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<document id="AC4B24DB05A389DF4A813C9DCDD5A1D2" ID-CLB-Dataset="4912" ID-DOI="10.11646/zootaxa.3854.1.1" ID-GBIF-Dataset="73bc52e8-f82a-4258-94fc-adcbdf637e54" ID-ISSN="1175-5326" ID-Zenodo-Dep="4929336" ID-ZooBank="7F874BB5-91EB-41CC-A039-E98E7B53F47C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623438164503" checkinUser="felipe" docAuthor="Robillard, Tony, Gorochov, Andrej V., Poulain, Simon &amp; Suhardjono, Yayuk R." docDate="2014" docId="03E687D65B0BDE7CFF10E998A181FD8C" docLanguage="en" docName="zt03854p104.pdf" docOrigin="Zootaxa 3854 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cardiodactylus obi Gorochov &amp; Robillard 2014, n. sp." docType="treatment" docVersion="8" lastPageNumber="77" masterDocId="FFDFFFAE5B41DE30FF87E867A032FFB1" masterDocTitle="Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species" masterLastPageNumber="104" masterPageNumber="1" pageNumber="75" updateTime="1699012453132" updateUser="ExternalLinkService">
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<mods:title id="07870136E08B4CC2801FC9D84B17D727">Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species</mods:title>
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<mods:namePart id="553A6512EF56F1AB10BBCBA175E86D45">Robillard, Tony</mods:namePart>
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<mods:namePart id="4039A260A4226DFC859441AA00E2415B">Gorochov, Andrej V.</mods:namePart>
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<mods:namePart id="D8DC2513C8AF89C4A40398A6878FA20D">Poulain, Simon</mods:namePart>
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<mods:namePart id="9656286AB2FDEB9C74D371389AA37F00">Suhardjono, Yayuk R.</mods:namePart>
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<mods:title id="A947CFCA37D9CE1675A1E7FE8701A8CF">Zootaxa</mods:title>
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<mods:date id="AD2BFC2CAC6D061A9627087AE0D65535">2014</mods:date>
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<mods:number id="B41185C5D886B9A50CE15F73DD6BAB94">2014-08-20</mods:number>
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<mods:classification id="B02FEFA411D93424F120B068158D9985">journal article</mods:classification>
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<mods:identifier id="74C15AC860C78F6A29E438D47761A72B" type="DOI">10.11646/zootaxa.3854.1.1</mods:identifier>
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<treatment id="03E687D65B0BDE7CFF10E998A181FD8C" ID-DOI="http://doi.org/10.5281/zenodo.5227378" ID-GBIF-Taxon="184626125" ID-Zenodo-Dep="5227378" LSID="urn:lsid:plazi:treatment:03E687D65B0BDE7CFF10E998A181FD8C" httpUri="http://treatment.plazi.org/id/03E687D65B0BDE7CFF10E998A181FD8C" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75">
<subSubSection id="C355654B5B0BDE7AFF10E998A2DFFDAB" box="[151,749,511,538]" pageId="74" pageNumber="75" type="nomenclature">
<paragraph id="8BF036C05B0BDE7AFF10E998A2DFFDAB" blockId="74.[151,749,511,571]" box="[151,749,511,538]" pageId="74" pageNumber="75">
<heading id="D0B881AC5B0BDE7AFF10E998A2DFFDAB" bold="true" box="[151,749,511,538]" fontSize="11" level="2" pageId="74" pageNumber="75" reason="4">
<emphasis id="B93BEAD25B0BDE7AFF10E998A2DFFDAB" bold="true" box="[151,749,511,538]" pageId="74" pageNumber="75">
<taxonomicName id="4C4F4D435B0BDE7AFF10E998A2ACFDAB" authority="Gorochov &amp; Robillard" authorityName="Gorochov &amp; Robillard" authorityYear="2014" box="[151,670,511,538]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="obi" status="sp. nov.">
<emphasis id="B93BEAD25B0BDE7AFF10E998A148FDA8" bold="true" box="[151,378,511,537]" italics="true" pageId="74" pageNumber="75">Cardiodactylus obi</emphasis>
Gorochov &amp; Robillard
</taxonomicName>
,
<taxonomicNameLabel id="A20857A95B0BDE7AFD2CEA67A2DFFDAB" box="[683,749,512,538]" pageId="74" pageNumber="75" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C355654B5B0BDE7AFF10EA45A272FD8A" box="[151,576,546,571]" pageId="74" pageNumber="75" type="description">
<paragraph id="8BF036C05B0BDE7AFF10EA45A272FD8A" blockId="74.[151,749,511,571]" box="[151,576,546,571]" pageId="74" pageNumber="75">
(
<figureCitation id="13742A455B0BDE7AFF18EA45A0D5FD8A" box="[159,231,546,571]" captionStart="FIGURE 4" captionStartId="9.[151,250,1958,1980]" captionTargetBox="[166,1423,428,1928]" captionTargetId="figure-102@9.[151,1436,410,1936]" captionTargetPageId="9" captionText="FIGURE 4. Distributions of Cardiodactylus species around the Wallace line. The species distributions are hypothesized broadly according to the material examined and type localities. The Walace line is represented in red." figureDoi="http://doi.org/10.5281/zenodo.4929352" httpUri="https://zenodo.org/record/4929352/files/figure.png" pageId="74" pageNumber="75">Figs 4</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFF74EA45A12AFD8A" box="[243,280,546,571]" captionStart="FIGURE 6" captionStartId="15.[151,250,1881,1903]" captionTargetBox="[248,1359,218,1849]" captionTargetId="figure-123@15.[228,1365,193,1860]" captionTargetPageId="15" captionText="FIGURE 6. Head of Cardiodactylus species in dorsal (left) and facial (right) views: Cardiodactylus lombrinjani Robillard, n. sp. (A); C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. palawan Gorochov, n. sp. (E); C. pelagus Otte, 2007 (F); C. reticulatus Gorochov, n. sp. (G); C. riga Otte, 2007 (H); C. rizali Robillard, n. sp. (I); C. sumba Robillard, n. sp. (J); C. talaudae Robillard, n. sp. (K); C. tangkoko Gorochov, n. sp. (L); C. tello Robillard, n. sp. (M); C. variegatus Gorochov, n. sp. (N). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929356" httpUri="https://zenodo.org/record/4929356/files/figure.png" pageId="74" pageNumber="75">6D</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFEA3EA45A164FD8A" box="[292,342,546,571]" captionStart="FIGURE 10" captionStartId="19.[151,250,1908,1930]" captionTargetBox="[162,1383,203,1876]" captionTargetId="figure-89@19.[151,1436,193,1886]" captionTargetPageId="19" captionText="FIGURE 10. Male forewing venation of Cardiodactylus species: Cardiodactylus lombrinjani Robillard, n. sp. (A), C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. pelagus Otte, 2007 (E); C. rizali Robillard, n. sp. (F); C. sumba Robillard, n. sp.(G); C. talaudae Robillard, n. sp. (H); C. tello Robillard, n. sp. (I). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929372" httpUri="https://zenodo.org/record/4929372/files/figure.png" pageId="74" pageNumber="75">10D</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFEE5EA45A1A1FD8A" box="[354,403,546,571]" captionStart="FIGURE 12" captionStartId="21.[151,250,1909,1931]" captionTargetBox="[192,1402,188,1891]" captionTargetId="figure-92@21.[174,1412,177,1916]" captionTargetPageId="21" captionText="FIGURE 12. Female forewing venation of Cardiodactylus species: Cardiodactylus kondoi Otte, 2007 (A); C. muiri Otte, 2007 (B); C. obi Gorochov &amp; Robillard, n. sp. (C), C. oeroe Robillard, n. sp. (D); C. lampongsi Robillard &amp; Gorochov, n. sp. (E); C. sumba Robillard, n. sp. (F); C. tello Robillard, n. sp. (G); C. pelagus Otte, 2007 (H); C. variegatus Gorochov &amp; Robillard, n. sp. (I). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929380" httpUri="https://zenodo.org/record/4929380/files/figure.png" pageId="74" pageNumber="75">12C</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFE19EA45A1E3FD8A" box="[414,465,546,571]" captionStart="FIGURE 13" captionStartId="22.[151,250,1815,1837]" captionTargetBox="[264,1326,246,1803]" captionTargetId="figure-164@22.[230,1351,193,1803]" captionTargetPageId="22" captionText="FIGURE 13. Ovipositor of Cardiodactylus species: Cardiodactylus borneoe Robillard &amp; Gorochov, n. sp. (A); C. buru Gorochov &amp; Robillard, n. sp. (B); C. erniae Robillard &amp; Gorochov, n. sp. (C); C. floresiensis Robillard, n. sp. (D); C. fruhstorferi Gorochov &amp; Robillard, n. sp. (E); C. halmahera Gorochov &amp; Robillard, n. sp. (F); C. kondoi Otte, 2007 (G); C. kotandora Robillard, n. sp. (H); C. lampongsi Robillard &amp; Gorochov, n. sp. (I); C. loboe Robillard, n. sp. (J); C. lombrinjani Robillard, n. sp. (K); C. muiri Otte, 2007 (L); C. muria Robillard, n. sp. (M); C. obi Gorochov &amp; Robillard, n. sp. (N); C. oeroe Robillard, n. sp. (O); C. pelagus Otte, 2007 (P); C. sumba Robillard, n. sp. (Q); C. tello Robillard, n. sp. (R); C. variegatus Gorochov &amp; Robillard, n. sp. (S). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929388" httpUri="https://zenodo.org/record/4929388/files/figure.png" pageId="74" pageNumber="75">13N</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFE5AEA45A222FD8A" box="[477,528,546,571]" captionStart="FIGURE 15" captionStartId="24.[151,250,1416,1438]" captionTargetBox="[208,1378,201,1380]" captionTargetId="figure-184@24.[196,1392,191,1387]" captionTargetPageId="24" captionText="FIGURE 15. Female copulatory papilla of Cardiodactylus species in ventral (left), dorsal (middle) and lateral (right) views: C. lombrinjani Robillard, n. sp. (A); C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. oeroe Robillard, n. sp. (E); C. pelagus Otte, 2007 (F); C. sumba Robillard, n. sp. (G); C. tello Robillard, n. sp. (H); C. variegatus Gorochov &amp; Robillard, n. sp. (I). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929398" httpUri="https://zenodo.org/record/4929398/files/figure.png" pageId="74" pageNumber="75">15D</figureCitation>
,
<figureCitation id="13742A455B0BDE7AFD9CEA45A20AFD8A" box="[539,568,546,571]" captionStart="FIGURE 45" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[210,1421,230,1880]" captionTargetId="figure-67@75.[156,1431,193,1932]" captionTargetPageId="75" captionText="FIGURE 45. Cardiodactylus obi Gorochov &amp; Robillard, n. sp.: male in dorsal (A) and lateral (B) views; female in dorsal (C) and lateral (D) views; male genitalia in dorsal (E), ventral (F) and lateral (G) views. Scale bars: AD: 5 mm; EG: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4930322" httpUri="https://zenodo.org/record/4930322/files/figure.png" pageId="74" pageNumber="75">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C355654B5B0BDE7AFF10EA0DA250FD5E" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="8BF036C05B0BDE7AFF10EA0DA302FD7A" blockId="74.[151,1437,618,2011]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF10EA0DA17CFD32" bold="true" box="[151,334,618,643]" pageId="74" pageNumber="75">Type material.</emphasis>
<materialsCitation id="3B273C9D5B0BDE7AFED8EA0DA181FD16" ID-GBIF-Occurrence="3336528310" collectingDate="1930-04-23" collectingDateMax="1930-04-27" collectingDateMin="1930-04-23" collectionCode="RMNH" collectorName="Male &amp; Obi I." country="Indonesia" location="Obi Latoe" pageId="74" pageNumber="75" specimenCount="1" stateProvince="Maluku Utara" typeStatus="holotype">
<collectorName id="26BA53165B0BDE7AFED8EA0DA1AAFD32" box="[351,408,618,643]" pageId="74" pageNumber="75">Male</collectorName>
<typeStatus id="54F488625B0BDE7AFE2EEA0DA222FD32" box="[425,528,618,643]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
:
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<collectingCountry id="F35876505B0BDE7AFDA4EA0DA2A8FD32" box="[547,666,618,643]" name="Indonesia" pageId="74" pageNumber="75">Indonesia</collectingCountry>
.
<collectingRegion id="498BF8225B0BDE7AFD28EA0DA3ECFD32" box="[687,990,618,643]" country="Indonesia" name="Maluku Utara" pageId="74" pageNumber="75">Maluku Utara Province</collectingRegion>
</emphasis>
,
<location id="8E90601B5B0BDE7AFC71EA0DA443FD33" LSID="urn:lsid:plazi:treatment:03E687D65B0BDE7CFF10E998A181FD8C:8E90601B5B0BDE7AFC71EA0DA443FD33" box="[1014,1137,618,643]" country="Indonesia" name="Obi Latoe" pageId="74" pageNumber="75" stateProvince="Maluku Utara">Obi Latoe</location>
[
<collectorName id="26BA53165B0BDE7AFB0FEA0DA4E5FD33" box="[1160,1239,618,643]" pageId="74" pageNumber="75">Obi I.</collectorName>
], Snellius Exp.,
<date id="FFF110005B0BDE7AFF10EAE9A10FFD16" box="[151,317,654,679]" pageId="74" pageNumber="75" value="1930-04-23" valueMax="1930-04-27" valueMin="1930-04-23">
<collectingDate id="EFB5E9E85B0BDE7AFF10EAE9A10FFD16" box="[151,317,654,679]" pageId="74" pageNumber="75" value="1930-04-23" valueMax="1930-04-27" valueMin="1930-04-23">2327.IV.1930</collectingDate>
</date>
(
<collectionCode id="ED5EAE055B0BDE7AFED7EAE9A199FD16" box="[336,427,654,679]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="74" pageNumber="75" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B273C9D5B0BDE7AFE46EAE8A0E9FD7A" ID-GBIF-Occurrence="3336528342" collectionCode="ZIN" collectorName="Obi I." country="Indonesia" location="Rolle" municipality="Obi" pageId="74" pageNumber="75" specimenCount="3" specimenCount-female="3" stateProvince="Maluku Utara" typeStatus="paratype">
<typeStatus id="54F488625B0BDE7AFE46EAE8A21CFD16" box="[449,558,655,679]" pageId="74" pageNumber="75" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D49FD495B0BDE7AFDC3EAE9A25AFD16" box="[580,616,654,679]" pageId="74" pageNumber="75" type="female">2♀</specimenCount>
):
<emphasis id="B93BEAD25B0BDE7AFD05EAE9A41FFD16" bold="true" box="[642,1069,654,679]" pageId="74" pageNumber="75">
<collectingCountry id="F35876505B0BDE7AFD05EAE9A2CBFD16" box="[642,761,654,679]" name="Indonesia" pageId="74" pageNumber="75">Indonesia</collectingCountry>
.
<collectingRegion id="498BF8225B0BDE7AFC8EEAE9A41FFD16" box="[777,1069,654,679]" country="Indonesia" name="Maluku Utara" pageId="74" pageNumber="75">Maluku Utara Province</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6B94ACBA5B0BDE7AFBC7EAE9A458FD16" box="[1088,1130,654,679]" pageId="74" pageNumber="75">Obi</collectingMunicipality>
[
<collectorName id="26BA53165B0BDE7AFBFBEAE9A4F4FD17" box="[1148,1222,654,679]" pageId="74" pageNumber="75">Obi I.</collectorName>
], 1904,
<specimenCount id="9D49FD495B0BDE7AFAADEAE9A57FFD16" box="[1322,1357,654,679]" pageId="74" pageNumber="75" type="female">1♀</specimenCount>
,
<location id="8E90601B5B0BDE7AFAD8EAE9A5A9FD16" LSID="urn:lsid:plazi:treatment:03E687D65B0BDE7CFF10E998A181FD8C:8E90601B5B0BDE7AFAD8EAE9A5A9FD16" box="[1375,1435,654,679]" country="Indonesia" municipality="Obi" name="Rolle" pageId="74" pageNumber="75" stateProvince="Maluku Utara">Rolle</location>
(
<collectionCode id="ED5EAE055B0BDE7AFF18EAD5A0E1FD7A" box="[159,211,690,715]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="74" pageNumber="75" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B273C9D5B0BDE7AFF60EAD5A31EFD7A" ID-GBIF-Occurrence="3336528458" box="[231,812,690,715]" collectionCode="MHNG" collectorName="Obi I. &amp; Fruhstorfer" country="Indonesia" pageId="74" pageNumber="75" specimenCount="1" specimenCount-female="1" stateProvince="Maluku Utara" typeStatus="paratype">
[
<collectorName id="26BA53165B0BDE7AFF6AEAD5A101FD7B" box="[237,307,690,715]" pageId="74" pageNumber="75">Obi I.</collectorName>
], Doherty,
<specimenCount id="9D49FD495B0BDE7AFE36EAD5A1E6FD7A" box="[433,468,690,715]" pageId="74" pageNumber="75" type="female">1♀</specimenCount>
, ex coll.
<collectorName id="26BA53165B0BDE7AFDBCEAD5A28BFD7A" box="[571,697,690,715]" pageId="74" pageNumber="75">Fruhstorfer</collectorName>
(
<collectionCode id="ED5EAE055B0BDE7AFD4FEAD5A316FD7A" box="[712,804,690,715]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="74" pageNumber="75" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036C05B0BDE7AFF42EAB1A250FD5E" blockId="74.[151,1437,618,2011]" box="[197,610,726,751]" pageId="74" pageNumber="75">
<materialsCitation id="3B273C9D5B0BDE7AFF42EAB1A26CFD5E" ID-GBIF-Occurrence="3336528352" box="[197,606,726,751]" country="Indonesia" location="Obi Island" pageId="74" pageNumber="75" specimenCount="1" typeStatus="holotype">
<emphasis id="B93BEAD25B0BDE7AFF42EAB1A151FD5F" bold="true" box="[197,355,726,751]" pageId="74" pageNumber="75">
<typeStatus id="54F488625B0BDE7AFF42EAB1A132FD5E" box="[197,256,726,751]" pageId="74" pageNumber="75">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35876505B0BDE7AFEEDEAB1A1E8FD5E" box="[362,474,726,751]" name="Indonesia" pageId="74" pageNumber="75">Indonesia</collectingCountry>
,
<location id="8E90601B5B0BDE7AFE62EAB1A26CFD5E" LSID="urn:lsid:plazi:treatment:03E687D65B0BDE7CFF10E998A181FD8C:8E90601B5B0BDE7AFE62EAB1A26CFD5E" box="[485,606,726,751]" country="Indonesia" name="Obi Island" pageId="74" pageNumber="75">Obi Island</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355654B5B0BDE7AFF42EA9DA2ADFCA2" box="[197,671,762,787]" pageId="74" pageNumber="75" type="etymology">
<paragraph id="8BF036C05B0BDE7AFF42EA9DA2ADFCA2" blockId="74.[151,1437,618,2011]" box="[197,671,762,787]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EA9DA17EFCA2" bold="true" box="[197,332,762,787]" pageId="74" pageNumber="75">Etymology.</emphasis>
Named after the
<typeStatus id="54F488625B0BDE7AFD89EA9CA20CFCA2" box="[526,574,763,787]" pageId="74" pageNumber="75">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C355654B5B0BDE7AFF42EB79A26FFC86" box="[197,605,798,823]" pageId="74" pageNumber="75" type="distribution">
<paragraph id="8BF036C05B0BDE7AFF42EB79A26FFC86" blockId="74.[151,1437,618,2011]" box="[197,605,798,823]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EB79A16CFC86" bold="true" box="[197,350,798,823]" pageId="74" pageNumber="75">Distribution.</emphasis>
<collectingCountry id="F35876505B0BDE7AFEE2EB79A1E7FC86" box="[357,469,798,823]" name="Indonesia" pageId="74" pageNumber="75">Indonesia</collectingCountry>
, Obi Island.
</paragraph>
</subSubSection>
<subSubSection id="C355654B5B0BDE7AFF42EB25A5B5FCCE" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="8BF036C05B0BDE7AFF42EB25A5B5FCCE" blockId="74.[151,1437,618,2011]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EB25A10DFCEA" bold="true" box="[197,319,834,859]" pageId="74" pageNumber="75">Diagnosis.</emphasis>
Species of average to large size, close to
<taxonomicName id="4C4F4D435B0BDE7AFC9CEB24A423FCEA" authority="Otte, 2007 a" authorityName="Otte" authorityYear="2007" box="[795,1041,834,859]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="manus">
<emphasis id="B93BEAD25B0BDE7AFC9CEB24A3B7FCEB" box="[795,901,835,858]" italics="true" pageId="74" pageNumber="75">C. manus</emphasis>
<bibRefCitation id="EFDE4B315B0BDE7AFC0AEB25A423FCEA" author="Otte, D." box="[909,1041,834,859]" pageId="74" pageNumber="75" pagination="341 - 400" refId="ref50945" refString="Otte, D. (2007 a) New species of Cardiodactylus from the western Pacific region (Gryllidae: Eneopterinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 156, 341 - 400. http: // dx. doi. org / 10.1635 / 0097 - 3157 (2007) 156 [341: nsocft] 2.0. co; 2" type="journal article" year="2007">Otte, 2007a</bibRefCitation>
</taxonomicName>
by the male genitalia but differing by coloration (male and female FWs, lateral lobes of pronotum mostly dark brown) and details of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C355654B5B0BDE7CFF42EBEDA181FD8C" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75" type="description">
<paragraph id="8BF036C05B0BDE7AFF42EBEDA4AAFB56" blockId="74.[151,1437,618,2011]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EBEDA165FC12" bold="true" box="[197,343,906,931]" pageId="74" pageNumber="75">Description.</emphasis>
Size average to large for the species group, shape stocky (
<figureCitation id="13742A455B0BDE7AFB9AEBEDA49CFC12" box="[1053,1198,906,931]" captionStart="FIGURE 45" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[210,1421,230,1880]" captionTargetId="figure-67@75.[156,1431,193,1932]" captionTargetPageId="75" captionText="FIGURE 45. Cardiodactylus obi Gorochov &amp; Robillard, n. sp.: male in dorsal (A) and lateral (B) views; female in dorsal (C) and lateral (D) views; male genitalia in dorsal (E), ventral (F) and lateral (G) views. Scale bars: AD: 5 mm; EG: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4930322" httpUri="https://zenodo.org/record/4930322/files/figure.png" pageId="74" pageNumber="75">Fig. 45AD</figureCitation>
). General coloration contrasted. Head dorsum almost entirely dark brown (
<figureCitation id="13742A455B0BDE7AFC83EBC9A353FC76" box="[772,865,942,967]" captionStart="FIGURE 6" captionStartId="15.[151,250,1881,1903]" captionTargetBox="[248,1359,218,1849]" captionTargetId="figure-123@15.[228,1365,193,1860]" captionTargetPageId="15" captionText="FIGURE 6. Head of Cardiodactylus species in dorsal (left) and facial (right) views: Cardiodactylus lombrinjani Robillard, n. sp. (A); C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. palawan Gorochov, n. sp. (E); C. pelagus Otte, 2007 (F); C. reticulatus Gorochov, n. sp. (G); C. riga Otte, 2007 (H); C. rizali Robillard, n. sp. (I); C. sumba Robillard, n. sp. (J); C. talaudae Robillard, n. sp. (K); C. tangkoko Gorochov, n. sp. (L); C. tello Robillard, n. sp. (M); C. variegatus Gorochov, n. sp. (N). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929356" httpUri="https://zenodo.org/record/4929356/files/figure.png" pageId="74" pageNumber="75">Fig. 6D</figureCitation>
), with 4 wide brown bands almost fused together separated by faint yellow lines; fastigium dark brown; yellow lines posterior to eyes. Scapes yellow brown, with a dark brown ring; rest of antennae orange brown. Front part of fastigium yellow with 2 dark brown spots. Face and mouthparts yellowish to brown, with a transverse dark brown band bellow antennae and a yellow band bellow eyes. Lateral side of head mostly dark brown dorsally, progressively lighter ventrally; eyes underlined by a yellow line ventrally. Pronotum: Dorsal disk almost homogeneously brown, anterior corners yellow, posterior edge dark brown. Lateral lobes mostly dark brown, ventral margin yellowish. Legs almost homogeneously orange brown, FIII knees darker. Hind wing tail dark brown with yellow transverse veins. Cerci light brown faintly mottled with brown. Abdomen mostly dark, with yellow stripes laterally. Subgenital plate yellow brown.
</paragraph>
<paragraph id="8BF036C05B0BDE7AFF42EC95A3CDFA52" blockId="74.[151,1437,618,2011]" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EC95A138FABA" bold="true" box="[197,266,1266,1291]" pageId="74" pageNumber="75">Male:</emphasis>
FW coloration dark brown with the following whitish areas (
<figureCitation id="13742A455B0BDE7AFC39EC95A415FABA" box="[958,1063,1266,1291]" captionStart="FIGURE 10" captionStartId="19.[151,250,1908,1930]" captionTargetBox="[162,1383,203,1876]" captionTargetId="figure-89@19.[151,1436,193,1886]" captionTargetPageId="19" captionText="FIGURE 10. Male forewing venation of Cardiodactylus species: Cardiodactylus lombrinjani Robillard, n. sp. (A), C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. pelagus Otte, 2007 (E); C. rizali Robillard, n. sp. (F); C. sumba Robillard, n. sp.(G); C. talaudae Robillard, n. sp. (H); C. tello Robillard, n. sp. (I). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929372" httpUri="https://zenodo.org/record/4929372/files/figure.png" pageId="74" pageNumber="75">Fig. 10D</figureCitation>
): base of FWs including bases of anal veins and CuA, wide transverve band posterior to mirror, part of chords, 1A(file) and harp veins. Orange brown sclerotized area near base of chords. M and R orange brown, the area between dark brown; Sc orange brown, its projections whitish, the membrane between them dark brown. FW venation (
<figureCitation id="13742A455B0BDE7AFB1AED39A539FAC6" box="[1181,1291,1374,1399]" captionStart="FIGURE 10" captionStartId="19.[151,250,1908,1930]" captionTargetBox="[162,1383,203,1876]" captionTargetId="figure-89@19.[151,1436,193,1886]" captionTargetPageId="19" captionText="FIGURE 10. Male forewing venation of Cardiodactylus species: Cardiodactylus lombrinjani Robillard, n. sp. (A), C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. pelagus Otte, 2007 (E); C. rizali Robillard, n. sp. (F); C. sumba Robillard, n. sp.(G); C. talaudae Robillard, n. sp. (H); C. tello Robillard, n. sp. (I). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929372" httpUri="https://zenodo.org/record/4929372/files/figure.png" pageId="74" pageNumber="75">Fig. 10D</figureCitation>
): 1A clearly bisinuated. CuP absent. Harp with 2 w-shaped veins, and a small straight anterior one. Mirror area: mirror (d1) oval, distinct, crossed at 2/3 by a transverse vein; d2 not distinct. Apical field with 4 cell alignments posterior to mirror (n = 1). Lateral field with 9 projections of Sc and 4 more ventral veins.
</paragraph>
<paragraph id="8BF036C05B0BDE7AFF42ED89A257F8FA" blockId="74.[151,1437,618,2011]" pageId="74" pageNumber="75">
Male genitalia (
<figureCitation id="13742A455B0BDE7AFEFFED89A230F9B6" box="[376,514,1518,1543]" captionStart="FIGURE 45" captionStartId="75.[151,250,1953,1975]" captionTargetBox="[210,1421,230,1880]" captionTargetId="figure-67@75.[156,1431,193,1932]" captionTargetPageId="75" captionText="FIGURE 45. Cardiodactylus obi Gorochov &amp; Robillard, n. sp.: male in dorsal (A) and lateral (B) views; female in dorsal (C) and lateral (D) views; male genitalia in dorsal (E), ventral (F) and lateral (G) views. Scale bars: AD: 5 mm; EG: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4930322" httpUri="https://zenodo.org/record/4930322/files/figure.png" pageId="74" pageNumber="75">Fig. 45EG</figureCitation>
): Pseudepiphallus as in
<taxonomicName id="4C4F4D435B0BDE7AFC94ED88A34CF9B7" authorityName="Otte" authorityYear="2007" box="[787,894,1519,1542]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="manus">
<emphasis id="B93BEAD25B0BDE7AFC94ED88A34CF9B7" box="[787,894,1519,1542]" italics="true" pageId="74" pageNumber="75">C. manus</emphasis>
</taxonomicName>
, shaped as a wide gutter. Apex of dorsal ridges with short setae, carinated innerly and folded laterally, the lateral triangular expansions clearly asymmetrical, the left one larger and more posterior, the right one smaller and more anterior. Dorsal ridges almost convex in lateral view. Pseudepiphallic sclerite with wide rectangular anterior expansions, slightly divergent, the membrane between them with thin setae. Rami with large preapical rectangular plates and with short globular convergent apical stems. Posterior pseudepiphallic apex rounded, with a triangular translucent area. Pseudepiphallic parameres with posterior arms thin and curved laterally. Ectophallic arc complete, curved posteriorly, without posterior expansion. Ectophallic fold with thin lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite small, with a wide posterior expansion. Endophallic apodeme with a short dorsal crest and lateral lamellas. Membrane of endophallic cavity smooth.
</paragraph>
<paragraph id="8BF036C05B0BDE7CFF42EF31A351FEAC" blockId="74.[151,1437,618,2011]" lastBlockId="76.[151,1436,151,573]" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75">
<emphasis id="B93BEAD25B0BDE7AFF42EF31A117F8DE" bold="true" box="[197,293,1878,1903]" pageId="74" pageNumber="75">Female:</emphasis>
Base of tegmina yellow; dorsal field variable,
<specimenCount id="9D49FD495B0BDE7AFCF3EF30A424F8DE" box="[884,1046,1878,1903]" pageId="74" pageNumber="75" type="generic">one specimen</specimenCount>
with practically uniform brown coloration (ZIN) with only one lateral yellowish spot, the other female specimen with an additional yellowish elongate pattern in anterior part of tegmina (MHNG) (
<figureCitation id="13742A455B0BDE7AFD72EFF9A369F806" box="[757,859,1950,1975]" captionStart="FIGURE 12" captionStartId="21.[151,250,1909,1931]" captionTargetBox="[192,1402,188,1891]" captionTargetId="figure-92@21.[174,1412,177,1916]" captionTargetPageId="21" captionText="FIGURE 12. Female forewing venation of Cardiodactylus species: Cardiodactylus kondoi Otte, 2007 (A); C. muiri Otte, 2007 (B); C. obi Gorochov &amp; Robillard, n. sp. (C), C. oeroe Robillard, n. sp. (D); C. lampongsi Robillard &amp; Gorochov, n. sp. (E); C. sumba Robillard, n. sp. (F); C. tello Robillard, n. sp. (G); C. pelagus Otte, 2007 (H); C. variegatus Gorochov &amp; Robillard, n. sp. (I). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929380" httpUri="https://zenodo.org/record/4929380/files/figure.png" pageId="74" pageNumber="75">Fig. 12C</figureCitation>
); lateral field of tegmina with proximal branches of Sc and crossveins between them yellowish white, distal branches of Sc and membranes between all Sc branches light brown (however R, stem of Sc, and area between them brown, as in female of
<taxonomicName id="4C4F4D435B0DDE7CFB62E8FFA50CFF01" authority="Robillard &amp; Gorochov &amp; Poulain &amp; Suhardjono, 2014" authorityName="Robillard &amp; Gorochov &amp; Poulain &amp; Suhardjono" authorityYear="2014" box="[1253,1342,152,176]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="buru" status="sp. nov.">
<emphasis id="B93BEAD25B0DDE7CFB62E8FFA50CFF01" box="[1253,1342,152,176]" italics="true" pageId="76" pageNumber="77">C. buru</emphasis>
</taxonomicName>
<emphasis id="B93BEAD25B0DDE7CFACFE8F0A5B8FF01" bold="true" box="[1352,1418,151,176]" pageId="76" pageNumber="77">
<taxonomicNameLabel id="A20857A95B0DDE7CFACFE8F0A5B8FF01" box="[1352,1418,151,176]" pageId="76" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); tegminal venation with 1112 longitudinal veins in dorsal field and with 89 branches on Sc and distinctly less numerous crossveins in lateral field. Ovipositor approximately equal or shorter to hind femur in length, its apex slightly denticulate on both dorsal and ventral edges (
<figureCitation id="13742A455B0DDE7CFD69E963A365FEAC" box="[750,855,260,285]" captionStart="FIGURE 13" captionStartId="22.[151,250,1815,1837]" captionTargetBox="[264,1326,246,1803]" captionTargetId="figure-164@22.[230,1351,193,1803]" captionTargetPageId="22" captionText="FIGURE 13. Ovipositor of Cardiodactylus species: Cardiodactylus borneoe Robillard &amp; Gorochov, n. sp. (A); C. buru Gorochov &amp; Robillard, n. sp. (B); C. erniae Robillard &amp; Gorochov, n. sp. (C); C. floresiensis Robillard, n. sp. (D); C. fruhstorferi Gorochov &amp; Robillard, n. sp. (E); C. halmahera Gorochov &amp; Robillard, n. sp. (F); C. kondoi Otte, 2007 (G); C. kotandora Robillard, n. sp. (H); C. lampongsi Robillard &amp; Gorochov, n. sp. (I); C. loboe Robillard, n. sp. (J); C. lombrinjani Robillard, n. sp. (K); C. muiri Otte, 2007 (L); C. muria Robillard, n. sp. (M); C. obi Gorochov &amp; Robillard, n. sp. (N); C. oeroe Robillard, n. sp. (O); C. pelagus Otte, 2007 (P); C. sumba Robillard, n. sp. (Q); C. tello Robillard, n. sp. (R); C. variegatus Gorochov &amp; Robillard, n. sp. (S). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929388" httpUri="https://zenodo.org/record/4929388/files/figure.png" pageId="76" pageNumber="77">Fig. 13N</figureCitation>
).
</paragraph>
<caption id="DF3066485B0ADE7BFF10EFC6A557F867" ID-DOI="http://doi.org/10.5281/zenodo.4930322" ID-Zenodo-Dep="4930322" httpUri="https://zenodo.org/record/4930322/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,250,1953,1975]" targetBox="[210,1421,230,1880]" targetPageId="75">
<paragraph id="8BF036C05B0ADE7BFF10EFC6A557F867" blockId="75.[151,1436,1953,2006]" pageId="75" pageNumber="76">
<emphasis id="B93BEAD25B0ADE7BFF10EFC6A112F807" bold="true" box="[151,288,1953,1975]" pageId="75" pageNumber="76">FIGURE 45.</emphasis>
<taxonomicName id="4C4F4D435B0ADE7BFEA1EFC6A2EAF806" authority="Gorochov &amp; Robillard" authorityName="Gorochov &amp; Robillard" authorityYear="2014" box="[294,728,1953,1975]" class="Insecta" family="Haglotettigoniidae" genus="Cardiodactylus" kingdom="Animalia" order="Orthoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="obi" status="sp. nov.">
<emphasis id="B93BEAD25B0ADE7BFEA1EFC6A1DAF806" box="[294,488,1953,1975]" italics="true" pageId="75" pageNumber="76">Cardiodactylus obi</emphasis>
Gorochov &amp; Robillard
</taxonomicName>
,
<emphasis id="B93BEAD25B0ADE7BFD64EFC5A32FF806" bold="true" box="[739,797,1954,1975]" pageId="75" pageNumber="76">
<taxonomicNameLabel id="A20857A95B0ADE7BFD64EFC5A32FF806" box="[739,797,1954,1975]" pageId="75" pageNumber="76" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: male in dorsal (A) and lateral (B) views; female in dorsal (C) and lateral (D) views; male genitalia in dorsal (E), ventral (F) and lateral (G) views. Scale bars: AD: 5 mm; EG: 1 mm.
</paragraph>
</caption>
<paragraph id="8BF036C05B0DDE7CFF42E940A45EFED4" blockId="76.[151,1436,151,573]" pageId="76" pageNumber="77">
Female genitalia (
<figureCitation id="13742A455B0DDE7CFE0AE940A1C7FEF1" box="[397,501,295,320]" captionStart="FIGURE 15" captionStartId="24.[151,250,1416,1438]" captionTargetBox="[208,1378,201,1380]" captionTargetId="figure-184@24.[196,1392,191,1387]" captionTargetPageId="24" captionText="FIGURE 15. Female copulatory papilla of Cardiodactylus species in ventral (left), dorsal (middle) and lateral (right) views: C. lombrinjani Robillard, n. sp. (A); C. muiri Otte, 2007 (B); C. muria Robillard, n. sp. (C); C. obi Gorochov &amp; Robillard, n. sp. (D); C. oeroe Robillard, n. sp. (E); C. pelagus Otte, 2007 (F); C. sumba Robillard, n. sp. (G); C. tello Robillard, n. sp. (H); C. variegatus Gorochov &amp; Robillard, n. sp. (I). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4929398" httpUri="https://zenodo.org/record/4929398/files/figure.png" pageId="76" pageNumber="77">Fig. 15D</figureCitation>
): Copulatory papilla longer than wide, trapezoidal, with elongate and basal sclerites mostly on dorsal side; apex elongate, not rounded, slightly sclerotized, folded ventrally.
</paragraph>
<paragraph id="8BF036C05B0DDE7CFF42E908A19DFE39" blockId="76.[151,1436,151,573]" box="[197,431,367,392]" pageId="76" pageNumber="77">
<emphasis id="B93BEAD25B0DDE7CFF42E908A100FE39" bold="true" box="[197,306,367,392]" pageId="76" pageNumber="77">Juvenile:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF036C05B0DDE7CFF42E9F3A1C3FE44" blockId="76.[151,1436,151,573]" pageId="76" pageNumber="77">
<emphasis id="B93BEAD25B0DDE7CFF42E9F3A149FE1C" bold="true" box="[197,379,404,429]" pageId="76" pageNumber="77">Measurements.</emphasis>
Length in mm. Body: male 27.2, female 2132.2; body with wings: male 34.2, female 3141.7; pronotum: male 3.5, female 3.34.2; tegmina: male 20, female 1823.5; hind femora: male 22.8, female 1924.2; ovipositor 17.518.5.
</paragraph>
<paragraph id="8BF036C05B0DDE7CFF42E998A295FDA9" blockId="76.[151,1436,151,573]" box="[197,679,511,536]" pageId="76" pageNumber="77">
<emphasis id="B93BEAD25B0DDE7CFF42E998A218FDA9" bold="true" box="[197,554,511,536]" pageId="76" pageNumber="77">Habitat and life history traits.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF036C05B0DDE7CFF42EA43A181FD8C" blockId="76.[151,1436,151,573]" box="[197,435,548,573]" pageId="76" pageNumber="77">
<emphasis id="B93BEAD25B0DDE7CFF42EA43A105FD8C" bold="true" box="[197,311,548,573]" pageId="76" pageNumber="77">Behavior.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>