215 lines
34 KiB
XML
215 lines
34 KiB
XML
<document id="FFC4F9DDB5E173C9B1D152019B994CC1" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF7F8306FF2E16ADFF581E4F" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia zaragozae Hershler, Liu & Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="9" updateTime="1698261240372" updateUser="plazi">
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<mods:title id="103AEFACF980262265464C7124725D3B">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:namePart id="20FBBC92D548F5F7BF4327AAC550F541">Hershler, Robert</mods:namePart>
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<mods:namePart id="8AEC13C11452BB38C80D0F57571AE03D">Liu, Hsiu-Ping</mods:namePart>
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<mods:namePart id="EE5FA62183FCA0B2859E8F1C46780A38">Landye, Jerry</mods:namePart>
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<mods:typeOfResource id="01A2D202BE8F2E1B7A6D8EDB6A8ED02E">text</mods:typeOfResource>
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<mods:title id="21DD2606D50119727802E0BE5DB487EE">Zootaxa</mods:title>
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<mods:part id="FFCBA1DFEB56D75E9846395C0C3C0759">
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<mods:date id="2519DC5B098B068CCAD1466326C76993">2011</mods:date>
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<mods:classification id="85CE103201E2978C9F37C37167BA42EA">journal article</mods:classification>
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<mods:identifier id="20F9233826C1451892EFF126B36D7ACB" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="CFB792B1350E9E401DB1A1ADAE48A79E" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
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<treatment id="03E687C5BF7F8306FF2E16ADFF581E4F" ID-DOI="http://doi.org/10.5281/zenodo.5629191" ID-GBIF-Taxon="119406531" ID-Zenodo-Dep="5629191" LSID="urn:lsid:plazi:treatment:03E687C5BF7F8306FF2E16ADFF581E4F" httpUri="http://treatment.plazi.org/id/03E687C5BF7F8306FF2E16ADFF581E4F" lastPageNumber="9" pageId="8" pageNumber="9">
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<subSubSection id="C3556558BF7F8306FF2E16ADFC211B8A" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BF036D3BF7F8306FF2E16ADFE7E189E" blockId="8.[151,474,439,498]" box="[151,474,439,465]" pageId="8" pageNumber="9">
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<heading id="D0B881BFBF7F8306FF2E16ADFE7E189E" bold="true" box="[151,474,439,465]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B93BEAC1BF7F8306FF2E16ADFE7E189E" bold="true" box="[151,474,439,465]" pageId="8" pageNumber="9">
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<taxonomicName id="4C4F4D50BF7F8306FF2E16ADFED2189E" box="[151,374,439,465]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="zaragozae" status="sp. nov.">
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<emphasis id="B93BEAC1BF7F8306FF2E16ADFED2189E" bold="true" box="[151,374,439,465]" italics="true" pageId="8" pageNumber="9">Tryonia zaragozae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF7F8306FEC416ADFE7E189E" box="[381,474,439,465]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF7F8306FF2E16C0FE6018BD" blockId="8.[151,474,439,498]" box="[151,452,474,498]" pageId="8" pageNumber="9">
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(
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<figureCitation id="13742A56BF7F8306FF2616C0FF4018BD" box="[159,228,474,498]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Figs 4</figureCitation>
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E–G, J–L, P–R, 5B)
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</paragraph>
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<paragraph id="8BF036D3BF7F8306FF2E1539FCAF1B77" blockId="8.[151,779,546,568]" box="[151,779,546,568]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="AB5F11FDBF7F8306FF2E1539FCAF1B77" box="[151,779,546,568]" pageId="8" pageNumber="9">
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<taxonomicName id="4C4F4D50BF7F8306FF2E1539FF411B77" box="[151,229,547,568]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus" status="sp. nov.">
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<emphasis id="B93BEAC1BF7F8306FF2E1539FF411B77" box="[151,229,547,568]" italics="true" pageId="8" pageNumber="9">Tryonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF7F8306FF551539FE861B77" box="[236,290,547,568]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
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2.—
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<treatmentCitation id="0AEE10C2BF7F8306FEEB1538FDFA1B77" author="Hershler" box="[338,606,546,568]" page="1757" pageId="8" pageNumber="9" year="2005">
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<bibRefCitation id="EFDE4B22BF7F8306FEEB1538FD851B77" author="Hershler" box="[338,545,546,568]" pageId="8" pageNumber="9" refString="Hershler, R., Mulvey, M. & Liu, H. - P. (2005) Genetic variation in the Desert springsnail (Tryonia porrecta): implications for reproductive mode and dispersal. Molecular Ecology, 14, 1755 - 1765." type="journal article" year="2005">
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Hershler
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<emphasis id="B93BEAC1BF7F8306FE0B1539FE7A1B77" box="[434,478,546,568]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
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. 2005
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</bibRefCitation>
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: 1757
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</treatmentCitation>
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(COI sequence).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF036D3BF7F8306FF2E157FFC211B8A" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
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<emphasis id="B93BEAC1BF7F8306FF2E157FFF451B32" bold="true" box="[151,225,613,637]" pageId="8" pageNumber="9">
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<typeStatus id="54F48871BF7F8306FF2E157FFF7E1B32" box="[151,218,613,637]" pageId="8" pageNumber="9">Types</typeStatus>
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.
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</emphasis>
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<typeStatus id="54F48871BF7F8306FF52157FFEF21B32" box="[235,342,613,637]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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, USNM 873286, springs ca.
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<quantity id="4CB79B36BF7F8306FD25157FFD4F1B32" box="[668,747,613,637]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="9" unit="km" value="4.8">4.8 km</quantity>
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north-east of Ignacio Zaragoza along the south side of Highway 5, Chihuahua,
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<geoCoordinate id="EE7B5014BF7F8306FEC61593FD801BEE" box="[383,548,649,673]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="29.663305">29º39'47.9" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF7F8306FD8D1593FD4F1BEE" box="[564,747,649,673]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="-107.74049">107º44'25.8" W</geoCoordinate>
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, leg. J.J.L.
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<emphasis id="B93BEAC1BF7F8306FCC11590FC0A1BEE" box="[888,942,649,673]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
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.,
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<date id="FFF11013BF7F8306FC711593FBF11BEE" box="[968,1109,649,673]" pageId="8" pageNumber="9" value="1971-08-25">25/viii/1971</date>
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.
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<typeStatus id="54F48871BF7F8306FBDD1590FB751BED" box="[1124,1233,650,674]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 873260, USNM 873289, USNM 874120, USNM 1153679.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF7F8306FF7E15C8FAF41BA5" box="[199,1360,721,746]" pageId="8" pageNumber="9" type="etymology">
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||
<paragraph id="8BF036D3BF7F8306FF7E15C8FAF41BA5" blockId="8.[151,1436,613,1792]" box="[199,1360,721,746]" pageId="8" pageNumber="9">
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<emphasis id="B93BEAC1BF7F8306FF7E15C8FEE21BA5" bold="true" box="[199,326,722,746]" pageId="8" pageNumber="9">Etymology</emphasis>
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. The species name is a geographic epithet referring to the nearby town of Ignacio Zaragoza.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF7F8306FF7E15EFFB5C1A42" box="[199,1272,757,782]" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BF036D3BF7F8306FF7E15EFFB5C1A42" blockId="8.[151,1436,613,1792]" box="[199,1272,757,782]" pageId="8" pageNumber="9">
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<emphasis id="B93BEAC1BF7F8306FF7E15EFFE3A1A41" bold="true" box="[199,414,757,782]" pageId="8" pageNumber="9">Referred material</emphasis>
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. CHIHUAHUA.
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<collectionCode id="ED5EAE16BF7F8306FDDA15EFFD101A42" LSID="urn:lsid:biocol.org:col:34871" box="[611,692,757,781]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
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854942, topotypes, leg. R.H. and J.J.L.,
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<date id="FFF11013BF7F8306FBC415EFFB501A42" box="[1149,1268,757,781]" pageId="8" pageNumber="9" value="1998-12-08">8/xii/1998</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF7F8306FF7E1403FAA41A1A" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BF036D3BF7F8306FF7E1403FAA41A1A" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
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<emphasis id="B93BEAC1BF7F8306FF7E1403FEE51A7D" bold="true" box="[199,321,793,818]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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Shell medium-sized, conic or turriform, apex usually eroded; penis having two distal papillae on inner edge and a basal papilla both on the inner and outer edges. Contrasted with
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<taxonomicName id="4C4F4D50BF7F8306FB9F1424FB091A1A" box="[1062,1197,830,853]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="peregrina">
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<emphasis id="B93BEAC1BF7F8306FB9F1424FB091A1A" box="[1062,1197,830,853]" italics="true" pageId="8" pageNumber="9">T. peregrina</emphasis>
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</taxonomicName>
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above.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF7F8306FF7E147BFE501F06" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BF036D3BF7F8306FF7E147BFA321D46" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
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<emphasis id="B93BEAC1BF7F8306FF7E147BFEF51A35" bold="true" box="[199,337,865,890]" pageId="8" pageNumber="9">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF7F8306FE1E147BFE4E1A36" box="[423,490,865,889]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
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E–G) up to
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<quantity id="4CB79B36BF7F8306FDD5147BFD661A36" box="[620,706,865,889]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="9" unit="mm" value="6.7">6.7 mm</quantity>
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tall, whorls 6.5–8.75, spire height 150–200% width of shell, sexual dimorphism of shells not obvious. Teleoconch whorls weakly convex to almost flat, evenly rounded or, less frequently, narrowly shouldered, with adpressed sutures. Aperture sharply angled adapically, parietal lip usually incomplete, usually adnate when complete, rarely slightly disjunct, umbilicus absent. Outer lip thin, orthocline or weakly prosocline. Sculpture of strong growth lines and well developed spiral threads (
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<figureCitation id="13742A56BF7F8306FBDE14EBFB0E1D46" box="[1127,1194,1009,1033]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
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L). Periostracum tan.
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||
</paragraph>
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<paragraph id="8BF036D3BF7F8306FF7E130FFA881D3A" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
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||
<emphasis id="B93BEAC1BF7F8306FF7E130FFE6D1D61" bold="true" box="[199,457,1045,1070]" pageId="8" pageNumber="9">Shell measurements (</emphasis>
|
||
mean in parentheses): height
|
||
<quantity id="4CB79B36BF7F8306FCA0130FFC1A1D62" box="[793,958,1045,1069]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.65" metricValueMin="3.85" pageId="8" pageNumber="9" unit="mm" value="4.25" valueMax="4.65" valueMin="3.85">3.85–4.65 mm</quantity>
|
||
(4.20), width
|
||
<quantity id="4CB79B36BF7F8306FBDA130FFAAC1D62" box="[1123,1288,1045,1069]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" metricValueMax="1.78" metricValueMin="1.6" pageId="8" pageNumber="9" unit="mm" value="1.69" valueMax="1.78" valueMin="1.6">1.60–1.78 mm</quantity>
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||
(1.69), body whorl height
|
||
<quantity id="4CB79B36BF7F8306FE941323FE741D1E" box="[301,464,1081,1105]" metricMagnitude="-3" metricUnit="m" metricValue="2.255" metricValueMax="2.41" metricValueMin="2.1" pageId="8" pageNumber="9" unit="mm" value="2.255" valueMax="2.41" valueMin="2.1">2.10–2.41 mm</quantity>
|
||
(2.21), body whorl width
|
||
<quantity id="4CB79B36BF7F8306FD401323FC391D1E" box="[761,925,1081,1105]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.67" metricValueMin="1.53" pageId="8" pageNumber="9" unit="mm" value="1.6" valueMax="1.67" valueMin="1.53">1.53–1.67 mm</quantity>
|
||
(1.60), aperture height
|
||
<quantity id="4CB79B36BF7F8306FB1E1323FAEE1D1E" box="[1191,1354,1081,1105]" metricMagnitude="-3" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.41" metricValueMin="1.23" pageId="8" pageNumber="9" unit="mm" value="1.3199999999999998" valueMax="1.41" valueMin="1.23">1.23–1.41 mm</quantity>
|
||
(1.31), aperture width
|
||
<quantity id="4CB79B36BF7F8306FEF91347FE471D3A" box="[320,483,1117,1141]" metricMagnitude="-4" metricUnit="m" metricValue="9.95" metricValueMax="10.8" metricValueMin="9.1" pageId="8" pageNumber="9" unit="mm" value="0.995" valueMax="1.08" valueMin="0.91">0.91–1.08 mm</quantity>
|
||
(0.99), total number of whorls 6.50–7.25 (6.78) (USNM 1153679,
|
||
<emphasis id="B93BEAC1BF7F8306FB6D1344FB461D3A" box="[1236,1250,1118,1141]" italics="true" pageId="8" pageNumber="9">n</emphasis>
|
||
= 15).
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF7F8306FF7E1398FC751DF2" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93BEAC1BF7F8306FF7E1398FDA71DD5" bold="true" box="[199,515,1153,1178]" pageId="8" pageNumber="9">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF7F8306FE251398FDA71DD5" box="[412,515,1154,1178]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF7F8306FDD9139BFD631DD6" box="[608,711,1153,1177]" metricMagnitude="-3" metricUnit="m" metricValue="4.39" pageId="8" pageNumber="9" unit="mm" value="4.39">4.39 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF7F8306FCA0139BFC241DD6" box="[793,896,1153,1177]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="9" unit="mm" value="1.68">1.68 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF7F8306FBD8139BFB6D1DD6" box="[1121,1225,1153,1177]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="9" unit="mm" value="2.3">2.30 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF7F8306FF2E13BFFF591DF2" box="[151,253,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="mm" value="1.6">1.60 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF7F8306FE0113BFFDBA1DF2" box="[440,542,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="9" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF7F8306FD6A13BFFC9D1DF2" box="[723,825,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="8" pageNumber="9" unit="mm" value="1.01">1.01 mm</quantity>
|
||
, 7.25 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF7F8306FF7E13D3FF5B1CF6" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
|
||
Edges of last operculum whorl strongly frilled on outer side (
|
||
<figureCitation id="13742A56BF7F8306FCC713D3FC601DAE" box="[894,964,1225,1249]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
J); muscle attachment scar margins somewhat thickened on inner side of operculum (
|
||
<figureCitation id="13742A56BF7F8306FD3E13F7FD6F1C4A" box="[647,715,1261,1285]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
K). Radula (
|
||
<figureCitation id="13742A56BF7F8306FCEC13F7FC3C1C4A" box="[853,920,1261,1285]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
P-R): dorsal edge of central radular teeth concave, basal tongue rounded or V-shaped, median cusps elongate, distally pointed, parallel-sided proximally, lateral cusps five–six, basal cusps two–three (innermost larger) (
|
||
<figureCitation id="13742A56BF7F8306FC9D122FFCCE1C02" box="[804,874,1333,1357]" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
Q). Lateral teeth having two–three cusps on inner and four–six cusps on outer side, length of outer wing 175–200% width of cutting edge, central cusp pointed (
|
||
<figureCitation id="13742A56BF7F8306FAD61243FF021CDA" captionStart="FIGURE 4" captionStartId="9.[151,244,1781,1802]" captionTargetBox="[151,1436,191,1759]" captionTargetId="figure@9.[151,1436,191,1760]" captionTargetPageId="9" captionText="FIGURE 4. Tryonia peregrina sp. nov. and Tryonia zaragozae sp. nov. A. Holotype, T. peregrina, USNM 874032. B – D. Shells, T. peregrina, USNM 873288 (C and D are males). E. Holotype, T. zaragozae, USNM 873286. F – G. Male shells, T. zaragozae, USNM 874120. H – I. Operculum (outer, inner sides), T. peregrina, USNM 1153677. J – K. Operculum (outer, inner sides), T. zaragozae, USNM 1153679. L. Apical view of shell showing spiral sculpture on teleoconch, T. zaragozae, USNM 1153679. M. Portion of radular ribbon, T. peregrina, USNM 1153677. N. Central radular teeth, T. peregrina, USNM 1153677. O. Lateral tooth, T. peregrina, USNM 1153677. P. Portion of radular ribbon, T. zaragozae, USNM 1153679. Q. Central radular teeth, T. zaragozae, USNM 1153679. R. Lateral tooth, T. zaragozae, USNM 1153679. Scale bars A – G = 1.0 mm, H – L = 250 µm, M, P = 25 µm, N – O, Q – R = 10 µm." httpUri="https://zenodo.org/record/278446/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
R). Inner marginal teeth with 24–35 cusps, outer marginal teeth with 30–40 cusps. Radula data are from USNM 1153679.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF7F8306FF7E12DFFE501F06" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF7F8306FDF812DFFD201C92" box="[577,644,1477,1501]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
B) having two distal papillae on inner edge and a basal papilla both on the inner and outer edges (30 of
|
||
<specimenCount id="9D49FD5ABF7F8306FDB212F3FD0D1F4E" box="[523,681,1513,1537]" pageId="8" pageNumber="9" type="generic">30 specimens</specimenCount>
|
||
). Papilla on outer edge sometimes very small. Distal bulb of penis expanded laterally on inner side, lightly pigmented; stylet small. Penial duct undulating for most of length. Penial data are from USNM 1153679.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF7F8306FF7E114DFF581E4F" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="8BF036D3BF7F8306FF7E114DFF581E4F" blockId="8.[151,1436,613,1792]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93BEAC1BF7F8306FF7E114DFE551F3F" bold="true" box="[199,497,1623,1648]" pageId="8" pageNumber="9">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C4F4D50BF7F8306FE411143FD6D1F3F" box="[504,713,1625,1648]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="zaragozae">
|
||
<emphasis id="B93BEAC1BF7F8306FE411143FD6D1F3F" box="[504,713,1625,1648]" italics="true" pageId="8" pageNumber="9">Tryonia zaragozae</emphasis>
|
||
</taxonomicName>
|
||
is known only from its
|
||
<typeStatus id="54F48871BF7F8306FC6F1143FBA21F3E" box="[982,1030,1625,1649]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality, a large (ca.
|
||
<quantity id="4CB79B36BF7F8306FB491142FAE81F3F" box="[1264,1356,1624,1648]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="km" value="0.01">0.01 km</quantity>
|
||
2) cienega and spring system (
|
||
<figureCitation id="13742A56BF7F8306FE261166FE451FDB" box="[415,481,1660,1684]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
|
||
C) in the upper Río Casas Grandes drainage (
|
||
<figureCitation id="13742A56BF7F8306FC641166FBBB1FDB" box="[989,1055,1660,1684]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
|
||
, locality 4). Snails were collected (on gravel and on soft substrates) from several springs and bogs (29–30°C) within this cienega. This habit has been severely degraded by overgrazing in recent years, although one of the larger head springs is now fenced (first noted in 1998).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |