detail using molecular data.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE026D76CFF4A15C10FFA9CC0" blockId="43.[151,1437,151,1257]" pageId="43" pageNumber="44">
|
||
<emphasis id="B93BEAF8E026D76CFF4A15C10A659CED" bold="true" box="[199,367,1196,1221]" pageId="43" pageNumber="44">Conservation.</emphasis>
|
||
<collectingCountry id="F358767AE026D76CFEF515C10A9D9CEC" box="[376,407,1196,1220]" name="American Samoa" pageId="43" pageNumber="44">As</collectingCountry>
|
||
interpreted here,
|
||
<taxonomicName id="4C4F4D69E026D76CFDE915C009F19CEC" box="[612,763,1197,1220]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E026D76CFDE915C009F19CEC" box="[612,763,1197,1220]" italics="true" pageId="43" pageNumber="44">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E026D76CFC8815C1086B9CED" box="[773,865,1196,1221]" pageId="43" pageNumber="44" rank="species">
|
||
<emphasis id="B93BEAF8E026D76CFC8815C1086B9CED" bold="true" box="[773,865,1196,1221]" pageId="43" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is a widespread and moderately common species occurring in a range of habitat
|
||
<typeStatus id="54F48848E026D76CFE7D15BC09249CC1" box="[496,558,1233,1257]" pageId="43" pageNumber="44">types</typeStatus>
|
||
.
|
||
<collectingCountry id="F358767AE026D76CFDB415BD09529CC0" box="[569,600,1232,1256]" name="American Samoa" pageId="43" pageNumber="44">As</collectingCountry>
|
||
such, its conservation is not at present a matter of concern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |