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<document id="61B9E2F651BA9D1EAC5DCB52BEB38BAA" ID-DOI="10.11646/zootaxa.4093.3.2" ID-GBIF-Dataset="9f88e467-a052-4233-af95-05206184d3d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="266589" ID-ZooBank="F03731C8-DE0E-4CCA-9E11-A2DF5994C9A9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458721627483" checkinUser="plazi" docAuthor="Prieto, Carlos &amp; Vargas, Maria A." docDate="2016" docId="03E687EBFFDF85507CD3FA37FE39F953" docLanguage="en" docName="zootaxa.4093.3.2.pdf.imf" docOrigin="Zootaxa 4093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhamma hybla Druce 1907" docType="treatment" docVersion="7" lastPageNumber="334" masterDocId="FFDFFF93FFD6855B7C44FFC7FFB8FFAC" masterDocTitle="Elfin butterflies of the genus Rhamma Johnson (Lepidoptera: Lycaenidae: Theclinae): A review of the Colombian species" masterLastPageNumber="342" masterPageNumber="323" pageNumber="332" updateTime="1698185951747" updateUser="plazi">
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<mods:title id="DA5C1B1288B3519CE1348E5061AD71F0">Elfin butterflies of the genus Rhamma Johnson (Lepidoptera: Lycaenidae: Theclinae): A review of the Colombian species</mods:title>
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<mods:namePart id="5BEAB9469BD8C22D18A65E4B9B03AE3C">Prieto, Carlos</mods:namePart>
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<mods:namePart id="9498005D3979E457A9FBE9783AC7AFC1">Vargas, Maria A.</mods:namePart>
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<mods:title id="105058EEB007020353BAA253E607884A">Zootaxa</mods:title>
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<mods:date id="E0B779E1299D4FA9781A9AEB70363864">2016</mods:date>
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<mods:number id="CD2A85927A09F3B551F3F390BBA4C62A">4093</mods:number>
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<mods:identifier id="A117DEE2A706936433299DDCD93C851F" type="DOI">10.11646/zootaxa.4093.3.2</mods:identifier>
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<treatment id="03E687EBFFDF85507CD3FA37FE39F953" ID-DOI="http://doi.org/10.5281/zenodo.6058879" ID-GBIF-Taxon="119257860" ID-Zenodo-Dep="6058879" LSID="urn:lsid:plazi:treatment:03E687EBFFDF85507CD3FA37FE39F953" httpUri="http://treatment.plazi.org/id/03E687EBFFDF85507CD3FA37FE39F953" lastPageId="11" lastPageNumber="334" pageId="9" pageNumber="332">
<subSubSection id="C3556576FFDF85527CD3FA37FE6EF987" pageId="9" pageNumber="332" type="nomenclature">
<paragraph id="8BF036FDFFDF85527CD3FA37FDB8F9A6" blockId="9.[151,512,1520,1579]" box="[151,512,1520,1546]" pageId="9" pageNumber="332">
<heading id="D0B88191FFDF85527CD3FA37FDB8F9A6" bold="true" box="[151,512,1520,1546]" fontSize="11" level="1" pageId="9" pageNumber="332" reason="1">
<taxonomicName id="4C4F4D7EFFDF85527CD3FA37FDB8F9A6" authority="Druce, 1907" authorityName="Druce" authorityYear="1907" box="[151,512,1520,1546]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="hybla">
<emphasis id="B93BEAEFFFDF85527CD3FA37FDB8F9A6" bold="true" box="[151,512,1520,1546]" pageId="9" pageNumber="332">
<emphasis id="B93BEAEFFFDF85527CD3FA37FEE8F9A6" bold="true" box="[151,336,1520,1546]" italics="true" pageId="9" pageNumber="332">Rhamma hybla</emphasis>
(Druce, 1907)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF036FDFFDF85527CD3F9D5FE6EF987" blockId="9.[151,512,1520,1579]" box="[151,470,1554,1579]" pageId="9" pageNumber="332">
(
<figureCitation id="13742A78FFDF85527CDAF9D5FE9FF987" box="[158,295,1554,1579]" captionStart="FIGURES 1 – 10" captionStartId="5.[151,264,1553,1576]" captionTargetBox="[154,1436,193,1527]" captionTargetId="figure@5.[151,1436,193,1533]" captionTargetPageId="5" captionText="FIGURES 1 – 10. Adults depicted in dorsal (left) and ventral (right) aspect: 1. Rhamma hybla male (Antioquia); 2. Rhamma hybla female (Antioquia); 3. Rhamma adunca male (Cauca); 4. Rhamma adunca female (Cauca); 5. Rhamma adunca male (Belmira, Antioquia); 6. Rhamma adunca female (Cauca); 7. Rhamma arria male (Valle); 8. Rhamma arria female (Valle); 9. Rhamma livida male (Cauca); 10. Rhamma livida female (Cauca)." httpUri="https://zenodo.org/record/266590/files/figure.png" pageId="9" pageNumber="332">Figures 1, 2</figureCitation>
,
<figureCitation id="13742A78FFDF85527D77F9D4FEE9F987" box="[307,337,1555,1579]" captionStart="FIGURES 30 – 35" captionStartId="9.[151,264,1361,1384]" captionTargetBox="[155,1431,193,1323]" captionTargetId="figure@9.[151,1436,193,1324]" captionTargetPageId="9" captionText="FIGURES 30 – 35. Male genitalia armatures in lateral (left) and ventral (right) views: 30. Rhamma hybla; 31. Rhamma adunca; 32. Rhamma arria; 33. Rhamma livida (arrows point a., vinculum strut strongly developed and b., vinculum forming a 90 degree angle); 34. Rhamma oxida; 35. Rhamma comstocki." httpUri="https://zenodo.org/record/266593/files/figure.png" pageId="9" pageNumber="332">30</figureCitation>
,
<figureCitation id="13742A78FFDF85527D19F9D4FEC3F987" box="[349,379,1555,1579]" captionStart="FIGURES 42 – 50" captionStartId="10.[151,264,1892,1915]" captionTargetBox="[379,1198,193,1851]" captionTargetId="figure@10.[379,1198,193,1851]" captionTargetPageId="10" captionText="FIGURES 42 – 50. Female genitalia (ductus bursae and bursa copulatrix): 42. Rhamma hybla (arrows point a., strongly developed lamella postvaginalis, terminating in serrate configurations b., ductus bursae lightly sclerotized in the middle so that it appears to be transparent and c., dendritic, fan shaped signa) 43. Rhamma adunca; 44. Rhamma arria; 45. Rhamma livida; 46. Rhamma oxida; 47. Rhamma comstocki; 48. Rhamma shapiroi; 49. Rhamma bilix; 50. Rhamma anosma." httpUri="https://zenodo.org/record/266594/files/figure.png" pageId="9" pageNumber="332">42</figureCitation>
,
<figureCitation id="13742A78FFDF85527DC3F9D5FE1DF987" box="[391,421,1554,1579]" captionStart="FIGURES 51 – 62" captionStartId="11.[151,264,1499,1522]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@11.[151,1436,193,1478]" captionTargetPageId="11" captionText="FIGURES 51 – 62. Androconial brands (arrows point the scent pad): 51. Rhamma hybla; 52. Rhamma adunca; 53. Rhamma arria; 54. Rhamma livida; 55. Rhamma oxida; 56. Rhamma comstocki; 57. Rhamma shapiroi; 58. Rhamma mishma; 59. Rhamma bilix; 60. Rhamma anosma; 61. Rhamma familiaris; 62. Rhamma commodus." httpUri="https://zenodo.org/record/266595/files/figure.png" pageId="9" pageNumber="332">51</figureCitation>
,
<figureCitation id="13742A78FFDF85527DF5F9D5FE76F987" box="[433,462,1554,1579]" captionStart="FIGURES 67 – 70" captionStartId="15.[151,264,1912,1935]" captionTargetBox="[159,1431,198,1884]" captionTargetId="figure@15.[151,1436,193,1892]" captionTargetPageId="15" captionText="FIGURES 67 – 70. Distributional maps of Colombian Rhamma. 67. Rhamma commodus, Rhamma familiaris; 68. Rhamma comstocki; 69. Rhamma hybla; 70. Rhamma livida." httpUri="https://zenodo.org/record/266597/files/figure.png" pageId="9" pageNumber="332">69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3556576FFDF85527CD3F99CFAD0F923" pageId="9" pageNumber="332" type="reference_group">
<paragraph id="8BF036FDFFDF85527CD3F99CFBE4F9DD" blockId="9.[151,1384,1627,1679]" box="[151,1116,1627,1649]" pageId="9" pageNumber="332">
<taxonomicName id="4C4F4D7EFFDF85527CD3F99CFE6BF9DD" ID-CoL="566H2" authority="Druce, 1907: 578" authorityName="Druce" authorityPageNumber="578" authorityYear="1907" box="[151,467,1627,1649]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="hybla">
<emphasis id="B93BEAEFFFDF85527CD3F99CFEA2F9DD" box="[151,282,1627,1649]" italics="true" pageId="9" pageNumber="332">Thecla hybla</emphasis>
Druce, 1907: 578
</taxonomicName>
, pl. 33, fig. 4; Johnson 1992: 142, fig. 153.
<typeStatus id="54F4885FFFDF85527FCAF99CFC54F9DD" box="[910,1004,1627,1649]" pageId="9" pageNumber="332" type="holotype">Holotype</typeStatus>
examined.
</paragraph>
<paragraph id="8BF036FDFFDF85527CD3F9BEFAD0F923" blockId="9.[151,1384,1627,1679]" box="[151,1384,1657,1679]" pageId="9" pageNumber="332">
<taxonomicName id="4C4F4D7EFFDF85527CD3F9BEFD31F923" authority="Salazar &amp; Johnson, 1997: 3" authorityName="Salazar &amp; Johnson" authorityPageNumber="3" authorityYear="1997" box="[151,649,1657,1679]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="emeraldina">
<emphasis id="B93BEAEFFFDF85527CD3F9BEFED3F923" box="[151,363,1657,1679]" italics="true" pageId="9" pageNumber="332">Rhamma emeraldina</emphasis>
Salazar &amp; Johnson, 1997: 3
</taxonomicName>
, photoplate IX, D, deposited in FSCA. Image of the
<typeStatus id="54F4885FFFDF855278DBF9BEFB40F923" box="[1183,1272,1657,1679]" pageId="9" pageNumber="332" type="holotype">holotype</typeStatus>
examined.
</paragraph>
</subSubSection>
<subSubSection id="C3556576FFDF85527CD3F97BFAD6F825" pageId="9" pageNumber="332" type="materials_examined">
<paragraph id="8BF036FDFFDF85527CD3F97BFBB5F954" blockId="9.[151,1436,1724,2001]" pageId="9" pageNumber="332">
<emphasis id="B93BEAEFFFDF85527CD3F97BFEFEF979" bold="true" box="[151,326,1724,1749]" pageId="9" pageNumber="332">
<typeStatus id="54F4885FFFDF85527CD3F97BFF6AF979" box="[151,210,1724,1749]" pageId="9" pageNumber="332">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F4885FFFDF85527D09F97BFE0DF979" box="[333,437,1724,1749]" pageId="9" pageNumber="332" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F358766DFFDF85527E47F97BFDDAF979" box="[515,610,1724,1749]" name="Ecuador" pageId="9" pageNumber="332">Ecuador</collectingCountry>
, Hewitson Coll. 79-60,
<taxonomicName id="4C4F4D7EFFDF85527F35F97BFB1FF979" authority="B.M. Type No. Rh." authorityName="B.M. Type No. Rh." box="[881,1191,1724,1749]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="genus">
Thecla, B.M.
<typeStatus id="54F4885FFFDF8552784AF97AFBFDF979" box="[1038,1093,1725,1749]" pageId="9" pageNumber="332">Type</typeStatus>
No. Rh.
</taxonomicName>
603., Genitalia by K. Johnson
<date id="FFF1103DFFDF85527CBEF926FEE1F954" box="[250,345,1760,1784]" pageId="9" pageNumber="332" value="1983-09">IX 1983</date>
,
<taxonomicName id="4C4F4D7EFFDF85527D20F927FE13F955" box="[356,427,1760,1785]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="hybla">Hybla</taxonomicName>
, male symbol,
<typeStatus id="54F4885FFFDF85527E11F926FD3DF955" box="[597,645,1761,1785]" pageId="9" pageNumber="332">type</typeStatus>
H. H Druce. Deposited in BMNH.
</paragraph>
<paragraph id="8BF036FDFFDF85527C83F8C3FAD6F825" blockId="9.[151,1436,1724,2001]" pageId="9" pageNumber="332">
<emphasis id="B93BEAEFFFDF85527C83F8C3FDE8F8B1" bold="true" box="[199,592,1796,1821]" pageId="9" pageNumber="332">Taxonomic history and remarks.</emphasis>
We were able to examine topotypical specimens of
<taxonomicName id="4C4F4D7EFFDF855278E5F8C2FEE6F8EC" authority="Salazar &amp; Johnson, 1997" authorityName="Salazar &amp; Johnson" authorityYear="1997" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="emeraldina">
<emphasis id="B93BEAEFFFDF855278E5F8C2FAFAF8B1" box="[1185,1346,1796,1821]" italics="true" pageId="9" pageNumber="332">R. emeraldina</emphasis>
Salazar &amp; Johnson, 1997
</taxonomicName>
, and all diagnostic features discussed in the original description of this species fall within the range of variation of
<taxonomicName id="4C4F4D7EFFDF85527DC0F88AFE5AF8C9" box="[388,482,1868,1893]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="hybla">
<emphasis id="B93BEAEFFFDF85527DC0F88AFE5AF8C9" box="[388,482,1868,1893]" italics="true" pageId="9" pageNumber="332">R. hybla</emphasis>
</taxonomicName>
. In Johnson
<emphasis id="B93BEAEFFFDF85527E35F88AFD12F8C9" box="[625,682,1868,1893]" italics="true" pageId="9" pageNumber="332">et al.</emphasis>
(1997), the authors illustrate a specimen from
<collectingCountry id="F358766DFFDF855278FCF88BFAB3F8C9" box="[1208,1291,1868,1893]" name="Bolivia" pageId="9" pageNumber="332">Bolivia</collectingCountry>
identified as
<taxonomicName id="4C4F4D7EFFDF85527CD3F8B6FEA8F825" box="[151,272,1904,1929]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="cassidyi">
<emphasis id="B93BEAEFFFDF85527CD3F8B6FEA8F825" box="[151,272,1904,1929]" italics="true" pageId="9" pageNumber="332">R. cassidyi</emphasis>
</taxonomicName>
(IX, E), however, based in our current knowledge of the group, this specimen is a male of
<emphasis id="B93BEAEFFFDF8552794DF8B6FAD6F825" box="[1289,1390,1904,1929]" italics="true" pageId="9" pageNumber="332">
<taxonomicName id="4C4F4D7EFFDF8552794DF8B6FAD2F825" box="[1289,1386,1904,1929]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="332" phylum="Arthropoda" rank="species" species="hybla">R. hybla</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3556576FFDF85527C83F853FA94F87D" pageId="9" pageNumber="332" type="distribution">
<paragraph id="8BF036FDFFDF85527C83F853FA94F87D" blockId="9.[151,1436,1724,2001]" pageId="9" pageNumber="332">
<emphasis id="B93BEAEFFFDF85527C83F853FDAFF801" bold="true" box="[199,535,1940,1965]" pageId="9" pageNumber="332">Distribution and variability.</emphasis>
This is the largest species of the genus. We have found it in several localities of the central cordillera in Antioquia where it is relatively abundant and with little morphological variation.
</paragraph>
</subSubSection>
<caption id="DF306675FFDC85517CD3F8A3FB00F87B" httpUri="https://zenodo.org/record/266594/files/figure.png" pageId="10" pageNumber="333" targetBox="[379,1198,193,1851]" targetPageId="10">
<paragraph id="8BF036FDFFDC85517CD3F8A3FB00F87B" blockId="10.[151,1436,1892,2007]" pageId="10" pageNumber="333">
<emphasis id="B93BEAEFFFDC85517CD3F8A3FEEAF8D6" bold="true" box="[151,338,1892,1915]" pageId="10" pageNumber="333">FIGURES 42–50</emphasis>
. Female genitalia (ductus bursae and bursa copulatrix):
<emphasis id="B93BEAEFFFDC85517FF2F8A2FC6EF8D6" bold="true" box="[950,982,1893,1914]" pageId="10" pageNumber="333">42.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517FA5F8A2FBC7F8D7" box="[993,1151,1893,1915]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="hybla">
<emphasis id="B93BEAEFFFDC85517FA5F8A2FBC7F8D7" box="[993,1151,1893,1915]" italics="true" pageId="10" pageNumber="333">Rhamma hybla</emphasis>
</taxonomicName>
(arrows point
<emphasis id="B93BEAEFFFDC85517960F8A2FA8FF8D6" bold="true" box="[1316,1335,1893,1914]" pageId="10" pageNumber="333">a.</emphasis>
, strongly developed lamella postvaginalis, terminating in serrate configurations
<emphasis id="B93BEAEFFFDC85517F1BF843FCCBF835" bold="true" box="[863,883,1924,1945]" pageId="10" pageNumber="333">b.</emphasis>
, ductus bursae lightly sclerotized in the middle so that it appears to be transparent and
<emphasis id="B93BEAEFFFDC85517D91F865FE5FF81B" bold="true" box="[469,487,1954,1975]" pageId="10" pageNumber="333">c.</emphasis>
, dendritic, fan shaped signa)
<emphasis id="B93BEAEFFFDC85517F4AF865FC96F81B" bold="true" box="[782,814,1954,1975]" pageId="10" pageNumber="333">43.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517F70F865FC67F814" box="[820,991,1954,1976]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="adunca">
<emphasis id="B93BEAEFFFDC85517F70F865FC67F814" box="[820,991,1954,1976]" italics="true" pageId="10" pageNumber="333">Rhamma adunca</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517FAFF865FBB3F81B" bold="true" box="[1003,1035,1954,1975]" pageId="10" pageNumber="333">44.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517854F865FB1CF814" box="[1040,1188,1954,1976]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="arria">
<emphasis id="B93BEAEFFFDC85517854F865FB1CF814" box="[1040,1188,1954,1976]" italics="true" pageId="10" pageNumber="333">Rhamma arria</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC855178F5F865FB69F81B" bold="true" box="[1201,1233,1954,1975]" pageId="10" pageNumber="333">45.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517892F865FAC8F814" box="[1238,1392,1954,1976]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="livida">
<emphasis id="B93BEAEFFFDC85517892F865FAC8F814" box="[1238,1392,1954,1976]" italics="true" pageId="10" pageNumber="333">Rhamma livida</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517938F865FA24F81B" bold="true" box="[1404,1436,1954,1975]" pageId="10" pageNumber="333">46.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517CD3F806FE97F87B" box="[151,303,1985,2007]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="oxida">
<emphasis id="B93BEAEFFFDC85517CD3F806FE97F87B" box="[151,303,1985,2007]" italics="true" pageId="10" pageNumber="333">Rhamma oxida</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517D79F806FEE4F87A" bold="true" box="[317,348,1985,2006]" pageId="10" pageNumber="333">47.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517D27F806FD90F87B" box="[355,552,1985,2007]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="comstocki">
<emphasis id="B93BEAEFFFDC85517D27F806FD90F87B" box="[355,552,1985,2007]" italics="true" pageId="10" pageNumber="333">Rhamma comstocki</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517E71F806FDEDF87A" bold="true" box="[565,597,1985,2006]" pageId="10" pageNumber="333">48.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517E18F806FCA8F87B" box="[604,784,1985,2007]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="shapiroi">
<emphasis id="B93BEAEFFFDC85517E18F806FCA8F87B" box="[604,784,1985,2007]" italics="true" pageId="10" pageNumber="333">Rhamma shapiroi</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517F58F806FC85F87A" bold="true" box="[796,829,1985,2006]" pageId="10" pageNumber="333">49.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517F07F806FC68F87B" box="[835,976,1985,2007]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="bilix">
<emphasis id="B93BEAEFFFDC85517F07F806FC68F87B" box="[835,976,1985,2007]" italics="true" pageId="10" pageNumber="333">Rhamma bilix</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDC85517F99F806FC45F87A" bold="true" box="[989,1021,1985,2006]" pageId="10" pageNumber="333">50.</emphasis>
<taxonomicName id="4C4F4D7EFFDC85517847F806FB0BF87B" box="[1027,1203,1985,2007]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="333" phylum="Arthropoda" rank="species" species="anosma">
<emphasis id="B93BEAEFFFDC85517847F806FB0BF87B" box="[1027,1203,1985,2007]" italics="true" pageId="10" pageNumber="333">Rhamma anosma</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF306675FFDD85507CD3FA1CFBBAF983" httpUri="https://zenodo.org/record/266595/files/figure.png" pageId="11" pageNumber="334" targetBox="[151,1436,193,1477]" targetPageId="11">
<paragraph id="8BF036FDFFDD85507CD3FA1CFBBAF983" blockId="11.[151,1436,1499,1583]" pageId="11" pageNumber="334">
<emphasis id="B93BEAEFFFDD85507CD3FA1CFEF7FA5D" bold="true" box="[151,335,1499,1522]" pageId="11" pageNumber="334">FIGURES 51–62</emphasis>
. Androconial brands (arrows point the scent pad):
<emphasis id="B93BEAEFFFDD85507F27FA1BFC3BFA5D" bold="true" box="[867,899,1500,1521]" pageId="11" pageNumber="334">51.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507FCFFA1BFB9EFA5E" box="[907,1062,1500,1522]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="hybla">
<emphasis id="B93BEAEFFFDD85507FCFFA1BFB9EFA5E" box="[907,1062,1500,1522]" italics="true" pageId="11" pageNumber="334">Rhamma hybla</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507871FA1BFBEDFA5D" bold="true" box="[1077,1109,1500,1521]" pageId="11" pageNumber="334">52.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507819FA1BFAB3FA5E" box="[1117,1291,1500,1522]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="adunca">
<emphasis id="B93BEAEFFFDD85507819FA1BFAB3FA5E" box="[1117,1291,1500,1522]" italics="true" pageId="11" pageNumber="334">Rhamma adunca</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD8550795EFA1BFA82FA5D" bold="true" box="[1306,1338,1500,1521]" pageId="11" pageNumber="334">53.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507906FA1BFF73F9BD" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="arria">
<emphasis id="B93BEAEFFFDD85507906FA1BFF73F9BD" italics="true" pageId="11" pageNumber="334">Rhamma arria</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507C99FA3CFF44F9BC" bold="true" box="[221,252,1531,1552]" pageId="11" pageNumber="334">54.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507D43FA3CFE1EF9BD" box="[263,422,1531,1553]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="livida">
<emphasis id="B93BEAEFFFDD85507D43FA3CFE1EF9BD" box="[263,422,1531,1553]" italics="true" pageId="11" pageNumber="334">Rhamma livida</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507DFCFA3CFE6FF9BC" bold="true" box="[440,471,1531,1552]" pageId="11" pageNumber="334">55.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507DA6FA3CFDC7F9BD" box="[482,639,1531,1553]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="oxida">
<emphasis id="B93BEAEFFFDD85507DA6FA3CFDC7F9BD" box="[482,639,1531,1553]" italics="true" pageId="11" pageNumber="334">Rhamma oxida</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507ED5FA3CFD09F9BC" bold="true" box="[657,689,1531,1552]" pageId="11" pageNumber="334">56.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507EF8FA3CFC3DF9BD" box="[700,901,1531,1553]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="comstocki">
<emphasis id="B93BEAEFFFDD85507EF8FA3CFC3DF9BD" box="[700,901,1531,1553]" italics="true" pageId="11" pageNumber="334">Rhamma comstocki</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507FD3FA3CFC0FF9BC" bold="true" box="[919,951,1531,1552]" pageId="11" pageNumber="334">57.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507F86FA3CFBC2F9BD" box="[962,1146,1531,1553]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="shapiroi">
<emphasis id="B93BEAEFFFDD85507F86FA3CFBC2F9BD" box="[962,1146,1531,1553]" italics="true" pageId="11" pageNumber="334">Rhamma shapiroi</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD855078C8FA3CFB14F9BC" bold="true" box="[1164,1196,1531,1552]" pageId="11" pageNumber="334">58.</emphasis>
<taxonomicName id="4C4F4D7EFFDD855078F3FA3CFAD3F9BD" box="[1207,1387,1531,1553]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="mishma">
<emphasis id="B93BEAEFFFDD855078F3FA3CFAD3F9BD" box="[1207,1387,1531,1553]" italics="true" pageId="11" pageNumber="334">Rhamma mishma</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507938FA3CFA24F9BC" bold="true" box="[1404,1436,1531,1552]" pageId="11" pageNumber="334">59.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507CD3F9DEFE9CF983" box="[151,292,1561,1583]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="bilix">
<emphasis id="B93BEAEFFFDD85507CD3F9DEFE9CF983" box="[151,292,1561,1583]" italics="true" pageId="11" pageNumber="334">Rhamma bilix</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507D75F9DEFEE9F982" bold="true" box="[305,337,1561,1582]" pageId="11" pageNumber="334">60.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507D13F9DEFDBFF983" box="[343,519,1561,1583]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="anosma">
<emphasis id="B93BEAEFFFDD85507D13F9DEFDBFF983" box="[343,519,1561,1583]" italics="true" pageId="11" pageNumber="334">Rhamma anosma</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507E50F9DEFD8CF982" bold="true" box="[532,564,1561,1582]" pageId="11" pageNumber="334">61.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507E7EF9DEFD44F983" box="[570,764,1561,1583]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="familiaris">
<emphasis id="B93BEAEFFFDD85507E7EF9DEFD44F983" box="[570,764,1561,1583]" italics="true" pageId="11" pageNumber="334">Rhamma familiaris</emphasis>
</taxonomicName>
;
<emphasis id="B93BEAEFFFDD85507F4EF9DEFC92F982" bold="true" box="[778,810,1561,1582]" pageId="11" pageNumber="334">62.</emphasis>
<taxonomicName id="4C4F4D7EFFDD85507F74F9DEFC44F983" box="[816,1020,1561,1583]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="species" species="commodus">
<emphasis id="B93BEAEFFFDD85507F74F9DEFC44F983" box="[816,1020,1561,1583]" italics="true" pageId="11" pageNumber="334">Rhamma commodus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3556576FFDD85507C83F991FE39F953" pageId="11" pageNumber="334" type="diagnosis">
<paragraph id="8BF036FDFFDD85507C83F991FE39F953" blockId="11.[151,1436,1622,1791]" pageId="11" pageNumber="334">
<emphasis id="B93BEAEFFFDD85507C83F991FEF9F9C3" bold="true" box="[199,321,1622,1647]" pageId="11" pageNumber="334">Diagnosis.</emphasis>
Males and females are immediately distinguished from all other
<taxonomicName id="4C4F4D7EFFDD85507817F991FB0FF9C3" box="[1107,1207,1622,1647]" class="Insecta" family="Lycaenidae" genus="Rhamma" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAEFFFDD85507817F991FB0FF9C3" box="[1107,1207,1622,1647]" italics="true" pageId="11" pageNumber="334">Rhamma</emphasis>
</taxonomicName>
by their large size (Forewing length&gt;
<quantity id="4CB79B18FFDD85507D34F9BCFE7AF93F" box="[368,450,1659,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="334" unit="mm" value="14.0">14 mm</quantity>
), green color and inconspicuous scent brand on the dorsal surface.
<emphasis id="B93BEAEFFFDD85507C83F959FE13F91B" bold="true" box="[199,427,1694,1719]" pageId="11" pageNumber="334">Material examined</emphasis>
(
<specimenCount id="9D49FD74FFDD85507DF9F958FE49F91B" box="[445,497,1695,1719]" pageId="11" pageNumber="334" type="generic">18♂</specimenCount>
, 1♀).
<emphasis id="B93BEAEFFFDD85507E7FF959FD58F91B" bold="true" box="[571,736,1694,1719]" pageId="11" pageNumber="334">ANTIOQUIA</emphasis>
:
<specimenCount id="9D49FD74FFDD85507EB5F958FCAEF91B" box="[753,790,1695,1719]" pageId="11" pageNumber="334" type="generic">8♂</specimenCount>
, CP: Bélmira, Páramo de Bélmira,
<quantity id="4CB79B18FFDD855078EBF958FAB0F91B" box="[1199,1288,1695,1719]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="334" unit="m" value="3200.0">3200 m</quantity>
,
<date id="FFF1103DFFDD85507952F959FA20F91B" box="[1302,1432,1694,1719]" pageId="11" pageNumber="334" value="2014-01-05">05/01/2014</date>
,
<date id="FFF1103DFFDD85507CD3F905FEA1F977" box="[151,281,1730,1755]" pageId="11" pageNumber="334" value="2015-08-01">01/08/2015</date>
, C. Prieto;
<specimenCount id="9D49FD74FFDD85507DDCF904FE05F977" box="[408,445,1731,1755]" pageId="11" pageNumber="334" type="generic">9♂</specimenCount>
, CP: 1♀, Yarumal, La Marconia,
<quantity id="4CB79B18FFDD85507F7CF904FC28F976" box="[824,912,1731,1755]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="334" unit="m" value="2700.0">2700 m</quantity>
,
<date id="FFF1103DFFDD85507FD8F905FBA7F977" box="[924,1055,1730,1755]" pageId="11" pageNumber="334" value="2015-07-28">28/07/2015</date>
;
<specimenCount id="9D49FD74FFDD8550786EF904FBF7F977" box="[1066,1103,1731,1755]" pageId="11" pageNumber="334" type="generic">1♂</specimenCount>
, CP: El retiro, San Sebastian,
<quantity id="4CB79B18FFDD85507CD3F920FF56F953" box="[151,238,1767,1791]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="334" unit="m" value="2800.0">2800 m</quantity>
,
<date id="FFF1103DFFDD85507CBEF920FEC5F953" box="[250,381,1767,1791]" pageId="11" pageNumber="334" value="2007-08-18">18/08/2007</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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